a A ag i A a I a - ~: “ Ot OER! a 8 re mie I pana PRA he os re be nd ee ae pity taf. —< See BULLETIN THE AMERICAN MUSEUM OF NATURAL HISTORY — VoLtumE XLII, 1920-1921 NEW YORK PUBLISHED BY ORDER OF THE TRUSTEES 1920-1921 er —en. a ee eee uae ate —_ THE AMERICAN MUSEUM OF NATURAL HISTORY SEVENTY-SEVENTH STREET AND CENTRAL Park West, New York City Boarp OF TRUSTEES (As of December 31, 1921) PRESIDENT ' HENRY FAIRFIELD OSBORN First VICE-PRESIDENT SeconpD VicEe-PRESIDENT CLEVELAND H. DODGE J. P. MORGAN TREASURER SECRETARY HENRY P. DAVISON PERCY R. PYNE Ex-oFFIc10 THE MAYOR OF THE CITY OF NEW YORK THE COMPTROLLER OF THE CITY OF NEW YORK THE PRESIDENT OF THE DEPARTMENT OF PARKS ELECTIVE GEORGE F. BAKER ARTHUR CURTISS JAMES GEORGE F. BAKER, Jr. WALTER B. JAMES FREDERICK F. BREWSTER CHARLES LANIER THOMAS DeWITT CUYLER OGDEN MILLS WALTER DOUGLAS A. PERRY OSBORN CHILDS FRICK GEORGE D. PRATT MADISON GRANT THEODORE ROOSEVELT WILLIAM AVERELL HARRIMAN LEONARD C. SANFORD ARCHER M. HUNTINGTON JOHN B. TREVOR ADRIAN ISELIN FELIX M. WARBURG ADMINISTRATIVE OFFICERS (As of December 31, 1921) DrrEcToR ASSISTANT SECRETARY FREDERIC A. LUCAS GEORGE H. SHERWOOD ASsISTANT TREASURER THE UNITED STATES TRUST COMPANY OF NEW YORK iii ScrenTiFic STAFF (As of December 31, 1921) DIRECTOR Freperic A. Lucas, Sc.D. GEOLOGY AND INVERTEBRATE PALEONTOLOGY Epmunp Otts Hovey, Ph. D., Curator Cuester A. Reeps, Ph.D., Associate Curator of Invertebrate Paleontology MINERALOGY Hersert P. Wurrtock, C.E., Curator WOODS AND FORESTRY (Curatorship Vacant) LOWER INVERTEBRATES Henry E. Crampton, Ph.D., Honorary Curator Roy W. Miner, A.B., Associate Curator (In Charge) Witiarp G. Van Name, Ph.D., Assistant Curator ENTOMOLOGY Frank E. Lutz, Ph.D., Curator A. J. Mutcuier, Assistant in Coleoptera Frank E. Watson, B.S., Assistant in Lepidoptera JoserH Bequaert, Ph.D., Assistant in Congo Zoology ICHTHYOLOGY Basurorp Dean, Ph.D., Honorary Curator Joun T. Nicuous, A.B., Associate Curator of Recent Fishes E. W. Gupaer, Ph.D., Associate in Ichthyology HERPETOLOGY ~— G. K. Nosie, A.M., Assistant Curator (In Charge) Karu Parrerson Scumipt, A.B,, Assistant Curator ORNITHOLOGY Frank M. Cuapman, Se.D., Curator W. DeW. Mitver, Associate Curator Rowert CusuMan Murpny, Se.D., Associate Curator of Marine Birds James P. Cuaprn, A.M., Assistant Curator, African Birds Loupitow Griscom, A.M., Assistant Curator MAMMALOGY Roy C. Anprews, A.M., Associate Curator of Mammals of the Kastern Hemisphere H. EF. Anwrnony, A.M., Associate Curator of Mammals of the Western Hemisphere Hensent Lana, Assistant Curator, African Mammals Cant FE. Axe ey, Associate in Mammalogy iv —_—-- > =o se “_—a Scientific Staff VERTEBRATE PALHONTOLOGY Henry FatrFretp Osporn, LL.D., D.Sc., Honorary Curator W. D. Marruew, Ph.D., Curator Wa ter GRANGER, Associate Curator of Fossil Mammals Barnum Brown, A.B., Associate Curator of Fossil Reptiles Wiuram K. Grecory, Ph.D., Associate in Paleontology COMPARATIVE ANATOMY Wituram K. GrEcoRY, Ph.D., Curator S. H. Cuvuss, Assistant in Osteology ANTHROPOLOGY Ciark Wisster, Ph.D., Curator Puiny E. Gopparp, Ph.D., Curator of Ethnology N.C. Netson, M.L., Associate Curator of North American Archeology Cuar.es W. Mrap, Assistant Curator of Peruvian Archeology Louis R. Suttrvan, A.M., Assistant Curator, Physical Anthropology COMPARATIVE PHYSIOLOGY Ratrpw W. Tower, Ph.D., Curator PUBLIC HEALTH CHARLEs-Epwarp Amory Wrinstow, D.P.H., Curator PUBLIC EDUCATION GerorGe H. Sxerwoop, A.M., Curator G. Criypk FisHer, Ph.D., Associate Curator Ruts Crospy Noste, A.B., Assistant Curator Grace FisHer Ramsey, Assistant LIBRARY AND PUBLICATIONS ; Raupu W. Tower, Ph.D., Curator Ipa Ricuarpson Hoop, A.B., Assistant Librarian RESEARCH ASSOCIATES ALEssSANDRO F asp, Physiology CLARENCE L. Hay, A.M.,.Mexican and Central American Archeology George F.. Kunz, Ph.D., Gems Cuares W. Lena, B.S., Coleoptera J. Howarp McGrecor, Ph.D., Human Anatomy FRANK J. Meyers, Rotifera HerBert F. Scowarz, A.M., Hymenoptera A. L. TreapwE Lt, Ph.D., Annulata Wituiam M. Wueeter, Ph.D., Social Insects CONTENTS OF VOLUME XLIII PAGE Title-pammess 5 << 6s Secacn es Mis 05s Ss op oa see Saar me ae ee 1 Officers:and Trustees... 2.525 osc se SR ee ae Se aha nag ae lil Sontific Staff. 3.3.25 en eS nz iv Contents... 2.65 0632 a eee ee ae vi Dates of Publication of Separata.............. Pee ey eR ar eee vi List of Illustrations. .. 25.60 Se ee eee eee Vii List of New Taxonomic Nameés;- >... 85 Ge PO ea eae es Pee ee Ferrata 555 on a ee a icc es ee ee a xii Art. I.—Parasitic Copepods From the Congo Basin. By CHARLES BRANCH Wiuson.::: (Plates THis Ss ea i oe eee 1 II.—Tipulide Collected by the American Museum Congo Expedition. By CHares P. ALEXANDER. (Plate IV, one text figure)... . 9 I1I.—Neuroptera, Panorpata, and Trichoptera Collected by the American Museum Congo Expedition, With Lists of the Species Known from the Belgian Congo. By NaTHaNn BANKs............ 21 1V.—African Stone-Flies and May-Flies Collected by the American Museum Congo Expedition. By James G. NEEDHAM. (Plate: Wes pies ees wee ce es Cts gn 35 V.—Isopods Collected by the American Museum Congo Expedition. By Wittarp G. Van Name. (One hundred and twenty-six GOEE TOG) circ hesis scm ae Lac ane agai eee Lae ae 41 V1I.—Lepidoptera of the Congo, Being a Systematic List of the Butter- flies and Moths Collected by The American Museum of Natural History Congo Expedition, Together with Description of Some Hitherto Undescribed Species. By W. J. HoLLAnp. (Plates VI-XIV and nine text figures). .................55. 109 VII.—Amphipods Collected by the American Museum Congo Expedi- tion, 1909-1915. By CLarence R. SHommaker. (Six text VIII.—The Brachyuran Crabs Collected by the American Museum Congo Expedition, 1909-1915. By Mary J. Rarssun. (Plates XV-LXIV, one map, and twenty-two text figures).. 379 Index to Article Vii 856i Hci vids be SON cL eee aE tie ere 333 Index to Article VIET: cei nievc Vink Rae Oa ee teat 469 Index to Articles I-V, ViVi). c:ciscasn dvd athe eh ie ae wks 475 DATES OF PUBLICATION OF SEPARATES The edition of separates is 300 copies, of which about 100 are mailed on the date of issue, and the others placed on sale in the Library. Art. I, April 22, 1920. Art. V, Sept. 27, 1920. OEE 5; a Ses * VI, Dec. 4, 1920. My AAG, a AO) TANG tO eee “ IV, “ 28, 1920. “ VIL, Sept. 30, 1921, vi era Se ee ee ee ee et LV ee LIST OF ILLUSTRATIONS PLATES I.—The male and female of Argulus reticulatus. II.—The male and female of Argulus ambloplites. III.—The females of Lernwa haplocephala and Brachyella macrura. IV.—Wings of Limnobia congoensis, Lecteria africana, Tipula dolichopezoides, T’. langi, Nephrotoma chapini, and N. ruwenzoriana. V.—Pentagenia schoutedeni, Cenopsis fugitans, Neoperla dubia, and N. excisa. VI.—Planema nelsoni, Papilio ridleyanus, P. r. fumosus, Hypolimnas bartteloti var. obliterata, Cymothoé diphyia, and Neptis agatha. VII.—Bicyclus medontias var. obsoletus, Euphedra imitans, Precis stygia stygia, P. s. gregorii, P. s. fuscata, Cymothoé cyclades, Kallimula osborni, Mycalesis chapini, and Euryphene maximiniana. VIII.—Cymothoé herminia, C. h. var. poénsis, Neptis biafra, Cymothoé capellides, C. capella, C. langi, and C. adelina. IX.—Euphedra cyanea, Cymothoé cenis var. conformis, Euryphene lucasi, Euphedra medon var. innotata, and Cymothoé cenis var. rubida. X.—Cymothoé aramis, C. ogova, C. Reginw-Elizabethe, C. anatorgis, C. angulifascia, Mycalesis langi, and Euryphene fulgurata. XI.—Euphedra preussi notata, E. inanoides, E. preussi subviridis, E. rezioides, E. preussi fulvofasciata, E. p. angustior, and E. p. latefasciata. XII.—Abantis rubra, Triclema lutzi, Leptalina niangarensis, Spindasis aderna, Oxylides homeyeri, Spindasis chapini, Telipna rothioides, T. medjensis, Pentila cletensi, Epitola langi, Pentila clarensis, Diestogyna kahli, and D. rotundata. XITI.—Xanthospilopteryx medjensis, Amphicollia pactolica, Pitthea famulita, Metarctia chapini, Pseudogonitis variabilis, Eutelia nigricans, Tolna bolengensis, Rhyn- chodes avakubi, Amphigonia hyalinata, Heterospila (?) rubida, Leocyma congoén- sis, Sphingomorpha aliena, Dermaleipa nubilata, Deinypena fulvida, D. trans- versata, D. morosa, and Pseudoterpna (?) chapinaria. XIV.—Leipoxais punctulata, Sapelia bipunctata, Deinypena multilineata, Osteosema phyllobrota, Lelia soloides, Callocossus langi, Gonanticlea (?) langaria, Ctenogyna (2) medjensis, Deinypena obscura, Proterozeuxis (?) medjensis, Rhamidava pieri- dara, and Phasicnecus grandiplaga. XV.—Geograpsus lividus and Callinectes latimanus. XVI.—Sarmatium curvatum and its habitat. XVII.—Habitat of Cardisoma armatum. XVII.—Pisa carinimana and Dromia atlantica. XIX.—Callinectes marginatus and C. gladiator. XX.—Callinectes marginatus and Potamon (Potamonautes) floweri. XXI.—Callinectes latimanus. XXII.—Callinectes latimanus and Geograpsus lividus. XXII.—Thalamita africana. XXIV.—Potamon (Potamonautes) dybowskii. XXV.—Potamon (Potamonautes) lirrangensis. XXVI.—Potamon (Potamonautes) stanleyensis and P. (P.) lirrangensis. XXVII.—Potamon (Potamon) ballayi. Vii Vill — Illustrations XXVIII.—Potamon (Potamon) ballayi, P. (Geothelphusa) perparvus, and P. (G.) congoénsis. XXIX.—Potamon (Geothelphusa) congoénsis. XXX.—Potamon (Geothelphusa) perparvus. XXXI.—Potamon (Acanthothelphusa) faradjensis. XXXII.—Potamon (Acanthothelphusa) langi. XXXIII.—Erimetopus brazze. XXXIV .—Deckenia mitis. XXXV.—Menippe nanus and Pilumnus verrucosipes. XXXVI.—Pilumnus verrucosipes and Eurypanopeus blanchardi (?). XXXVII.—Panopeus africanus. XXXVIII.—Grapsus grapsus. XXXIX.—Goniopsis cruentata. XL.—Pachygrapsus gracilis and P. transversus. XLI.—Sesarma (Chiromantes) africanum. XLII.—Sesarma (Chiromantes) alberti, S. (C.) africanum, and Sarmatium curvatum, XLIII.—Sesarma (Holometopus) angolense. XLIV.—Sesarma (Holometopus) elegans. XLV.—Sesarma (Holometopus) angolense, S. (H.) eats and Uca tangeri. XLVI.—Sarmatium curvatum. XLVII.—Sarmatium curvatum, Cyclograpsus occidentalis, and Sesarma (Holometopus) biittikoferi. XLVIII.—Cardisoma.armatum and Sesarma (Chiromantes) atherts. XLIX.—Cardisoma armatum. L.—Cardisoma armatum. LI.—Cardisoma armatum. LII.—Ocypode ippeus. LIII.—Ocypode africana. LIV.—Uca tangeri. LV.—Shore of the Atlantic Ocean near San Antonio, Angola. Sandy cove on left bank of Congo estuary near San Antonio. LVI.—Shore on east side of Banana peninsula, at incoming tide. Shore vegetation along bay north of San Antonio, at low tide. LVII.—Wild date palms, north of Banana, after a grass fire. Laterite blocks on the beach of the Atlantic near Moanda. LVIL1.—Forest of mangroves bordering a creek near Malela, at high tide. LIX.—Interior of a mangrove forest near Malela, at low tide. LX.—Forest of mangroves bordered by Raphia near Malela, at incoming tide, Edge of bushy mangrove formation along Banana Creek, at low tide. LXI.—Bank of the Congo about 17 miles from Banana, between Malela and Ponta da Lenha, at low tide. LXII.—Bank of the Congo just below Malela, at outgoing tide. LXIII.—Tschopo River near Stanleyville, at low water, in March, LXIV.—Forest brook in the Rain Forest of the Northeastern Belgian Congo south of Poko, at the end of January. Text Fiaures Pace Details of pupa of Tipula species from Mt. Ruwenzori.......0. 066.666. .00 5. 18 Eurydice carangia, new species: female and second pleopod of male,......... 50 te oe Illustrations : ix ' PaGE Eurydice carangis, new species: mouth-parts of female.................--. 51 Eurydice carangis, new species: legs of female................006.00000005 52 PACH, CODMMNMON! TOMAR? 26555. 256:i5cior stag Bal ete ea SANS Nb ele ele aes 54 Nerocila cephalotes: young individuals. ................ 0622 e cece cece eens 56 es RRR i ari pate Po 3 Oar arta 58 RES * Sa SS Sen cal yor a a re nn 60 Ichthyoxenos expansus, new species: female..............022 0000002 eee eens 61 Ichthyoxenos expansus, new species: female...............605:00+ ee eeeees 62 SERINE CRIP ICSE” LOTIONO So Sg oii 5 Ale att vn CREE Ce ao gel She oie woe 64 Spheroma destructor: mouth-parts of female... ........00.0 6066s 65 Spheroma destructor: legs of females. ...... 2.0.20. ccc cere sheen ees 66 Pleurocrypta langi, new species: dorsal and ventral aspects and first opercular NT ee een og AES A So WP EO GT mw Bly ioe Gk MOEN OE GE 68 Pseudione chapini, new species: dorsal and ventral aspects of femaleand male.. 70 RINE, RUM URAR INN ES J.D Cote ete et xs WAU ewes) VAs OSs Sele aCe va ais 74 Ligyda exotica: legs of female and male................... 0.04 ee cee eens 75 OT NT SRS at a Ne ae ai nn ae re 76 Ligyda olfersii: legs of female and male....................00 0 cece eee ees 77 Eubelum stanleyanum, new species: female and details of structure of female and ee amas rae trp Ether DOLE cicd ow Sole leSve x was ou whi ees 79 Eubelum stanleyanum, new species: female................0 0.0 cc cece ee 80 Eubelum stipulatum: female and details of structure of femaleand male........ 82 CII! SURMMB a ee lignes canes oe en Cause eid wendgaus be 83 Eubelum propinquum, new species: details of structure of male.............. 85 Eubelum asperius, new species: details of structure of female................ 86 Eubelum garambe, new species: details of structure of female................ 88 Eubelum garambe, new species: female. .................00 0 cee cece eee ee 89 Eubelum tenebrarum, new species: female and details of structure of male and eS SO Sire ny Slee PET he ODE IY oie Wats a 9 bw bd CaN he 90 Eubelum tenebrarum, new species: female................. 2.0002 e eee eee 91 Synarmadillo globus: male and details of structure of male.................. 93 SOP MARIN MEDEA? INGLES 6d ia. ye Goat ares Ga FO aks SE a ya Nae Bae 94 Synarmadillo lubilensis, new species: female and details of structure of female. 96 Synarmadillo lubilensis, new species: female... ...........0000 6c cee eee 97 Cubaris banane, new species: female and details of structure of female and male 98 Cubaris regulus, new species: female and details of structure of female........ 101 en MANNIE TOMNGIO 60. cig sates sins ale aia howe eee Ould eats Paw ies eee shee 103 Niambia squamata: male and details of structure of female and male......... 104 Deudoriz batikelides, new species: female from Mombasa....... RS See raion 222 PEADMMA Lae, NEW BPCGIOS: MAIO. 6.) . ac wa es seta esc kt eee 263 Preis RO OW BNOUIESS MAIO. ss eos ek ae bb Ge dle ak ences ee 266 Deilemera anomala, new species: male and female......................... 269 Lelta hildoides, new species: male... . 2.5. ce ccc ees cece eens 298 Acidalia medjaria, new species: male................ FEC ae oe ae 312 Chrysopoloma nubila, new species: female...................00 0000 e eee eee 322 Saliunca rubriventris, new species: female....................0 0000-0020 eee 324 Phryganodes sex-guttata, new species: male... ........6.. 02 ee eee 328 x Illustrations PAGE Elasmopus congoénsis, new species: details of structure of male.............. 372 Elasmopus congoénsis, new species: details of structure of male......... Se ciee
and
projects strongly from the posterior margin.
Color, the same as in the female except that the black pigment is in widely
scattered spots instead of a continuous network.
Total length, 6mm. Carapace, 5 mm. long, 4mm. wide. Abdomen, 1.25 mm.
long, 1.30 mm. wide.
- Remarks.—Mention has been made of eight African species already
reported; from them and from all other species, the present one is
distinguished by the intricate reticulation of the dorsal surface of the
female, by the squarely truncated teeth on the basal plate of the maxil-
lipeds, by the peculiar pattern of the respiratory areas of the carapace,
by the structure of the supporting rods of the marginal membranes of the
sucking disks, and by the peculiar lobes on the fourth legs of the female.
Argulus ambloplites,! new species
Plate II
Host and Record of Specimens——A male and female were secured
from the outside surface of the gills of Ophiocephalus obscurus Giinther
at Faradje on the Dungu River, very near the headwaters of that part
of the Congo River Basin which is north of Lake Albert Nyanza and
close to the watershed separating the Congo from the Nile Basin.
Specific Characters of Female
Carapace ovate, considerably longer than wide and reaching beyond the center
of the abdomen, with the posterior lobes considerably narrowed and turned inward;
cephalic area broadly triangular and projecting moderately; lateral areas spindle-
shaped, narrowed both anteriorly and posteriorly; posterior sinus a little more than
one-third the length of the carapace and somewhat enlarged at its base.
Abdomen ovate, widest at its base, with broadly rounded sides and posterior
lobes; anal sinus less than one-eighth the length of the abdomen, as wide as deep,
squarely truncated at its base, anal lamin basal.
First antennz of medium size, basal joint with a blunt posterior process re-
_1Ambloplites: G&uBdbs, blunt, and drXires, armed, alluding to the spines on the antenne and
maxillipeds and the accessory processes.
4 Bi lletin American Museum of Natural History {Vol. XLII
enforced by a much larger one behind the insertion of the antenna; second segment
with stout anterior and lateral claws and a small posterior spine opposite the anterior
claw; second antennz with a long blunt process on the basal segment. Eyes rather
small, separated by a distance a little less than one-fourth the width of the carapace,
and situated just in front of the lateral sinuses; median eye very small and far
behind the lateral eyes. Sucking disks about one-eighth the diameter of the cara-
pace and opposite the lateral sinuses and widely separated. Maxillipeds of medium
size, the basal plate broadly triangular, produced posteriorly into three flattened
lamin, which are squarely truncated and pressed so tightly together that the
sinuses between them are mere lines. On the ventral surface inside the base of each
maxilliped is an accessory process, also flattened into a wide, squarely truncated
lamina. The first two pairs of swimming legs with flagella; the basal lobes on the
posterior pair small and rectangular.
Respiratory areas divided, the anterior portion a little more than half the diam-
eter of the posterior, ovate, ae its long axis inclined backwards and inwards; the
posterior portion somewhat enlarged at either end.
Color (preserved material) a uniform gray-brown.
Total length, 5 mm. Carapace, 4.50 mm. long, 3.75 mm. wide. Abdomen,
1.12 mm. long, 1 mm. wide.
Specific Characters of Male
Carapace elliptical, not narrowed posteriorly and not reaching the base of the
abdomen, but distinctly longer than wide; cephalic area projecting more than in the
female but relatively narrower; posterior sinus a little more than one-fourth the
length of the carapace, with parallel sides.
Abdomen ovate, a little longer than wide, less than a quarter of the entire length,
lobes broadly rounded; anal sinus one-fifth the length of the abdomen and narrower
than in the female.
Lobes on the posterior swimming legs extended laterally into a broad and blunt
cone; of the accessory sexual apparatus the peg on the fourth legs is exceptionally
large, as is also the semen receptacle on the third legs, and there is a long rounded
process extending backward from the center of the ventral surface of the second legs.
Color, a little lighter than that of the female.
Total length, 7 mm. Carapace, 5.30 mm. long, 4.50 mm. wide. Abdomen,
1,60 mm. long, 1.50 mm. wide.
Remarks.—This species is clearly distinguished from all the African
forms by the conspicuous overlapping of the abdomen by the carapace
lobes in the female, by the wide and shallow anal sinus of both’ sexes,
and by the blunt armature, especially of the maxillipeds.
These two argulids, coming as they do from widely distant localities,
one at the very headwaters of the Congo River and the other close to its
mouth, suggest that the whole basin may yield specimens upon further
examination. They are closely related to Thiele’s A. africanus and to
the present author’s A. americanus, but may be recognized by the
characters above given,
Or
1920} Wilson, Parasitic Copepods from the Congo Basin
Lernea haplocephala (Cunnington)
Plate III, Figures 20 to 22
Host and Record of Specimens.—EHight females were obtained from
the outside skin and flesh of Polypterus ornatipinnis Boulenger at Faradje
on the Dungu River, one of the headwaters of the Congo. One was ob-
tained January 1912, four in February 1912, one in January 1913, and
two in February 1913. They were all found on the sides of the body
near the tail; they had penetrated the skin between the scales, and their
cephalothorax with its processes was enclosed in a tough cyst just
beneath the skin. In obtaining the four, Feb. 25, 1912, the skin, the
scales, and a thin layer of flesh were removed entire with the parasites
in situ; these specimens are excellently preserved.
Specific Characters of Female
General form thick and stout; cephalothorax with four large fleshy processes
or horns, two ventral and two dorsal, very similar to those of L. cruciata. Each
horn is conical, thick, and often flattened laterally at the base and tapered to a
_ bluntly rounded point. The ventral horns are swollen a little more than the dorsal,
especially on their ventral margins and toward their tips. The four horns are
arranged like the letter X and each is a little more than one-fifth of the entire length.
The neck behind the cephalothorax increases gradually in diameter back to the
pregenital prominence. Just behind the horns and external to the second pair of
legs there is a swelling on either side of the neck which projects ventrolaterally, the
two coming together across the midline on the ventral surface, but not extending
on the dorsal surface. These accessory processes correspond to those found in
L. variabilis behind the primary horns.
The pregenital prominence is exceptionally large, not bilobed, and spherical in
shape. It projects more than half its diameter, giving the body a decided boot or
foot shape. The toe of the boot is the abdomen, which is somewhat inclined to the
body axis and whose diameter is nearly equal to that of the body in front of the
pregenital prominence. It is a half longer than wide, the same diameter throughout,
bluntly rounded, and shows no signs of anal laminz.
The first pair of swimming legs is on the ventral surface of the cephalothorax
between the bases of the ventral horns; the second pair is on the ridge connecting the
two accessory prominences just behind the horns. The relative distances of the four
pairs of legs from the anterior margin of the cephalothorax, calling the entire length
of the body 100, are 11.15:35:70.
In the specimen from which the drawing of the entire animal was made there
was a torsion of 180, the fourth legs being apparently on the side of the body opposite
the first pair.
No egg-strings were found upon any of the present specimens, but Cunnington
stated that the egg-sacs were moderately long, about one-fifth the length of the
body, and tapering. Each contains four.or five rows of eggs, with twenty-five or
thirty in a row.
6 Bulletin American Museum of Natural History [Vol. XLII
Head fairly large and ovate, being narrowed anteriorly between the bases of the
antennz; the anterior margin of the narrowed portion is somewhat reéntrant at the
center. The first antenne are three-jointed and well armed with sete, the second
pair are two-jointed, the joints of the same length, the terminal one armed at the tip
with curved claws and several short sete. The maxillipeds are stout, the terminal
joint small, spherical, and tipped with four stout curved claws.
Color (preserved material), a uniform yellowish white.
Total length, 10 mm. Transverse length of dorsal and ventral arms and the
head, 3.80 mm. Greatest diameter of the body, 1 mm.
Remarks.—This is one of the three species established by/Dr. Cun-
nington in 1914; his type specimen was taken from the soft region at the
junction of the pelvic fins of a large Polypterus congicus Boulenger.
Other specimens were obtained from different species of Polypterus in
the White Nile, and, so far as recorded, they were found in the soft
flesh at the junction of paired fins. But the present specimens, instead
of choosing such a locality, burrowed in between the hard scales on the
side of the body. Coming from the Dungu River, they extend the
habitat of the species to the Congo Basin, and make it reasonably
certain that this parasite is as widely distributed in Africa as is the
genus of fish (Polypterus) upon which it isfound. The present host is the
fourth species of the genus from which this parasite has been obtained.
Cunnington gave some excellent photographs of the species and a
good general description, but he did not locate the swimming legs, except
the second pair, nor did he describe any of the appendages. These
omissions are here supplied and, fortunately, they only emphasize the
validity of the species.
Cunnington stated: “The appendages appear to show compara-
tively minor differences within the limits of this genus, and have not
been appealed to for the purpose of establishing new species. Thus I
have not deemed it necessary to study in detail the head appendages
of my new forms since they are by no means easy to investigate, and my
material, with one exception, was very scanty”’ (p. 283).
While this may be true in a measure, and while it may be advisable
to base an artificial key upon characters easily observed, it should never
be forgotten that the final decision of the validity of a species must
always rest upon the structure of the appendages. Consequently, a
description and figures of the appendages are more or less indispensable
for the establishment of the species. Even in dealing with such bizarre
forms as these lernwids, where species may be distinguished fairly well
without resort to the appendages, the presentation of their details will
avoid all question.
1920] Wilson, Parasitic Copepods from the Congo Basin 7
The distinguishing characters of this species are the well-developed
and undivided pregenital prominence, the four horns of equal length
arranged like the letter X, and the small spherical terminal joint of the
maxillipeds, with its four curved claws.
Brachiella macrura,' new species
Plate III, Figures 23 to 28
Host and Record of Specimens.—Three females with egg-strings
were obtained from the gills of an African snapper, Neomenis fulgens
Cuvier and Valenciennes, at Banana on the mouth of the Congo River,
July-August 1915.
Specific Characters of Female —
Cephalothorax about the same length as the trunk, cylindrical and the same
_ diameter throughout, and inclined backward at right angles to the trunk. Head
neither enlarged nor separated in any way from the thorax, covered with an indistinct
carapace. Maxillipeds nearly even with the anterior margin of the head and forming
apparently a heavy under jaw; second maxille at the angle between the cephalo-
thorax and the trunk. Trunk narrowed anteriorly into a very short neck, more or less
wrinkled posteriorly, flattened dorsoventrally.
Four posterior processes of about the same length, two dorsal and two ventral;
a genital process between the two latter and a little ventral to them, about half their
length and the same diameter throughout, which is equal to that of the posterior
processes.
Egg-strings arising between the ventral and dorsal processes and considerably
more than twice their length, tapering gradually to a blunt point. Each string has a
diameter at its base one-fourth greater than that of the processes, and contains five
rows of eggs, forty to forty-five in a row, making the aggregate of each string a little
over two hundred eggs.
_ First antennz swollen at the base and three-jointed; second antenne biramose
and turned down squarely across the frontal margin of the head, the endopod (dorsal
ramus) large and bluntly rounded, one-jointed and armed at the very tip with tiny
spines, the exopod smaller, two-jointed, and tipped with a tuft of finger-like sete.
Mouth-tube short and wide, extending forward between the tips of the second anten-
ne. First maxille tripartite, the third division lateral and much smaller than the
two terminal ones; palp one-jointed, bipartite, the rami very short and stout, each
tipped with a single spine. Second maxillz about one-third the length of the cephalo-
_ thorax, slender, fused at the base and then separate as far as the bulla. Maxillipeds
large and stout, the basal joint armed on its ventral surface near the inner margin
with two large knobs, covered with small spines; the terminal claw has an accessory
spine at its base.
\Macrura: paxpds, long, and dupa, tail, alluding to the genital process.
Bulletin American Museum of Natural History (Vol. XLII
v2)
Color (preserved material), a brownish yellow.
Cephalothorax, 4 mm. long, 0.85 mm. in diameter. Trunk, 4 mm. long, 2 mm.
wide, 1.75 mm. thick. Posterior processes, 4 mm. long. Egg-strings, 9 mm. long.
Remarks.—This species is most closely related to Beneden’s chev-
reuxti, which is the only other one possessing a long genital process.
But in chevreuxii the head is much enlarged and distinctly separated from
the thorax, the trunk is as wide as long and nearly twice as wide as thick,
and the egg-strings are only a trifle longer than the posterior processes:
The name macrura is especially appropriate for this species because
not only is the genital process (tail) longer than in any other species
except chevreuxii, but the posterior processes and egg-strings are also
exceptionally long. The combination of the three makes it a question
as to whether in this case “the dog wags the tail or the, tail wags the
dog.”’
Buuretin A. M. N.H.
Vor. XLIII, Prare I
The Male and Female of Argulus reliculatus
9
Dorsal view of male.
First and second antennx.
Mazxilliped
il -
. Third and fourth swimming legs.
Dorsal view of female.
Fig. 7.
Fig. 8.
Fig. 9.
Fig. 10.
Rods supporting the membrane of
a the sucking dis se
espiratory areas of the carapace.
Maxilliped.
Fourth swimming leg.
5
“any,
ney
ap
sal
i 4
: iid
i
ay
dij
Buititetiy A. M.N. HH.” : Vor. XLII, Prats Il
The Male and Female of Argulus ambloplites
- 11. Dorsal view of male. Fig. 16. Rods supporting the membrane of
Fig. 12. First and second antenne. . the sucking disks.
Fig. 13. Second and third swimming legs. Fig. 17. Respiratory areas of carapace.
14. Fo eg. } Fig. 18. Maxilliped.
15. Dorsal view of female. Fig. 19. Fourth swimming leg.
me
ped ie
eae sss
ha
Buuretin A. M. N. H. ; VOL. we Prate Lil
24 22
The Females of Lernea haplocephala and Brachiella macrura
Side view of female of L. haplocephala. Fig. 25. Second antenna.
Dorsal view of head showing first and Fig. 26. Maxilla.
second antenne. Fig. 27. Maxilliped.
Maxilla. Fig. 28. Ventral view, showing genital and
Side view of female of B. macrura. posterior processes.
Dorsal view of head showing carapace.
59.57, 71 (67.5)
Article 11.—TIPULIDH COLLECTED BY THE AMERICAN
MUSEUM CONGO EXPEDITION’
By Cuarves P. ALEXANDER, Pu.D.
Piate IV
The crane-flies collected in the Belgian Congo by Messrs. Lang,
Chapin, and Bequaert were kindly submitted to me for determination
by Dr. Frank E. Lutz. The few specimens included represent only large
and rather conspicuous forms but some of these were of exceptional in-
_ terest. The number of species in the collection is seven, distributed
in five genera. Our knowledge of the Tipulide of the Congo is almost
nil, and further collections will be awaited with considerable interest and
anticipation.
The crane-fly fauna of the Ethiopian region exhibits some interest-
ing features that I have endeavored to summarize in the appended
table. The family Tanyderide and the Cylindrotomine and Pediciini
of the Tipulidz have not been recorded as yet from Africa or its islands,
and the Ptychopteride are represented only by a single Ptychoptera
from the Cape. The Limnobiini (Dicranomyia, Libnotes), Eriopterini
(Erioptera, Molophilus, Trentepohlia, Gonomyia), and Hexatomini (Lim-
nophila) are well represented in species. The Tipuline, likewise, are
- common and include a few curious endemic types. Genera that abound
in the American tropics and in the Orient (Geranomyia, Eriocera) are
here represented by few species. Still others that, in the Neotropical
region, occur in a variety of forms are here indicated only by a few,
often non-typical, species (Rhipidia, Teucholabis, Gnophomyia).
The apparent centers of distribution of some of the genera are
indicated by the abbreviations, as follows: E=Ethiopian; H= Hol-
arctic; N=Neotropic; Nea=Nearctic; O=Oriental; P= Palearctic;
A=Australasian; *= doubtful genera.
I. Cosmopolitan, or nearly so; representatives occurring in
most of the faunal regions of the world.
Ptychoptera (H) Subg. Empeda (H)
Dicranomyia Molophilus (H)
Geranomyia (N) Gonomyia (H)
: P scctentaaes results of The American Museum of Natural History Congo Expedition. Entomology
9
10
Il.
III.
VI.
VII.
Vill.
Bulletin American Museum of Natural History [Vol. XLII
Rhamphidia (A) Subg. Lezponeura (N)
Limnophila (H) Subg. Gonomyella (N)
Eriocera (N) Trimicra
Elephantomyia Longurio (E) ,
Orimarga (N) Tipula (H)
Erioptera Nephrotoma (H)
Tropicopolitan.
Styringomyia (EK, O) Trentepohlia (OY
Megistocera (E)
Oriental and Ethiopian; especially southern and eastern
Africa.
Thrypticomyia (QO) Scamboneura (O)
Libnotes (O) Tipulodina*
Conosia (QO) Ctenacroscelis (QO)
Neotropical and Ethiopian.
Ceratocheilus Lecteria
Center of Distribution in the Australasian region.
Tasiocera Habromastix
Dolichopeza
Center of Distribution in the Neotropical region.
Rhipidia Atarba
Teucholabis Toxorhina
Gnophomyia
Center of Distribution in the Holarctic or Palearctic region.
Limnobia (FL) Hexatoma (P)
Adelphomyia * (A) Ormosia * (1)
Orimarqula UT)
Endemie (South Africa),
Platylimnobia Leplotipula
Podoneura Idiotipula
1920] Alexander, Tipulide from the Belgian Congo 11
The types of the new species described in this paper are deposited
in the collection of The American Museum of Natural History. Para-
types of Lecteria africana are in the author’s collection.
TIPULIDZ
Limnobiine
Limnosia Meigen
Limnobia MxiGEn, 1818, System. Beschr., I, p. 92.
Limnobia congoensis, new species
Plate IV, Figure 1
Thorax shiny black and yellowish, the pro- and metapleura yellow; halteres
dark, pale at the base; legs dark; wings yellow, the outer margin broadly infumed
with darker, the veins narrowly seamed with dark brown; abdomen black, the
hypopygial region yellowish.
Female (?).—Length, about 10.2 mm.; wing, 12.6 mm.
Rostrum and palpi dark brown, the latter short. Antenne brownish yellow,
the scape above-dark brown; flagellar segments oval, each with a very long bristle.
Eyes large, the space on the vertex between them very narrow but little wider than
the diameter of the first antennal segment. Head brownish yellow, the area sur-
rounding the insertion of the antennze brown.
Pronotum dull yellow, above on the mid-line narrowly dark brown. Mesonotal
prescutum shiny; the usual three stripes very broad, black, entirely confluent
behind; the space anterior to and behind the lateral stripes yellowish. Scutal lobes
black, the median area paler; scutellum light yellow; postnotum black. Pleura
variegated yellowish and black; propleura, including the cox of the fore legs, deep
yellowish; mesopleura, including the coxe of the middle legs, black, excepting a
restricted dull yellowish area below the wing-root; metapleura, including the base of
the halteres, pale yellowish, continued onto the mesosternum between the middle
and hind cox. Halteres dark brown, except the basal third of the stem which is
conspicuously light yellow. Legs with the coxe as described above, the hind cox
black; posterior trochanters blackish, anterior trochanters reddish; femora dark
brown, most intense apically; tibie brown; tarsi dark brown. Wings with a strong
yellow tinge, the apical region and the posterior margin tinged with brown; stigma
dark brown, solid; narrow, dark brown seams along the cord and the longitudinal
veins, more conspicuous along Cu. Venation: Sc terminating opposite the end of
the sector; r at the tip of Ri; Rs moderately elongated, a little longer than cell
1st M2, angulated and slightly spurred at the origin; basal deflection of Cu at the
fork of M.
Abdomen black, the hypopygium and the sclerites immediately basad of it
orange-yellow. The tip of the abdomen of the unique type is slightly injured and I
cannot determine the sex with certainty but believe that it is a female. The very
narrow vertex suggests that the specimen may be a male.
12 Bulletin American Museum of Natural History [Vol. XLIIL
Habitat.—Belgian Congo.
Holotype.— 2 (?); Medje, 27° 30’ E., 2° 25’ N.; Sept. 15-30, 1910. (Lang and
Chapin Coll.)
This is a true member of the genus Limnobia and perhaps the first
to be described from the Ethiopian region although L. uniflava Riedel
may correctly be referred here. The present species bears a slight resem-
blance to forms like L. indigena Osten Sacken, etc.
The rather numerous species referred to this genus by Bergroth,
Speiser, Edwards, and others seem rather to be referable to Dicranomyia,
Rhipidia and Libnotes. Dr. Bergroth has written several important
papers and notes on tipulid synonymy. While admitting many of these
points, there are others to which I can by no means agree and it seems
opportune to speak of one of these now. In his latest paper on the
subject (‘Some Tipulid Synonymy,’ Psyche, XXII, pp. 54-59, April
1915) Dr. Bergroth, in speaking of the genera Limnobia and Dicra-
nomyia, writes as follows: “Alexander’s conception of these two genera
seems to be so different from that of Osten Sacken and all other authors
that an explanation of how he distinguishes them is much to be desired.’’
On the contrary, I maintain that my conception of the limits of
Dicranomyia corresponds very closely with that of Osten Sacken, and
that it is Bergroth and some other European writers who deviate from
that definition. The most important single feature entering into this
discussion concerns the relative length of the subcostal vein, it being
long and extending far beyond the origin of the sector in Limnobia,
but short and ending about opposite or before the origin of the sector in
most species of Dicranomyia. Dr. Bergroth has apparently overlooked
the group of species in the genus Dicranomyia (Osten Sacken, 1869,
Monogr., Smithsonian Miscell. Coll., No. 219, pp. 73-76) in which the
subcosta is fully as long as in Limnobia. The species included by Osten
Sacken, D. pubipennis, D. globithorax, D. rara, and D. simulans, are
from eastern America, but species in the same category are found in
practically all parts of the world where the genus occurs. In some coun-
tries they are more abundant than the species with the short subcosta,
but in the Holarctic region are generally very much less common, The
species that Dr. Bergroth would place in Limnobia (D. nebulosa,
eiseni, gloriosa, ete.) are quite comparable to the four species listed
above. In Europe, D. pilipennis Egger, which is apparently the same as,
or very closely related to, D. pubipennis, is a common species and must
be entirely familiar to Dr. Bergroth, and yet I believe he would hesitate
to refer it to Limnobia, in spite of the length of its subcostal vein.
1920] Alexander, Tipulide from the Belgian Congo 13
The American students of the Tipulide seem to have less difficulty
in reconciling the group of species with the long subcosta, as discussed
above, to the genus Dicranomyia, but among the European workers
several take a stand similar to that of Dr. Bergroth, especially Edwards
and Speiser. DeMeijere described his D. umbrata (Java) with a long
subcosta and correctly referred it to Dicranomyia, yet Edwards later
referred it to the genus Limnobia. Brunetti describes a variety of forms
as species of Limnobia; those with a short subcosta (tinctinervis, festiva)
are undoubted species of Dicranomyia, while his L. indica with the sub-
costa long is, in my opinion, possibly a Dicranomyia; L. longinervis,
on the other hand, is evidently a Libnotes. One of Bergroth’s species,
D. venusta, of western North America, exhibits a marked variability in
the length of the subcosta, ranging from specimens where the vein ends
opposite the origin of the sector, to others where it extends notably
beyond. The question arises: is this a Dicranomyia or a Limnobia,
in the sense of Bergroth? Indeed, what is apparently the same, or a very
closely related, species was desneiied later on by Dr. Williston as Lim-
nobia concinna.
I have stated elsewhere, and maintain again, that when the fauna
of the world is considered, the only characters that are available for
distinguishing Limnobia from Dicranomyia are so slight as to be difficult
of definition, yet perceptible. In such cases the student has to depend
largely on the habitus of the species concerned. This, of course, brings
in intuition and the personal opinion of the authority, but the only other
course left is to refer all of these genera, Dicranomyia, Rhipidia, Peri-
pheroptera, Dapanoptera, Goniodineura, Libnotes, etc., to Limnobia.
A somewhat similar condition obtains in the Eriopterini, and here Dr.
Bergroth has deplored the possibility of lumping the complex into one
or few genera. I, likewise, believe that this is to be done only as a last
resort. The constant accession of exotic Tipulide has by no means eased
_ the burden of the taxonomist and the difficulty of exactly defining
genera is even greater than it was a half century ago, at the time of the
appearance of Osten Sacken’s monograph. Recently the writer has
undertaken a study of the immature stages of the Tipulide and related
families. Representatives of over seventy species, including forty
genera and all the tribes and major groups, have been obtained.
A careful study of this material has thrown much light on the subject
of classification, and it seems that its continuance will do much
toward solving the problem. The Tesults of this see will appear in
another paper.
14 Bulletin American Museum of Natural History (Vol. XLII
Lecteria Osten Sacken
Lecteria OSTEN SACKEN, 1887, Berliner Ent. Zeitschr., XX XI, part 2, p. 206.
Lecteria africana, new species
Plate IV, Figure 2
Coloration brownish yellow or brownish gray, the mesonotal prescutum with
narrow, indistinct stripes; pseudosutural fovezs conspicuous; legs reddish brown
with the tips of the segments darker; wings pale yellow, deepest along the costa;
pale gray and dark brown spots at the forks and tips of many of the Moa J abdomen
very elongate, reddish brown.
Male.—Length, 29-30 mm.; wing, 22-23 mm.; abdomen, 24 mm.; hind leg,
femur, 13.3 mm.; tibia, 12.4 ina tarsus, 11.2 mm.
Rostrum soak palpi dark bec Antennz with the first scapal smash dark
brown, faintly gray pruinose; second segment brown; flagellum pale brownish
yellow; first scapal segment elongate, as long as the succeeding three taken together;
basal flagellar segments rounded or oval, beyond the fourth elongate-oval with long,
delicate verticils. Head with the eyes large, protuberant; vertex moderate in width;
head strongly narrowed behind, light brownish gray with delicate transverse lines;
an indistinct narrow, median longitudinal stripe.
Mesonotum large. Prescutum brownish yellow, sometimes slightly grayish,
with three indistinct stripes that are narrowly and indistinctly margined with darker;
the middle stripe is bisected by a narrow dark brown stripe; pseudosutural fovex
large, subreniform, dark brown; scutum yellow or slightly grayish, the lobes a little
darker medially; scutellum small, brownish yellow, narrowly brown medially, with
numerous long yellow hairs; postnotum grayish yellow with a narrow dark brown
median stripe. Pleura brownish gray. Halteres brown, the knobs darker. Legs with
the cox# brownish gray; trochanters brown; femora and tibie reddish brown, the
tips narrowly dark brown; two basal segments of the tarsi reddish tipped with dark;
apical tarsal segments dark brown; tibie# unarmed. Wings pale yellow,- the
costal region more intensely yellow; small dark brown seams at r, Sea, base
of R4+5 and base of 2s; paler brown seams at the tip of Ra, fork of M,+2 and less
indistinctly at the ends of most of the longitudinal veins and along Cu. Venation:
Sc very long, almost as long as R; Rs very long, greatly arcuated at its origin and
running very close to Ri, even more than is usual in the genus; basal deflection of
R4 +5 short, strongly arcuated; Rez long, strongly upcurved at the tip; r inserted on Rg
near its base and on FR a short distance from the tip; cell 1st Mz elongate, irregularly |
hexagonal; petiole of cell M, a little longer than this cell; fusion of Ms and Cw
slight.
Abdomen very long, reddish brown, the lateral margins of the tergites narrowly
dark brown; seventh tergite with a narrow brown or black median stripe; abdominal
segments with numerous yellow set#; a narrow basal ring on either side destitute of
seta,
Hatitat.—Belgian Congo.
Holotype.— #; Faradje, 29° 40’ 0., 3° 40’ N.; January 1913, (Lang and Chapin
Coll.)
Paratopotypes.— fo; January 1913 and April 1911,
Paratype.— i; Bagboro, 29° K., 4° 18’ N.; October 16, 1911.
a ee
1920) Alexander, Tipulide from the Belgian Congo 15
The hitherto known species of this genus are Neotropical but the
occurrence of undescribed forms in Africa was mentioned by the writer
some years ago (1913, Proc. U. S. Nat. Mus., XLIV, p. 493). The re-
semblance of L. africana to species of the tropical American genus
Psaronius Enderlein and especially to the genotype, P. obscurus
(Fabricius), isstriking and difficult of explanation. In the paper just
cited I expressed the belief that the two genera are confluent but they
seem to be readily separable, the species of Psaronius having very long
tibial spurs while the tibie in Lecteria are entirely unarmed. The
similarity in venation and general habitus is remarkable.
Tipuline
Meaistocera Wiedemann
Megistocera WimpEMANN, 1828, Aussereur. Zweifl. Ins., I, p. 55.
Megistocera bicauda Speiser
Megistocera bicauda SrrtsER, 1909, in Sjéstedt’s Kilimandjaro-Meru Exped., I,
part 10, pp. 538 and 54. Type locality: Mombo in Usambara.
A single female that agrees well with Speiser’s description, except
as to the color of the abdomen, which is dark brown above, the basal
tergites with a rather bright yellowish brown median stripe which
becomes obsolete about the fourth segment.
Belgian Congo: Stanleville, 25° 15’ E., 0° 30’ N.; February 1915.
(Lang and Chapin Coll.)
Trputa Linnzus'
Tipula Linn.xus, 1758, Syst. Nat., 10th Ed., p. 585.
Tipula dolichopezoides, new subspecies
Plate IV, Figure 3
General coloration dark brown, the prescutum without distinct stripes; pleura
' yellow with indistinct brownish blotches; legs dark brown; wings with a brown tinge,
most intense along the anterior margin to beyond the wing apex; a pale area before
and beyond the stigma and in cell 1st M2; cell 1st M2 small, pentagonal; abdomen
banded brown and yellow.
Fémale.—Length, about 12 mm.; wing, 11.8 mm.; middle leg, femur, 8.3 mm.;
tibia, 9 mm.; hind leg, femur, 8.6 mm.; tibia, 8.7 mm.
1Tipula brunettiana, new name for Tipula splendens a cag we Fauna British India,
Wiehe St (India); non Tipula splendens Doane, 1901, Jou _ ¥. Ent. Soc., p. 107
ni ates
16 Bulletin American Museum of Natural History (Vol. XLIII
Frontal prolongation of the head short, brownish yellow above, the sides darker
brown; nasus represented only by a few stout hairs. Palpi moderately elongated,
dark brown, paler at the joints. Antenne with the scape light yellow; first flagellar
segment elongate, yellow; remaining segments brown, the basal portion a little
darker; flagellum with long verticils. Head broad behind, dark brown, the front
and the anterior portion of the vertex yellowish.
Mesonotum dark brown without distinct stripes; abundant long black hairs on
the interspaces. Pleura dull yellow, indistinctly marked with brown, the mesopleura
largely dark; a dark area on the mesosternum between the middle and hind coxe.
Halteres slender, dark brown, the extreme base and apex paler. Legs with the coxz
and trochanters yellow; remainder of the legs dark brown. Wings with a slight
brown tinge; the costal region, including the wing apex, dark brown; broad brown
seams along the cord and less distinctly along the longitudinal veins; a pale oblitera-
tive area before and beyond the oval dark brown stigma and another completely
filling the small cell 1st Me. Venation: Rs short, gently arcuated, only a little longer
than R2+3; Re long, persistent, bisecting the outer obliterative area; cell /st Me
very small, pentagonal; cell M, long-petiolate, the petiole about equal to Rz+3;
fusion of Cu; and M3+4 rather extensive, beginning at the fork of M.
Abdominal tergites dark brown, excepting the broad basal and lateral margins.
Sternites yellow, the segments with the posterior margin rather broadly ringed with
brown; a narrow sub-basal transverse band of brown. Female ovipositor rather
large, with a long dorsal basal shield, castaneous; the tergal valves elongate, slender,
divergent apically, with the tips rounded; sternal valves a little shorter, compressed,
the tips subacute.
Habitat.—Belgian Congo.
Holotype.— 9 ; Stanleyville, 25° 15’ E., 0° 30’ N.; February 1915. (Lang and
Chapin Coll.)
This small species curiously suggests the genus Oropeza Needham,
but is apparently a true Tipula. Its resemblance to certain African
species of Dolichopeza, belonging to the subgenus T'richodolichopeza
Alexander, is also noteworthy.
Tipula langi, new species
Plate LV, Figure 4
General coloration dark brown; pronotal scutum narrow and very high; a
rounded dark brown spot on the margin of the prescutum at about mid-length;
seutellum yellow; wings brownish gray, a brown seam along r-m and the basal
deflection of Cu; abdomen reddish brown, with a dark brown lateral stripe.
Female.—Length, about 18.5 mm.; wing, 19.5 mm.
Frontal prolongation of the head elongate, dark brown above, paler, reddish
yellow on the sides; nasus elongate, slender; palpi dark brown, Antenne with the
scape brown; first flagellar segment elongate, dark brown, the base a little paler;
remainder of the flagellum broken. Head dark brown.
Pronotum dark brown, the scutum highly projecting. Mesonotal prescutum
yellowish brown, brighter auteriorly; a narrow indistinct median brown line; a
rounded blackish spot on the lateral margin of the prescutum at about mid-length
an bc ce kell
ee eee
1920] Alexander, Tipulide from the Belgian Congo 17
and just behind the pseudosutural fovee; scutum dark blackish brown, the centers
of the lobes yellowish brown; scutellum yellowish brown; postnotum similar, the
basal third dark brown. Pleura dull yellowish brown, the region near the anterior
spiracle more cinereous; a dark brownish black area on the lateral regions of the
postnotum behind the wing-root. Halteres elongate, dark brown, the knobs brighter
reddish. Legs with the coxe# and trochanters dull yellowish; femora and tibie
reddish brown, the tips dark brown; tarsi dark brown. Wings with a strong brown-
ish gray tinge, the costal region more brownish; stigma dark brown, narrow; a
narrow brown seam along r—m and the basal deflection of Cu. Venation: Rs shorter
than R,+;; R2 present but weak and cell R; consequently small and with its inner end
acute; R; short, straight; cell 1st M, large, ample.
Abdominal tergites reddish brown, a little brighter basally; lateral margins of
the segments broadly blackish, producing a dark lateral stripe; an indistinct blackish
median spot near the base of the segments; sternites yellowish. Female ovipositor
with the tergal valves very long and slender, the inner margin at the base fringed
with long hairs; sternal valves short, compressed, pale yellow, the tips broadly
rounded.
Habitat.—Belgian Congo.
Holotype.— 9 ; Avakubi, 27° 40’ E., 1° 20’ N.; Oct. 4-8, 1909. (Lang and
Chapin Coll.)
This species is closest to 7’. jocosa Alexander (Cape Colony) in the
peculiar venation, but cell R, is smaller and with its inner end even
more acute, while cell 1st Mz is comparatively small.
Tipula langi rubricapilla, new subspecies
Female.—Length, 18 mm.; wing, 17 mm.
Differs from the typical langi as follows: the head is much brighter in color,
reddish, as is also the dorsal shield of the ovipositor. The flagellum is brownish
black, the segments rather elongated and with long verticils. The high pronotal
scutum is broadly yellow medially, the sides dark brown, the mesonotum largely
dark, but the scutellum is entirely yellow. Pleura yellowish, with extensive dark
brown markings including the propleura, most of the mesopleura and the outer
face of the coxe. Abdomen dark brown, especially on the sternites.
Habitat.—Belgian Congo.
Holotype.— 9; Medje, 27° 30’ E., 2° 25’ N.; Sept. 1-7, 1910. (Lang and
Chapin Coll.)
Pupa of Tipula Species
A large tipuline pupa was secured by Dr. Joseph Bequaert in the
Butagu Valley at 2200 meters, western slope of Mt. Ruwenzori. The
specimen was taken in April 1914 from wet moss near a brook. There
can be little doubt but that the specimen belongs to a large species of the
restricted genus Tipula Linneus but any further determination is im-
possible at this time. The fly that would emerge from this pupa would
18 Bulletin American Museum of Natural History [Vol. XLIII
be almost as large as Tipula abdominalis (Say) of northeastern North
America. It would naturally be expected that this might be the pupa
of the large and vigorous Ctenacroscelis albovittatus (Macquart) which
has an extensive range in eastern Africa, but the venation of the wing-
pad precludes this reference. This specimen may be briefly described
as follows: :
A B Cc
Fig. 1. Details of pure of Tipula species from Mt. Ruwenzori. A, pronotal horn; B, ventral
aspect of the caudal end; C, lateral aspect of the caudal end.
Pupa.—Length, 37 mm.
Width, dextro-sinistral, at the wing-pad, 5.2 mm.
Depth, dorso-ventral, at the wing-pad, 5.3 mm.
Pronotal breathing horn, 2.5 mm.
Coloration of the alcholic pupa very dark reddish brown. The specimen was
evidently nearly ready to emerge when it was killed.
Labral sheath very broad, transversely wrinkled Sheaths of the maxillary
palpi strongly recurved at their tips. Pronotal breathing-horns short, equal in
length, cylindrical, the tips slightly. expanded (Fig. 1A). The mesonotum is ex-
pansive, with about eight prominent but depressed tubercles, the anterior four of
which extend across the dorsum in a semicircular transverse row; the outermost pair
are located just above the root of the wing. Wing-sheaths showing the venation
clearly, vein R2 not dipping strongly toward vein Rs; cell M; petiolate; vein m-cu
short or punctiform. Wing-sheaths reaching the base of the third abdominal seg-
ment. Leg-sheaths attaining the base of the fourth abdominal segment, the fore
tarsal sheaths shorter than the other legs.
Abdothinal tergites with a row of twelve small spines across the caudal margin
of the posterior ring, on either side of the median line with an additional slightly
larger spine grouped close to two of the series to form a close triangle. Pleurites
with a large spine on the anterior ring and two small spines placed side by side on
the posterior ring. Abdominal sternites with about eleven spines on the posterior
ring, the two outermost on either end of the row small and paired; two widely
separated larger spines on the basal half of the posterior ring. Female cauda sur-
rounded by ten powerful spines, four being on the sternum of the eighth segment,
two on the pleura of the eighth segment and four larger spines, set close together,
located at the base of the ninth tergum (Fig. 1 B and C). In addition to the four
1920} Alexander, Tipulide from the Belgian Congo 19
tergal spines there is an additional long,powerful lobe on either side at the base of
the tergal valves, these directed caudad and slightly dorsad. Tergal valves of the
ovipositor considerably longer than the sternal valves.
Specimen in the collection of the American Museum.
NEPHROTOMA Meigen
Nephrotoma Metcen, 1803, Illiger’s Mag., II, p. 262.
Nephrotoma chapini, new species
Plate IV, Figure 5
Head orange-yellow, including the frontal prolongation; no dark occipital
mark; mesonotum dull yellow with three broad black stripes; scutellum and post-
notum yellow, the apical third of the latter black; pleura yellow, spotted with
brown; wings brownish yellow, the cord indistinctly seamed with brown; abdomen
yellow, transversely banded with black.
Female.—Length, 15 mm.; wing, 12.4 mm.
Frontal prolongation of the head very short, yellow; nasus short with a tuft of
long black hair; palpi dark brown. Antenne with the first segment yellow; second
segment dark brown; flagellum very dark brown. Head rich orange-yellow, duller
. on the genx; vertical tubercle rather prominent; no shiny area on occiput.
Pronotum broadly yellow, the sides dark brownish black. Mesonotum dull
yellow with three very broad black stripes that are almost confluent behind, sepa-
rated only by a very indistinct line of chestnut brown; the space before the lateral
stripe triangular, dull yellow; scutum almost entirely black, the median area paler;
scutellum dull brownish yellow, the sides darker; postnotum yellow, the apical
third dark brownish black. Pleura yellow, the mesepimeron paler, almost whitish;
a large brown spot on the mesepisternum and another on the mesosternum, suffusing
the base of the middle coxe. Halteres dark brown, the extreme base and the knobs
a little paler. Legs with the coxe more or less infumed with brown, darkest on the
posterior coxe, which are almost entirely of that color; trochanters dull yellow;
femora black, the basal portion yellowish, this narrowest on the fore femora, broadest
on the hind femora; tibiz and tarsi black. Wings with a strong grayish yellow tinge,
costal area a little brighter; stigma brown; the cord and the longitudinal veins
indistinctly seamed with very pale brown. Venation: Rs very short, scarcely longer
than the basal deflection of Ry+5; cell M, broadly sessile.
Abdominal tergites banded black and yellow, the first tergite and the apical
half of segments two to four black, the apical two-thirds of segment five black; all of
segment six black except a small yellow spot on the sides at the base; segment
seven entirely black; segment eight yellow, margined with black; segment nine
yellow; ovipositor chestnut; lateral margins of the tergites broadly dark brownish
black; sternites approximately similar to the tergites.
Habitat—Belgian Congo.
Holotype.— 9 ; Stanleyville, 25° 15’ E., 0° 30’ N.; February 1915. (Lang and
Chapin Coll.)
20 Bulletin American Museum of Natural History [Vol. XLIII
In its banded abdomen this species suggests N. tigrina Alexander
of Portuguese East Africa (1917, Ann. South African Mus., XVII,
pp. 177-179) but in other respects there is not a great resemblance
between the two.
Nephrotoma ruwenzoriana, new species
Plate IV, Figure 6
Head orange-yellow, the frontal prolongation black above; thorax entirely
blue-black except the basal two-thirds of the postnotum which are orange-yellow;
halteres and legs black; wings strongly infumed with blackish; abdomen blatk with
only the last segment tedidixh.
Female.—Length, 17.5 mm.; wing, 12.8 mm.
Frontal prolongation of the head short, jet black above, reddish yellow on the
sides; nasus long, slender, black; palpishort, black. Antenne with the first segment
brownish orange; second segment brown; flagellar segments black. Head orange-
yellow, the occiput with a brown triangular mark that runs forward onto the vertex.
Pronotum very dark reddish brown; mesonotum entirely deep blue-black with
only the basal two-thirds of the postnotum bright yellow. Pleura blue-black.
Halteres and legs black. Wings broad, strongly tinged with blackish, especially
along the costal region and on the basal third, whence it is continued outward along
veins Cu and 2nd A; stigma dark brown; tip of the wing darkened, a broad seam
along the cord. Venation: Rs short, straight, oblique; cell Mj, rather narrowly °
sessile.
Abdomen entirely blue-black, with the exception of the ninth segment and the
valves of the ovipositor, which are yellowish chestnut.
Habitat.—Belgian Congo.
Holotype.— 2 ; Mt. Ruwenzori, 29° 50’ E., 0° 30’ N., on the western ner in the
Butagu Valley at 3000 m.; April 15, 1914. (J. Bequaert Coll.)
Most closely related to N. fuscipennis (Karsch) from Portuguese
West Africa (1886, Ent. Nachricht., XII, pp. 52 and 53). The two
species may be separated by the following key:
1. Female with the scutellum yellowish red; a yellow spot on pleura above the
middle cox; second and third abdominal segments more or less yellowish
TOM, . s:6:c'sisie wap a aaah es seb hindi N. fuscipennis (Karsch),
Female with the scutellum black; pleura and abdomen uniformly blue-
TURK, «5.1.0 ois one acbin ade eeione ss cae ea N. ruwenzoriana, new species.
‘sotoeds Mou ‘puDwozuamns DUOpLydaNT JO BUIM “9 “BIA ‘sotoeds Mou ‘sapiozadoyoyop vyndty jo Buty ‘“g ‘BIW
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‘sowoeds mou ‘26un) nyndty jo Bum ‘p ‘Big ‘soroeds mou ‘stsuaobuos viqouwV] JO Zum “1 “BI
*(u194Shg UTeYPe2 N-Y90}sUIOD) suleA [VUY= V ‘snyIqnD= np ‘vIpey= p ‘snipey = y !vysooqng= og
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-
59.57,4 (67.5)
Article III—NEUROPTERA, PANORPATA, AND TRICHOPTERA
COLLECTED BY THE AMERICAN MUSEUM CONGO
EXPEDITION, WITH LISTS OF THE SPECIES KNOWN
FROM THE BELGIAN CONGO!
By NatuHan BANKS
‘The collections brought from the Congo by Messrs. Lang and
Chapin contain nineteen species of true Neuroptera, one of Panorpata,
and five of Trichoptera. Many of these forms are widely distributed in
tropical Africa, but the collection, small as it is, adds several species to
the fauna of the Belgian Congo. A list of the Congo species hitherto
known from the three orders here considered has been compiled by Dr.
J. Bequaert and incorporated in the paper. While the neuropterous
fauna of that country is too incompletely known to give much informa-
tion with regard to its affinities, yet the occurrence of several forms from
southern Abyssinia in the extreme northeastern Congo Basin, as shown
by this collection, is noteworthy.
APPROXIMATE LOCATION OF PLACES MENTIONED IN THIS PAPER?
Barumbu.—1° 10’ N., 23° 20’ E. Lisala.—2° 10’ N., 21° 30’ E.
Beni.—0° 30’ N., 29° 30’ E. Matadi.—5° 50’ S., 13° 35’ E.
Bumba.—2° 10’ N., 22° 30’ E. Medje.—2° 25’ N., 27° 30’ E.
Faradje.—3° 40’ N., 29° 40’ E. Niangara.—3° 40’ N., 27° 50’ E,
Garamba.—4° 10’ N., 29° 40’ E. Rungu.—3° N., 28° E.
Kabare.—0° 35’ S., 29° 30’ E. Rutshuru.—1° 15’ S., 29° 30’ E,
Kasindi.—0°, 29° 40’ E. Stanleyville—0° 30’ N., 25° 15’ E.
NEUROPTERA
Myrmeleonide
Patpares Rambur
1. Palpares hamatus Kolbe
Palpares hamatus Kouse, 1898, Entom. Zeitg. Stettin, LIX, p. 232.
Twenty-four specimens: Faradje, November 1912; Niangara,
November 1910; Rungu, October 1910.
Not previously recorded from the Belgian Congo.—Described from
Abyssinia. According to P. Esben-Petersen, 1916, Ark. f. Zool., X, No.
15, p. 10, this is a synonym of P. tigris Dalman; if the other synonymy
Scientific Results of the Congo Expedition. Entomology, No. 4.
2Most of these localities are shown on the map accompanying the reports on Congo Vespid (1918,
a are Mus. Nat. Hist., XX XIX, p. 9) and on land mollusks (1919, Bull. Amer. Mus. Nat. Hist.,
Ps : :
21
22 Bulletin American Museum of Natural History [Vol. XLIII
proposed by that author prove correct, the species has a wide distribu-
tion on the African continent. Navas’ genus Nosa (1911, Rev. Zool.
Afric., I, p. 239) is based on a specimen of this species.
2. Palpares abyssinicus Kolbe
Palpares abyssinicus Ko.px, 1898, Entom. Zeitg. Stettin, LIX, p. 233.
Six specimens: Barumbu, July 1909; Niangara, November 1910;
Faradje, December 1910.
Not previously recorded from the Belgian Congo.—Abyssinia,
Eritrea.
3. Palpares latro Nav4s
Palpares latro NavAs, 1911, Rev. Zoolog. Afric., I, p. 238, Pl. xv1, fig. 3.
One specimen: Niangara, April 1913.
This species was described from a specimen without locality, prob-
ably from Central Africa; no other records of it have been published
thus far. But, according to P. Esben-Petersen (1916, Ark. f. Zool., X,
No. 15, p. 11), it is identical with P. festivus Gersteecker.
Centrociisis Nav4s, 1909
[(=Sogra Navas, 1912)
4. Centroclisis mordax (Navas)
Sogra mordax Nav&s, 1912, Mem. R. Ac. Cienc. Barcelona, (3) X, p. 151, fig. 7.
One specimen: Matadi, June 1915. :
Not previously recorded from the Belgian Congo.—Described from
German East Africa. According to P. Esben-Petersen (1916, Ark. f.
Zool., X, No. 15, p. 14) a synonym of C. distincta (Rambur) which has a
wide range in Africa.
Myrmeteon Linné
5. Myrmeleon obscurus Rambur
Myrmeleon obscurus Rampur, 1842, Hist. Nat. Ins. Névroptéres, p. 403. *
One specimen: Stanleyville, March 1915.
Recorded from the Kasai (Lukombe, Dima), Manyema (Kibombo),
and Katanga (Bukama).—A common species in tropical and South
Africa.
Cueta Nayvis
6. Cueta mysteriosa (Gerstecker)
Myrmeleon mysteriosus GuersrvmcKer, 1894, Mitth, naturw. Ver. Neu-Vorpom-
mern, XXV (1893), p. 141.
1920] | Banks, Congo Neuroptera, Panorpata, and Trichoptera 23
One specimen: Niangara, April 1913.
Not previously recorded from the Belgian Congo.—German and
British East Africa. .
Nevroteon Navas |
7. Neuroleon filiformis (Gerstzcker)
Myrmeleon filiformis Gerstmcker, 1885, Mitth. naturw. Ver. Neu-Vorpom-
mern, XVI, p. 31.
One specimen: Faradje, January 1913.
Not previously recorded from the Belgian Congo.—Sierra Leone,
Cameroon, French Congo. ‘
FORMICALEON Banks
8. Formicaleon harpalyce Banks
Formicaleon harpalyce BAnxs, 1911, Ann. Entom. Soc. America, IV, p. 17.
Four specimens: Niangara, November 1910; Faradje, December
1910. .
Not previously recorded from the Belgian Congo.—Togo.
9. Formicaleon turbidus (NavAs)
Formicaleo turbidus Nav4s, 1915, Mem. Acc. Rom. Nuovi Lincei, X XXIII,
ms.13,
Two specimens: Medje, March 1910; Stanleyville, March 1915.
Not previously recorded from the Belgian Congo.—Described from
the Congo without more definite mention of locality; probably from the
French Congo.
Macronemvrvs Costa
10. Macronemurus lepidus Kolbe
Macronemurus lepidus Kose, 1898, Deutsch Ost Afr., IV, Netzfliigler, p. 23.
; Two specimens: Kabare, August 1914 (J. Bequaert Coll.); Faradje,
December 1912.
Not previously recorded from the Belgian Congo.—German East
Africa. |
Creacris Hagen
11. Creagris nubifer Kolbe
Creagris nubifer Kousn, 1898, Deutsch Ost Afr., IV, Netzfliigler, p. 25.
One specimen: between Beni and Kasindi, August 1914 (J. Bequaert
Coll.). ;
24 Bulletin American Museum of Natural History [Vol. XLIII
Known from the northeastern Congo (Kinjawanga, north of Lake
Albert Edward, a locality close to Beni) and Katanga (Bunkeya).—
Togo, Abyssinia, German and British East Africa, South Africa.
12. Creagris africana (Rambur)
Myrmeleon africanus RamBur, 1842, Hist. Nat. Ins. Névroptéres, p. 395.
Eleven specimens: Faradje, January 1913; Niangara, November
1910. .
Not previously recorded from the Belgian Congo.—Widely distrib-
uted in Africa south of the Sahara.
CrmoTHates Gerstzecker
13. Cymothales johnstoni Kirby
Cymothales johnstoni Krrspy, 1902, in H. Johnston, The Uganda Protectorate,
I, p. 469.
One specimen: Stanleyville, March 1915.
Not previously recorded from the Belgian Congo.—The species
was known from Uganda (Entebbe) only.
This is the only specimen I have seen, except the type, with which it
agrees closely. It is most nearly related to C. dulcis Gersteecker, but is
paler throughout.
Ascalaphidee
Tmesripasis MacLachlan
14. Tmesibasis waelbroecki van der Weele
Tmesibasis waelbroecki van der Weele, 1908, Cat. Coll. Zoolog. de Selys Long-
champs, Fasc. 8, Ascalaphiden, p. 92, fig. 58. .
Garamba, June 1912.
Known only from the Belgian Congo: Kinshasa.
Heuticomitus MacLachlan
15. Helicomitus festivus (Rambur)
Bubo festivus Rampur, 1842, Hist. Nat. Ins. Névroptéres, p. 356.
Three specimens: Niangara, November 1910; Faradje, November
1912.
Recorded from the Manyema(Kibombo, 245 kilom.south of Kindu),
Uele (Bambili) and Katanga (Bukama, Watura-Katwe, Kapiri).—
A common species in Africa from the Senegal and Egypt to Natal;
also on Madagascar.
1920] Banks, Congo Neuroptera, Panorpata, and Trichoptera 25
16. Helicomitus rutilus (Gerstecker)
Suphalasca rutila GerstacKer, 1894, Mitth. naturw. Ver. Neu-Vorpommern, -
XXYV (1893), p. 105.
Two specimens: Faradje, December 1912.
Not previously recorded from the Belgian Congo, but doubtfully
distinct from H. festivus, with which it is united by van der Weele
(1908, Cat. Coll. Zoolog. de Selys Longchamps, Fasc. 8, p. 175).—
German East Africa.
DIsPAROMITUS van der Weele
17. Disparomitus bacillus (Gersteecker)
Suphalasca bacillus Gprst@cKER, 1885, Mitth. naturw. Ver, Neu-Vorpommern,
XVI, p. 5.
Three specimens: Medje, April 1910.
Recorded from the Lower Congo (Congo da Lemba). a, West
African species: Togo, Sierra Leone, Cameroon, French Congo.
Chrysopidz
Curysopa Leach
18. Chrysopa bequaerti Navas
Chrysopa bequaerti Navas, 1912, Rev. Zoolog. Afric., I, p. 409, fig. 4.
One specimen: Faradje, November 1912.
Known from the Belgian Congo only, where it is recorded in the
Lower Congo (Kitobola) and Katanga (Kongolo).
NotHocurysa MacLachlan
19. Nothochrysa rufostigma (MacLachlan)
' Chrysopa rufostigma MacLacuian, 1868, Journ. Linn. Soc. London, Zool., [X,
p. 253.
One specimen: Faradje, November 1912.
Not previously recorded from the Belgian Congo.—South Africa..
NEUROPTERA RECORDED FROM THE BELGIAN CONGO
Myrmeleonide
PatparREs Rambur
Africa, Madeira, Southern Europe, Persia, India. -
Palpares abyssinicus Kolbe. See above.
Palpares egrotus Gersteecker.—Congo (?)
26 Bulletin American Museum of Naturai History [Vol, XLIII
Palpares bayeri Navis.—Northeastern Congo (Beni). Probably a synonym of
. P. obsoletus Gerstecker.
' Palpares cognatus Rambur.—Belgian Congo.
Palpares hamatus Kolbe. . See above.
Palpares ictericus Navés.—Lower Congo — Probably a synonym of
P. egrotus Gerstecker.
Palpares latipennis Rambur. —Lower Cana (Banana), between Leopoldville
and Stanleyville.
Palpares latro Navas. See above.
Palpares nigrescens Navis.—Katanga (Sankisia).
Palpares normalis Navés.— Katanga.
Palpares obsoletus Gersteecker.—Stanley Pool.
Palpares radiatus Rambur.—Congo (?).
Palpares stuhlmanni Kolbe.—Northeastern Congo (Vichimibi).
Palpares tigris Dalman.—Katanga (Punga, Sankisia). According to P. Esben-
Petersen, Nosa leonina Navas was based on an aberrant specimen of the same species.
See above., f
PALPARELLUS NavAs
Africa. P. Esben-Petersen (1916, op. cit., p. 11) is of the opinion that this is not
generically distinct from Palpares.
Palparellus obscuripennis (FE. Schmidt).—Kasai, Katanga (Lusindoi).
Hacenomyia Banks
(=Nelees Navas)
Hagenomyia imperator (Navas).—Katanga (Kapiri).
Hagenomyjia indistincta (Navds).—Katanga (Sankisia).
Hagenomyia lyncea (Fabricius).—Kasai (Lukombe).
Hagenomyia tristis (Hagen).—Lower Congo (Kunga, Leopoldville, Tchoa),
Kasai (Lukombe), Katanga (Kongolo, Katwamba, Katolo, Kapiri, Shinsenda).
Syncenes Kolbe
Syngenes longicornis (Rambur).—Congo (?).
Centrociisis Navis
(including Sogra Navas),
Africa,
Centroclisis brachygaster (Rambur),—Lower Congo (Leopoldville), Kasai (Lulua-
bourg).
Centroclisia lineatipennis (Peringuey).—Kasai (Hemptinne-St.-Benoft near
Luluabourg).
Centroclisis mordax Navas. See above.
Centroclisis perversa (Navas).—Uele.
Centroclisis rufeacens (Gerstwcker) —Lake Albert.
1920] - Banks, Congo Neuroptera, Panorpata, and Trichoptera
MyrMeteon Linné
Cosmopolitan.
Myrmeleon atlas Banks.—Northeastern Congo (Kwidjwi).
Myrmeleon lethifer Walker.—Northeastern Congo (Kwidjwi).
Myrmeleon obscurus Rambur. See above.
Myrmeleon simplicissimus Gerstecker.—Katanga (Kapiri).
Gymno.teon Banks
Africa,
Gymnoleon (?)cognatus Navés.—Katanga (Bukama).
Gipretta Navas
Africa.
Gibrella congolana Navds.—Katanga (Kapiri).
Macroteon Banks
Macroleon polyzonus (Gerstecker).—Equator (Busira).
Creacris Hagen
Creagris africanus (Rambur). See above.
Creagris nubifer Kolbe. See above.
Netesa Navas
Neteja sollicita Naviés.—Katanga (Kapiri).
ForRMICALEON Banks
Formicaleon harpalyce Banks. See above.
Formicaleon lethalis (Walker.—Kasai (Dima), Lake Leopold II, Katanga.
Formicaleon recurvus (Navads).—Kasai (Lodima).
Formicaleon scolius (Navés).—Lower Congo (Congo da Lemba).
Formicaleon turbidus (Navis). See above.
Formicaleon tholloni (Navads).—Kasai.
GRIALA Navas
Griala macilenta Navads,—Katanga (Kapiri).
Nemo.eon Navis
Nemoleon kitwanus (Kolbe).—Northeastern Congo (Kwidjwi).
, Macronemourus Costa
Macronemurus interruptus Kolbe.—Northeastern Congo (Buginda).
Macronemurus iolanthe Banks.—Belgian Congo.
Macronemurus lepidus Kolbe. See above.
28 Bulletin American Museum of Natural History [Vol. XLIII
Cyrmotuates Gerstecker
Africa, including Madagascar.
Cymothales congolensis Navis.—Katanga (Kapiri).
Cymothales delicatus Banks.—Congo (?).
Cymothales johnstoni Kirby. See above.
Cymothales liberiensis van der Weele.—Lower Congo (Lukula).
Cymothales mirabilis Gersteecker—Congo (?).
Mossa Navas
Africa. '
Mossa externa Navés.—Katanga (Bukama).
Mauta Navas
Africa.
Maula stigmatus Navés.—Katanga (Kalengwe).
Cueta Navas
Cueta cridai Navdés.—Kasai.
Cueta mysteriosa (Gersteecker.) See above.
Cueta punctatissima (Gerstzecker).—Katanga (Elisabethville),
Cueta styczynskii Navés.—Lower Congo (Boma). .
Nevro.eon Navas
Neuroleon alcidice (Banks) (=Creagris latens Navds).—Lower Congo (Boma,
Kitobola, Leopoldville), Katanga (Bukama).
Neuroleon filiformis (Gerstecker). See above.
Nyutus Navis
Nyutus lombardi Navaés.—Lower Congo (Matadi).
Banyvutus Navis
Banyulus acutus Navés.—Katanga (Katwamba, Kapiri, Lubumbashi River),
Banyutus insidiosus Navds.—Katanga (Katwamba, Kapiri),
Banyutus lethalis (Walker).—Katanga (Kasenga).
Banyutus maynei Navés.—Lower Congo (Malela).
Banyutus newer Navéis —Katanga (Katolo).
Nemopteride :
Nemopistaa Navis
Africa,
Nemopistha hennini Navés.—Belgian Congo; exact locality unknown,
1920) . Banks, Congo Neuroptera, Panorpata, and Trichoptera 29
Ascalaphide
AtLocormopes MacLachlan
Africa.
Allocormodes intractabilis (Walker).—Lower Congo (Lukula), [Kwango (Popa-
cabacca), Kasai (Kondue, Lodima).
TmesiBasis MacLachlan
Africa.
T mesibasis alberti Navis.—Katanga (Kasenga).
T'mesibasis waelbroecki van der Weele. See above.
Sunpatacsa Lefébvre
Australia, Malayan Archipelago, West Africa.
Suhpalacsa haullevillei (Navds).—Kasai (Dima).
Suhpalacsa recondita (Navds).—Lower Congo (Malela).
Suhpalacsa subcostalis (Navds).—Kasai (Dima).
Heticomitus MacLachlan
Africa, Southern Asia.
Helicomitus (?) bequaerti Navés.—Katanga (Sankisia).
Helicomitus festivus (Rambur). See above.
Nanomitus Navas
Africa.
Nanomitus sellatus Navis.—Katanga (Lukonzo).
DisPparomitus van der Weele
Africa. .
Disparomitus bacillus (Gersteecker). See above.
Disparomitus longus Navis.—Katanga (Kapiri).
Nacacta Navas
; Africa.
Nagacta leplaei Navés —Katanga (Kapiri).
Nagacta schoutedent Navds.—Katanga (Kapiri).
NeprHoneurA MacLachlan
Africa.
Nephoneura clavata Navdis.—Katanga (Lukonzolwa).
Encyoposis MacLachlan
Africa.
Encyoposis hemistigma van der Weele.—Katanga (Sankisia).
Encyoposis nigrostigma Navés.—Northeastern Congo (Gangara between Dungu
and Faradje).
30 Bulletin American Museum of Natural History — (Vol. XLIII
Puatascusa Kolbe
Africa.
Phalascusa vassei van der Weele.—Katanga (Kasenga, Elisabethville).
Dicotpus Gersteecker
Tropical Africa. oi
Dicolpus volucris Gersteecker—Manyema (Nyangwe).
Osmylide
Lysmus Navas
‘Lysmus leucomatodes Navis.—Congo (?).
Mantispide
Necyta Navas
Necyla cercata Navds.—Katanga (Mufungwa-Sampwe).
Necyla perparva (Esben—Petersen).—Lower Congo (Boma).
Mantispitta Enderlein
Mantispilla umbripennis Navés.—Katanga (Elisabethville).
Hemerobiidz
Nosysus Navis
Nosybus nobilis Navas.—Katanga (Bukama).
Chrysopidee
Curysopa Leach
Chrysopa bequaerti Navas. See above.
Chrysopa congrua Walker.—Lower Congo (Ganda Sundi).
Chrysopa ducissa Navdés.—Katanga (Kapiri).
NotHocurysa MacLachlan
Nothochrysa rufostigma (MacLachlan). See above.
Nothochrysa temerata Navdés.—Katanga (Kapiri).
PANORPATA
Bittacusidee
Birracus Latreille
1. Bittacus weelei Msben-Petersen
Bittacus weelei Esnun-Perensen, 1913, Rev. Zoolog, Afric., III, p. 142, figs. 7
and 8,
1920] Banks, Congo Neuroptera, Panorpata, and Trichoptera . 31
One specimen: Rutshuru, September 1914 (J. Bequaert Coll.).
Recorded from the Manyema (Kindu) and Katanga (Lubumbashi
River) —German East Africa.
PANORPATA RECORDED FROM THE BELGIAN CoNGO
Bittacusida2
Birtacus Latreille
Bittacus montanus van der Weele (B. schoutedeni Esben-Petersen).—North-
eastern Congo (between Beni and Lesse), Katanga (Mufungwa), Manyema (Kindu,
Vieux-Kassongo).
Bittacus pobeguini Navias.—Katanga (Kongolo).
Bittacus weelei Esben-Petersen. See above.
TRICHOPTERA
Polycentropide
Dipseupopsis Walker
1. Dipseudopsis fasciata Brauer
Dipseudopsis fasciata BRauER, 1875, Verh. Zool. Bot. Ges. Wien, XXV, p. 69,
Pl. rv, fig. 3.
Faradje, March and November 1912.
Recorded from various localities of the Belgian Congo: Lower Congo
(Leopoldville), island of the Congo River at the confluence of the
Ubangi River, Uele River.—Tropical Africa from the Senegal, Sudan
and Abyssinia to Rhodesia and German East Africa,
Hydropsychide
Hypropsycuopes Ulmer
2. Hydropsychodes sexfasciata Ulmer
Hydropsychodes seafasciata Utmer, 1904, Ark. f. Zool., I, p. 421, figs. 10-12.
Stanleyville, January to March 1915.
Recorded frém the Aruwimi River (Banalia).—Cameroon, Belgian
Congo.
ZETHALOPTERA Brauer
3. thaloptera dispar Brauer
Aithaloptera dispar BRAUER, 1875, Verh. Zool. Bot. Ges. Wien, XXV, p. 72, Pl
1v, figs. 4-4 e.
Matadi, June 24, 1909; Stanleyville, March 1915.
32 Bulletin American Museum of Natural History [Vol. XLII
Lower Congo (Boma, Leopoldville, Kinshasa, Kwamouth), Middle
Congo (Bolobo), Lake Albert Edward, Kwango (Popocabacca).—West
Africa: Senegal, Cameroon, French Congo, Anglo-Egyptian Sudan,
Belgian Congo.
PoLyMorpPHANIscus Walker
4. Polymorphaniscus bipunctatus (Brauer)
(Cstropsis bipunctata BRAvER, 1875, Verh. Zool. Bot. Ges. Wien, XXV, p. 73.
Bumba, May 15, 1915; Lisala, May 6, 1915.
Common in the Belgian Congo: Lower Congo (Kinshasa, Boma),
Middle Congo (Irebu), Kasai, Bangala, Aruwimi (Basoko).—Tropical
Africa from the Niger and Anglo-Egyptian Sudan to Natal.
PROTOMACRONEMA Ulmer
5. Protomacronema hyalinum Ulmer
Protomacronema hyalinum Ulmer, 1904, Zool. Anzeiger, XXVIII, p. 354, fig. 1.
Stanleyville, August 10, 1909.
Common in the Belgian Congo: Lower Congo (Boma, Kinshasa,
Leopoldville), Equator (Ikelemba, Baringa), Bangala (Umangi).—
French Congo.
TRICHOPTERA RECORDED FROM THE BELGIAN CoNGO
Hydroptilide
Caroxyetuira Ulmer
Catoxyethira fasciata Ulmer.—Lower Congo (Kinshasa).
Polycentropidz
NYCTIOPHYLAX Brauer
Nyctiophylax occidentalis Ulmer.—Lower Congo (Kinshasa).
Protopipseupopsis Ulmer
Protodipseudopsis sjastedti Ulmer.—Congo.
Dieseupopsis Walker
Dipseudopsis africana Ulmer.—Belgian Congo.
Dipseudopsis fasciata Brauer. See above.
Dipseudopsis lata Ulmer. Kasai (Dima),
Dipseudopsis schoutedeni Lestage.—Belgian Congo.
Dipseudopsis simplex Ulmer,—Belgian Congo,
59.57,34 (67.5)
. Article IV.—AFRICAN STONE-FLIES AND MAY-FLIES
COLLECTED BY THE AMERICAN MUSEUM
CONGO EXPEDITION!
By James G. NEEDHAM
Puate V
The few specimens of the above-named groups collected by the
Congo Expedition have proved of very great interest and are described
herewith.
PLECOPTERA
Two, or possibly three, species of stone-flies of the genus Neoperla
were taken by Messrs. Lang and Chapin at Faradje, Belgian Congo, 29°
40’ E., 3° 40’ N. The African species of this genus are inadequately
described. The first to be made known was Newman’s N. spio from Sierra
Leone (Newman, 1839, p. 86). Six others are briefly characterized in an
analytical key by Klapalek (1909 b, p. 218), four of them as new species,
without any description other than diagnostic characters (some of which
are mere color characters), without measurement, and without more
specific designation of localities. Two species of the Congo Expedition
collection are represented by both sexes and, in both, are so strongly
marked that it seems possible to identify them with two that are named
‘in Klapalek’s key. These two are more fully characterized herewith.
The third, somewhat larger, form (possible species) is represented by a
single female in bad condition, and is specifically unidentifiable.
NEoPERLA Needham
Pseudoperla Banks, 1892, Trans. Amer. Ent. Soc., XIX, p. 332 (preoccupied).
Neoperla Nerpuam, 1905, Proc. Ent. Soc. Washington, p. 108. Klapalek’ 1909,
Wien. Entom. Zeitg., XXVIII, p. 216.
Ochthopetina ENDERLEIN, 1909, Entom. Zeitg. Stettin, LXX, p. 324.
This genus includes all the African Plecoptera and is also fdund in
America, India, Indomalaya, and Japan. It is the dominant genus of
stone-flies in the tropics of the world.
Since so little is known of the African Plecoptera the following list of .
species described from the Ethiopian region may prove useful. The years
and pages refer to the appended bibliography.
1§cientific Results of the Congo Expedition. Entomology, No. 5.
35
36 Bulletin American Museum of Natural History {Vol. XLIII
Neoperla africana Klapalek, 1909 a, p. 56.—Cameroon (Johann bey ck
Kribi); German East Africa (Ukami, Langenburg).
Neoperla camerunensis (Enderlein), 1909 a, p. 343.—Cameroon (Barombi);
Belgian Congo (Kitobola).
Neoperla conradti (Enderlein), 1909 a, p. 335.—Cameroon (Barombi).
Neoperla didita (Enderlein), 1909 a, p. 345.—Cameroon (Barombi).
Neoperla dubia Klapalek, 1909 b, p. 218.—Africa; Belgian Congo (Faradje).
Neoperla excisa Klapalek, 1909 b, p. 218.—Africa; Belgian Congo (Faradje).
Neoperla laticollis Klapalek, 1909 b, p. 218.—Africa.
Neoperla leroiana Klapalek, 1911, p. 103.—Anglo-Egyptian Sudan (Redjaf).
Neoperla nigricauda Klapalek, 1909 a, p. 218. —Africa.
Neoperla sjéstedti Klapalek, 1909 a, p. 55.—Mt. Kilimanjaro (Kibonoto, 1300—
1900 m.); Mt. Ruwenzori (western slope, at 2000 m.).
Neoperla spio (Newman), 1839, p. 86.—Sierra Leone.
Neoperla tenera (Navas), 1915, p. 4.—Abyssinia (Endessa, Haut Aouache).
Neoperla transvaalensis (Enderlein), 1909 b, p. 402.—Transvaal (Zoutpansberg).
Neoperla excisa Klapalek
Plate V, Figures 13, 15 and 17
Length to wing tips 12 mm. Expanse 21 mm. in male. Color uniformly dull
yellowish over body, wings, and appendages, the tips of the last hardly darker in color,
the veins of the fore wings brownish. Ocelli large, close-set, these being separated
by a space that is less than half the diameter of one of them,
The distinctive characters of this species lie almost wholly in the secondary sexual
characters. In the male, segment ten of the abdomen is divided on the mid-dorsal
line, and the two anteriorly directed dorsal horns are slender, nearly smooth, approxi-
mated in their basal half and divergent toward the tips. Between the bases of these
horns, the free, flaplike, inner ends of the divided dorsum are thickly studded with
minute, roundish, brown, button-like, chitinous nodules, covering the convex upper
surface, one hundred or more on each flap. On the dorsum of the eighth segment a
conic-triangular, brown, heavily chitinized process arises under the tips of the paired
horns, and like them is directed forward. The tip of this median process is directed
toward a shallow notch in the emarginate chitinized apical ring of abdominal seg-
ment seven, and beside the emargination and beneath the tip of the process are a few
larger and darker chitinous nodules.
The female is similar to the male in coloration, slightly larger in size, and bears
no well-marked external sex characteristics, the apical margin of the eighth abdominal
segment on the ventral side is slightly emarginate toward the median line and more
strongly chitinized over a minute and very shallow median concavity in this margin.
The shell of the ovarian egg is marked by about eight to twelve straight, low longi-
tudinal ridges that divide the surface into areas like the staves on a barrel, that extend
over four-fifths the length of the egg, but that do not reach either the shell aperture
at the micropyle or the more pointed opposite end.
One male and one female. Faradje, Belgian Congo.
1920] Needham, African Stone-flies and May-flies 37
Neoperla dubia Klapalek
Plate V, Figures 12, 14 and 16
Length to tip of wings 13 mm. Expanse 23 mm. in male. Color brownish,
becoming yellowish below and on bases of all appendages. Legs yellowish, suffused
with brown on knees externally and on tarsi beyond the basal segment. Both fore
and hind wings smoky yellowish brown with dark brown veins.
Like the preceding species the critical diagnostic characters are found in the
genitalia. The horns on the dorsally cleft tenth abdominal segment are slender and
parallel beyond their converging bases. Like the horns, the internal flaps between
their bases are bare. The dorsum of the ninth segment is somewhat saddle-shaped,
with a raised, broadly rounded area each side, that is studded with some forty to
fifty setigerous punctures, bearing long divaricate bristles. At the rear of the
seventh segment rises a long flattened conical process that is directed backward. Its
tridentate tip lies between the tips of the paired horns.
In the female the apical margin of the eighth abdominal segment is slightly
produced in the mid-ventral line, in a minute, roundish, scale-like flap covering the
genital aperture, and the tip of this scale-like portion shows a more or less evident
median apical emargination.
The lower portion of the oviduct is densely clothed within by minute conic
prickles, whose points are directed outward. This chitin-lined portion is coiled
several turns, and has a length equal to that of several abdominal segments.
The shell of the egg of this species shows some thirty to forty very oblique
longitudinal strix, spirally wound about it, and reaching to its obtuse ends.
Several specimens of both sexes from Faradje, Belgian Congo.
PLECOPTERA RECORDED FROM THE BELGIAN CONGO
Perlide
NEoreRLA Needham
Neoperla camerunensis (Enderlein).—Lower Congo (Kitobola).
Neoperla dubia Klapalek. See above.
Neoperla excisa Klapalek. See above.
Neoperla sjéstedti Klapalek.—Western slope of Mt. Ruwenzori, at 2000 m.
Klapalek has also described three nymphs of Perlide taken in the
Belgian Congo (forest 90 kilom. west of the southern shore of Lake
Albert Edward) and Lestage has described and figured three additional
species of nymphs from the Maba River, near Mlonda (western shore
of Tanganyika). All these are undetermined species of Neoperla.
BIBLIOGRAPHY OF AFRICAN PLECOPTERA
ENDERLEIN, G. 1909 a. Plecopterologische Studien, II. Entom. Zeitg. Stettin, LXX,
pp. 324-352.
1909 b. Klassifikation der Plecopteren, sowie Diagnosen neuer Gattungen
und Arten. Zoolog. Anz., XXIV, pp. 385-419,
6. Bulletin American Museum of Natural History [Vol. XLIII
Kuapatex, F. 1909 a. Plecoptera. Zoolog. Ergebn. Sjéstedt Kilimandjaro Exp., IT,
14, 3, pp. 55-58.
1909 6. Vorldiufiger Bericht iiber exotische Plecopteren. Wien. Entom.
Zeitg., XXVIII, pp. 215-232.
1911. Neoperla'leroiana n. sp. Wien. Entom. Zeitg., XXX, pp. 103-104.
1912. Plecoptera. Wiss. Ergebn. Deutsch. Zentr. Afr. Exp. (1907-08),
III, pp. 447-452. ~
Lestace, J. A. 1917. Deuxiéme contribution 4 I’étude des larves des Ephéméres
et Perlides du Congo Belge. Rev. Zoolog. Afric., V, pp. 134-140.
NavdAs, L. 1912. Sur quelques Névroptéres d’ Afrique. Rev. Zoolog. Afric., I, pp. 401—
410.
1915. Neurépteros nuevos 0 poco conocidos, IV. Mem. Ac. Cienc.
Barcelona, (3) XI, No. 23, pp. 1-28.
NEwMaN, E. 1839. On the synonymy of the Perlites, together with brief characters
of the old, and of afewnewspecies. Mag. Nat. Hist., N.S., III, pp. 84-90.
EPHEMERIDA
A single male adult May-fly appeared among the pinned specimens
brought from the Belgian Congo by Messrs. Langand Chapin. However,
when a dilapidated and crumpled specimen of the stone-fly Neoperla
excisa was boiled for study, two additional minute specimens of another
May-fly were loosened from some place of concealment underneath the
stone-fly, and floated freely away from it in the water. These, thus
softened, were fit for study, and upon examination proved to be appar-
ently a nymph and a male imago of the same species. They represent
a new genus, as well—one closely allied to the cosmopolitan genus Cenis.
This is one of the smallest of May-flies. The other pinned specimen is
one of the largest of May-flies. It is the fine species for which Navas
proposed the generic name Eatonica. Its generic position has been dis-
cussed by Eaton, Nav&s, Ulmer, and Lestage without much agreement.
Lestage has summarized the evidence in the Revue Zoologique Africaine,
VI, 1918, pp. 82 to 89, and we follow him in calling this species Penta-
genia schoutedeni. The nymphs of all the genera in question are much
more strongly characterized than are the adults; but the nymph of this
species is as yet unknown. Both this species and the new one are
herewith described and illustrated.
Pentagenia schoutedeni (Navas)
Plate V, Figures 1 and 2
Length about 17 mm., tails 40 mm. additional. Expanse of wings 30 mm.
Color brownish, darker on dorsum and on all carinmw, paler beneath, Fore and
middle legs are lacking from the single specimen; the single hind leg present is pale
and concolorous, as are the setw. The middle tail, in the male, aborted to a few-
segmented rudiment. Claws very dissimilar, one of each tarsus being broadly
ala
a
1920] . Needham, African Stone-flies and May-flies 39
flabellate; the other, sharply uncinate. Venation as shown in Plate V, figure 1.
Wings opalescent and irridescent; fore wings strongly marked with a broad costal
band of brown. All veins tinged with a deeper shade of purplish brown, the trans-
verse veins about the base of both wings and a few others near the middle of the hind
wings more broadly margined with the same color.
Forceps of the male strong, the elongate middle segment curved almost in a
semicircle, the single small terminal joint twice as long as wide. Penes separated at
tip in less than half their length, in a V-shaped notch; each bearing a low subterminal
obtuse hook just below the aperture of the recurved sperm duct.
One male. Faradje, Belgian Congo. Lestage reports this species
as being distributed throughout equatorial Africa.
CZNOPsIs, new genus
Allied to Cenis. Tails three, middle one longest. Forceps of the male at least
three-jointed, the third joint very long and flexible. Claws dissimilar, one sharply
hooked, one blunt. Terminal tarsal segment as long as the three basal segments.
Wings two. Venation as shown in Plate V, figure 3, differing from Cenis in better
preservation of typical forks and in a wider band of cross-veins.
Cenopsis fugitans, new species
Plate V, Figures 3 to 11
Length 4mm., tails 9 mm. (middle one 10 mm.) additional. Expanse of wings 10
mm.
Color blackish, with pale setze and whitish wings.. Head blackish, paler in rear,
antenne pale. Thorax and abdomen black, the latter with sooty black patches on
sides of segments one to eight and on mid-dorsum of segments nine and ten. Tibi
each with a minute sooty patch just below the knee joint. Wings with smoky costal
band that is darkest along the subcostal vein. Male forceps wholly pale, basal seg-
ment not longer than wide; middle segment five times as long, cylindric, end seg-
ment longer than all the basal parts collectively, tapering and flexible in its terminal
portion. Penes fused into a single pyriform organ except at tip where a median cleft
remains, and surrounded beneath by a V-shaped ring of chitin at the apex of the tenth
abdominal segment, the arms of the V reaching laterally to the base of the forceps
each side. Fore tibia of the male three to four times as long as the other tibiz.
One specimen found as detailed above, with its nymph. Faradje,
Belgian Congo.
Nymph.—Length about 7 mm., antennz 2mm., sete broken, perhapsa little longer
than the antenne, width 2mm. Color apparently greenish black. Body depressed,
rather smooth with short legs and thin lateral edges to abdomen.
Head small and rather compact, with small rounded eyes capping its lateral
angles. Mouth-parts as shown in Plate V, figures 8 to 11. Prothorax wider than
the head with thin flaring anterolateral angles.
Abdomen with a conspicuous mid-dorsal hook on the second abdominal segment
and with thin flat lateral spines on segments four to nine, the series on each side
curving outward like a segment of a circular saw. Gills on segments one and three
40 Bulletin American Museum of Natural History [Vol. XLIII
to seven, those of segment one simple tapering filaments set erect upon a pedunculate
base, those on segment three elytroid and covering the others to rear, those of seg-
ments four to seven, thin, flat translucent, whitish plates, obliquely oval in form,
the margins fringed with long, forking, respiratory filaments.
One nymph, taken with the adult from the body of the stone-fly
Neoperla excisa, as noted above.
A complete summary of what has hitherto been published concerning African
May-flies will be found in a paper by J. A. Lestage entitled: “Les Ephéméres
d’Afrique: Notes critiques sur les espéces connues” in Revue Zoologique Africaine,
VI, 1918, pp. 65-114.
Puate V
Congo May-flies and Stone-flies.
Fig. 1. Wings of Pentagenia schoutedeni (Navas), male.
Fig. 2. Forceps and penes of the same.
Fig. 3. Wing of Cenopsis fugitans, new species, male.
Fig. 4. Forceps and penes of the same.
Fig. 5. Nymph of Cenopsis fugitans.
Fig. 6. Hind foot of same.
Fig. 7. Gill of first abdominal segment of same.
Fig. 8. Mandible of same.
Fig. 9. Maxilla of same.
Fig. 10. Hypopharynx of same.
Fig 11. Labium of same.
Fig. 12. Neoperla dubia Klapalek, mid-dorsal aspect of rear segments of the
abdomen in the male.
Fig. 13. Neoperla excisa Klapalek, showing corresponding parts.
Fig. 14. Egg-shell of Neoperla dubia, showing spiral striations.
Fig. 15. Egg-shell of Neoperla excisa, showing longitudinal ridges.
Fig. 16. Neoperla dubia, aspect of the eighth ventral segment of the female,
showing lamina.
Fig. 17. Neoperla excisa, showing the retuse corresponding border.
yo’
Buuuetin A. M. N. HA. Vor, XLIII, Puars V
‘
i, g X
» SP
Bc AS
CONT
1920] Banks, Congo Neuroptera, Panorpata, and Trichoptera 33
Psychomyidze
Ecnomvus MacLachlan
Ecnomus tropicus Ulmer.—Lower Congo (Kinshasa).
Ecnomus deceptor MacLachlan.—Lake Kivu.
Hydropsychide
Hypropsycuopes Ulmer
Hydropsychodes diminuta (Walker).—Lower Congo (Kinshasa)
Hydropsychodes albomaculata Ulmer.—Lower Congo (Kinshasa).
Hydropsychodes sexfasciata Ulmer. See above.
AETHALOPTERA Brauer
Aithaloptera dispar Brauer (=Primerenca maesi Navas). See above.
PotymorpuHaniscus Walker
Polymorphaniscus bipunctatus (Brauer.) See above. '
PuHanostoma Brauer
Phanostoma senegalense Brauer.—Lower Congo (Kinshasa), Bangala (Lisala).
ProTromacronema Ulmer
Protomacronema hyalinum Ulmer. See above.
Protomacronema pubescens Ulmer.—Lower Congo (Kinshasa), island of the Congo
River at the confluence of the Ubangi.
Macronema Pictet
Macronema capense Walker var. signatum Walker.—Ubangi.
Leptoceride
LepTrocervus Leach
Leptocerus trivittatus Ulmer.—Lower Congo (Kinshasa).
PsEUDOLEPTOCERUs Ulmer
Pseudoleptocerus squamosus (Ulmer).—Lower Congo (Kinshasa).
@cetis MacLachlan
(Ecetis fasciata Lestage.—Belgian Congo.
SETODELLINA Lestage’
Setodellina albopunctata Lestage.—Belgian Congo.
59.53, 72 (67.5)
Article V—ISOPODS COLLECTED BY THE AMERICAN
MUSEUM CONGO EXPEDITION!
By Witiarp G. VAN NAME
INTRODUCTION
Twenty-one species, representing three of the primary subdivisions
or superfamilies of the Isopoda, were collected by the Congo Expedi-
tion. Twelve of these species appear to be new to science. Of the total
number, nine are marine or are confined to the immediate vicinity of the
sea, one is a true fresh-water form, and the rest are terrestrial. Seven
of the aquatic species collected are parasitic on fishes or on other Crusta-
cea. This large proportion is, however, easily explained by the fact that
the isopods were largely obtained incidentally to the collection of other
animals and that the parasitic forms were thus more likely to be found
than free-living ones equally or even more abundant.
While the collection is interesting on account of the comparatively
little that is recorded concerning the Isopoda of the Congo region and
on account of the new forms contained in it, it is by no means a represen-
tative one and the species it comprises cannot be more than a small
percentage of those actually found there. Leaving out of account marine
forms, which are generally of more or less wide distribution, many land
isopods are known from other parts of tropical Africa, not too far distant
from the Congo region to make it probable that they may eventually
be found to range within its limits, especially as the comparative uni-
formity of wide stretches of country and the absence of mountain bar-
riers in Africa is favorable to a wide distribution of such animals. No
systematic collecting of Isopoda was done; nearly all the specimens were
obtained and preserved by Mr. Herbert Lang himself, in intervals
when his time and efforts were not occupied in obtaining or caring for
more important material; but the fact that even under such conditions
a larger collection was not brought back would seem to indicate that the
Isopoda do not constitute a very large or conspicuous part of the fauna.
From an economic point of view, the importance of the isopods in
the Congo region, as in most other parts of the world, is insignificant.
The terrestrial forms are apparently not harmful either from their abun-
dance or habits. The wood-boring aquatic species (Spheroma) was
ee Results of The American Museum Congo Expedition. General Invertebrate Zoology,
o. 4.
41
42 Bulletin American Museum of Natural History {Vol. XLIII
found only in mangrove roots and was not reported as attacking the
piling of wharves or bridges, as these animals do to a serious extent in
some places. Those parasitic on market fishes are undoubtedly harmful
to human interests; though they probably seldom kill the fish, except
perhaps very young or weak individuals, they must be very troublesome
to them, reducing them in flesh and food value. These parasites for-
tunately do not appear to be very abundant, as those collected were
found only by examining a large number of fishes. If they were numer-
ous, it would seem difficult to devise any way of combating such pests.
As food for larger animals, birds, and fishes, the isopods doubtless play
some part, but most of them would appear to contain little nourishment
and many of the terrestrial species are probably quite unpalatable.
LITERATURE
Though the literature concerning the isopods of South Africa, East
Africa, and some of the extreme western parts of the continent is rather
extensive, the published records and notices referring to these animals
in the Congo region are comparatively few. The more important of
them are, for the most part, contained in or cited in the general mono-
graphs of Schicedte and Meinert (1879-1884) and Hansen (1890); in
the works of Stebbing (1908, 1910) on South African Crustacea, as far
as the marine forms are concerned; and, for the land isopods, especially
in the monographs and reports of the late Prof. G. Budde-Lund (see
the bibliography at the end of this article). According to his published
statements, Budde-Lund had prepared descriptions and drawings of
many other previously unknown African species of land isopods, but the
publication of these was prevented by his untimely death. A number of
land isopods have been described from Togo Land by Hilgendorf (1893),°
from Assinie by Dollfus (1892), and from Liberia and Sierra Leone by
Richardson (1908), but these have not, as far as I am aware, been found
in the Congo region. The following are the species of isopods that I
have found reported from this region, taking as limits the west coast of
Africa from the latitude of Cape Verde to that of Cape Frio for the
marine forms, and for the terrestrial forms the Congo basin and the
colonies on the west coast from Cameroon to Portuguese West Africa
inclusive.
— we
— se ee ee
ln lis ih ie
1920] Van Name, Isopods of the Belgian Congo 43
Marine Isopoda
Aiga deshayesiana (Milne-Edwards), 1840.
Aiga deshayesiana Schicedte and Meinert, 1879, Naturhist. Tidsskr., “(3) XI,
p. 360, Pl. vii, figs. 7-9.
Rocinela deshayesiana Studer, 1884, Abh. k. Akad. Wiss. Berlin, 1883, phys.-
chem. Abt., Suppl., p. 22.
Azores and Cape Verde Islands; also Mediterranean.
Aga webbii (Guérin), 1836.
Aga webbii Schicedte and Meinert, 1879, Naturhist. Tidsskr., (3) XII, p. 347,
Pl. x, figs. 1-4.
Cape of Good Hope to Portugal.
Cirolana cranchii Leach, 1818.
Cirolana swainsonii Miers, 1881, Ann. Mag. Nat. Hist., (5) VIII, p. 370.
Cirolana cranchii Hansen, 1890, Danske Vidensk. Selsk. Skr., (6) V, p. 343,
Pl. 1m, figs. 3-31.
Senegal (Gorée Island); a widely distributed species.
Nerocila rhabdota Keelbel, 1879."
Nerocila rhabdota Kcelbel, 1879, Sitzungsber. Akad. Wiss. Wien, (math.-nat. K1.),
LXXVIII (Abt. 1), p. 409, Pl. 11, figs. 2a-2c.
Nerocila rhabdota Schicedte and Meinert, 1881, Naturhist. Tidsskr., (3) XIII,
p. 39, Pl. n, figs. 5, 6.
Senegal.
Nerocila cephalotes Schicedte and Meinert, 1881.!
Nerocila cephalotes Schicedte and Meinert, 1881, Naturhist. Tidsskr., (3) XIII,
pp. 8, 9, 60, Pl. rv, figs. 16-18.
Senegal (Gorée Island); Gaboon; Cape of Good Hope.
Anilocra capensis Milne-Edwards, 1840.
Anilocra capensis Schicedte and Meinert, 1881, Naturhist. Tidsskr., (3) XIII,
p. 146, Pl. x, figs. 4, 5.
Teneriffe to Java.
Glossobius linearis (Dana), 1853.
Glossobius linearis Schicedte and Meinert, 1883, . Naturhist. Tidsskr., (3) XIII,
p. 301, Pl. xu, figs. 1-9.
Widely distributed parasite of flying fishes; Cape Verde.
Glossobius laticauda (Milne-Edwards), 1840.
Glossobius laticauda Schicedte and Meinert, 1883, Naturhist. Tidsskr., (3)
XIII, p. 309, Pl. vir, figs. 10-16.
Widely distributed parasite of flying fishes; Cape Frio.
Cymothoa plebeia Schicedte and Meinert, 1884.'
Cymothoa plebeia Schicedte and Meinert, 1884, Naturhist. Tidsskr., (3) XIV,
p. 236, Pl. rx, figs. 1, 2.
Cape Verde.
Collected also by the American Museum Congo Expedition.
44 Bulletin American Museum of Natural History [Vol. XLIII
Terrestrial Isopoda
Ligyda gracilipes (Budde-Lund), 1885.
Ligia gracilipes Budde-Lund, 1885, Crust. Isop. Terrestr., p. 270.
Ligia gracilipes Dollfus, 1878, Bull. Soc. Zool. France, XXIII, p. 126.
Portuguese Congo (Landana), Senegal.
Eubelum stipulatum Budde-Lund, 1899.1
Eubelum stipulatum Budde-Lund, 1899, Rev. Crust. Isop. Terrestr., Part I, p. 71,
Pl. 1, figs. 1-16.
West Africa; Cameroon (Bonge).
Eubelum lubricum Budde-Lund, 1885.
Eubelum lubricum Budde-Lund, 1885, Crust. Isop. Terrestr., p. 392,
Eubelum lubricum Budde-Lund, 1899, Rev. Crust. Isop. Terrestr., Part I,
p. 72, Pl. 11, figs. 1-1.
Portuguese Congo (Landana, Chinchoxo).
Eubelum (Mesarmadillo) albicorne Budde-Lund, 1899.
[E.] Mesarmadillo albicornis Budde-Lund, 1899, Rev. Crust. Isop. Terrestr., Part
- I,p.79, Pl. m1, figs. 1-9.
Cameroon (N’dian).
Eubelum (Mesarmadillo) quadrimaculatum, Budde-LunP, 1899.
[E.] Mesarmadillo quadrimaculatus Budde-Lund, 1899, Rev. Crust. Isop.
Terrestr., Part I, p. 80, Pl. m1, figs. 13-17.
Cameroon.
Eubelum (Periscyphops) sileanum Budde-Lund, 1899.
[E.] Periscyphops silvanus Budde-Lund, 1899, Rev. Crust. Isop. Terrestr., Part I,
p. 82, Pl. m1, figs. 25-27.
Cameroon (Kitta, N’dian, Bibundi, Bonge).
Eubelum (Periscyphops) bizonatum Budde-Lund, 1899.
[E.] Periscyphops bizonatus Budde-Lund, 1899, Rev. Crust. Isop. Terrestr.,
Part I, p. 84, Pl. 1, figs. 18-24.
Cameroon (Kitta, Bibundi, Bonge).
Eubelum (Periscyphops) gibbosum Budde-Lund, 1899.
[E.] Periscyphops gibbosus Budde-Lund, 1899, Rev. Crust. Isop. Terrestr., Part
I, p. 85, Pl. v, figs. 23-25.
Cameroon (Bibundi).
Eubelum (Periscyphops) squamatum Budde-Lund, 1899.
[Z| Periscyphops squamatus Budde-Lund, 1899, Rev. Crust. Isop. Terrestr.,
Part I, p. 86, Pl. v, figs. 1-6. :
Cameroon (Bibundi).
Eubelum (Periscyphops) squamosum Budde-Lund, 1899.
(E.) Periscyphops squamosus Budde-Lund, 1899, Rev. Crust. Isop. Terrestr.,
Part I, p. 87, Pl. v, figs. 7-10.
Cameroon (Bibundi).
Synarmadillo albinotatus Budde-Lund, 1908.
Synarmadillo albinotatus, Budde-Lund, 1908, in Voeltzkow, Reise in Ostafrika,
II, p. 277, Pl. xim, fig. 47.
a
‘Collected also by the American Museum Congo Expedition,
—_— as oe a eee!
1920] Van Name, Isopods of the Belgian Congo 45
Synarmadillo cristifrons (Hilgendorf), 1893.
Periscyphis cristifrons Hilgendorf, 1893, Sitzungsber. Gesell. nat. Freunde,
Berlin, p. 152.
Synarmadillo cristifrons Budde-Lund, 1908, in Voeltzkow, Reise in Ostafrika,
p. 277, Pl. xi, fig. 48.
Cameroon (Kribi, Buea, Barombi Station).
Synarmadillo globus Budde-Lund, 1908.'
Synarmadillo globus Budde-Lund, 1908, in Voeltzkow, Reise in Ostafrika, ITI,
p. 276, Pl. xin, figs. 37-44.
Cameroon (Bibundi, Bonge).
Cubaris (Diploexochus) aculeata (Budde-Lund), 1885.
Armadillo aculeatus Budde-Lund, 1885, Crust. Isop. Terrestr., p. 289.
Diploexochus aculeatus Stebbing, 1910, Ann. South African Mus., VI, p. 446.
Portuguese Congo (Landana, Chinchoxo).
Niambia squamata (Budde-Lund), 1885.! .
Porcellio (Leptotrichus) squamatus, Budde-Lund, 1885, Crust. Isop. Terrestr.,
p. 196.
Niambia squamata Budde-Lund, 1909, in Schultze, Reise in Siidafrika, p. 60,
Pl. vi, figs. 1-3.
Portuguese Congo (Landana, Chinchoxo).
Rhyscotus globiceps Budde-Lund, 1908.
Rhyscotus globiceps, Budde-Lund, 1908, in Voelzkow, Reise in Ostafrika, IT,
p. 301, Pl. xvn, figs. 41-45.
French Congo (Loango).
Philoscia (Anchiphiloscia) cunningtoni Stebbing, 1908.
Anchiphiloscia cunningtoni Stebbing, 1908, Proc. Zool. Soc. London, p. 557,
Pl. xxvui, B.
North East Rhodesia (Niamkolo Bay, southern shore of Lake Tanganyika).
The following additional species are from localities but little beyond
the above limits.
Eubelum pila Budde-Lund, 1898.
Eubelum pila Budde-Lund, 1898, Deutsch Ost-Afr., IV, Land-Isopoden, p. 4.
Mt. Ruwenzori, on the western slope at 2600 m.
Eubelum hilgendorfii Budde-Lund, 1898.
Ewhelum hilgendorfii Budde-Lund, 1898, Deutsch Ost-Afr., 1V, Land-Isopoden,
p. 4, figs. 1, 2.
Mt. Ruwenzori, on the westérn slope at 200-3100 m.
Periscyphis nanus Budde-Lund, 1898.
Periscyphis nanus Budde-Lund, 1898, Deutsch Ost-Afr., IV, Land-Isopoden,
p. 5, fig. 6.
Mt. Ruwenzori, on the western slope at 3100 m.
Periscyphis pygmeus Budde-Lund, 1898. ‘
Periscyphis pygmeus Budde-Lund, 1898, Deutsch Ost-Afr., IV, Land-Isopoden,
p. 6, figs. 10, 11.
Mt. Ruwenzori, on the western slope at 3100 m.
1Collected also by the American Museum Congo Expedition.
46 Bulletin American Museum of Natural History (Vol. XLIII
Synarmadilloides roccatii Nobili, 1906.
Synarmadilloides roccatii Nobili, 1906, Boll. Mus. Zool. Anat. Torino, XXI,
No. 544, p. 2.
Eastern slope of Mt. Ruwenzori at 3000 m., Toro.
Porcellio (Porcellionides) pruinosus (Brandt), 1833.
Metoponorthus pruinosus Budde-Lund, 1898, Deutsch Ost-Afr., IV, Land-
Isopoden, p. 8.
Region of Mt. Ruwenzori. (This species is of almost world-wide distribution.)
Philoscia maculicornis Budde-Lund, 1898.
Philoscia maculicornis Budde-Lund, 1898, Deutsch Ost-Afr., IV, Land-Isopoden,
p. 9.
On the shore of Lake Albert, November 26, 1891. According to Stuhlmann’s diary
this locality is on the southwestern shore of the lake, in Belgian territory.
Philoscia mendica Budde-Lund, 1898.
Philoscia mendica Budde-Lund, 1898, Deutsch Ost-Afr., IV, Land-Isopoden,
p. 9, figs. 14, 15.
Region of Mt. Ruwenzori.
oe lt a
1920}
Van Name, Isopods of the Belgian Congo 47
Tue ConGco EXPEDITION COLLECTION
The following are the species comprised in the collection and
described and figured in the present article.
ae Se
Superfamily FLABELLOIDEA
PAGE
Pernt OOTIINONE, DOW GOOCIOE i oioas « viace eos oe akin lepers eee nape nsys 49
Nerocila cephalotes Schicedte and Meinert, 1881....................... 53
Nerocila rhabdota Koelbel, 1879.................000.45.. feahtetid wpe tcce, 57
Cymothoa plebeia Schicedte and Meinert, 1884. ............0.0.00.000055 59
Ichthyoxenos expansus, MEW SPECIES. ..... 266. c ene eee ene 60
Spheroma destructor Richardson, 1897...............0600 cece ee ee eens 63
Superfamily BopyrorpEa
Piemrdorynte angi, NOW BPeCHEO oe i ES ee 67
Paedione Chapint, NOW Species ....5 oe ee ee ee Senden bes 69
Superfamily OniscorpEA
UNE REIMAN SONU gl ge OE 5 yy gio owe gicle's woe Medea & KeaoeD 72
MMMM RIOT ANE CISERINOUS, LIBRE sie sik 5.6. scans eacy oo ow Ane ein dn ed hee ae wanes 77
Eubelum stanleyanum, new species... ...........0.. 0 cece cece eee 78
Eubelum stipulatum Budde-Lund, 1899... 2.20... 0. eee 81
Eubelum propinquum, new species.............6. 60. 84
Hubelign Geperive, NEw Species. 650)5 0. se ee be ee Recep sees 86 -
TNL CUPID: OW BOGCIOM 63) ok ins han ok ce eR hae ne bbe es 89
Hubelum tenebrarwm, New Species... 6. 6 ee cane eee e ewes 91
Synarmadillo globus Budde-Lund, 1908... ..............0.0 20220 e eae 92
Synarmadillo lubilensis, new species.............0.00 00 cece eee eee aee 95
Cubaris (Diploexochus) banana, new species. ..............0 0000 cece eee 97
Cubaris (Diploexochus) regulus, new species... .... 2.06.06. 0 0 eee eee 100
Niambia squamata (Budde-Lund), 1885... .........0.00 00020 ees 102
48 Bulletin American Museum of Natural History [Vol. XLIII
APPROXIMATE LOCATION OF PLACES WHERE SPECIMENS
WERE COLLECTED
Avakubi.—1° 20’ N., 27° 40’ E. Medje.—2° 25’ N., 27° 30’ E.
Banana.—6° S., 12° 20’ E. Poko.—3° 10’ N., 26° 50’ E.
Bengamisa.—1° N., 25° 10’ E. St. Antonio.—6° 10’ S., 12° 20’ E.
Garamba.—4° 10’ N., 29° 40’ E. Stanleyville—0° 30’ N., 25° 15’ E.
Lubila R.—1° N., 26° 30’ E. ~ Thysville.—5° 30’ S., 15° E.
Malela.—6° S., 12° 40’ E. Uele River.—3° 30’ N., 30° to 30° E.
Zambi.—6° S., 12° 50’ E.
The preponderance of parasitic forms among the aquatic species,
has been noted and an explanation offered. Naturally the majority of
new species are among the terrestrial forms; no less than five of them,
besides one previously described, belong to the typical subgenus of
Eubelum, a large genus particularly characteristic of tropical Africa,
its typical subgenus being confined to that region. An interesting and
unexpected discovery is that of a species assignable to the genus Ich-
thyoxenos, a group parasitic on fresh-water fishes, hitherto known only
from East Indian and Asiatic fresh waters. The fact that there is
considerable resemblance between the isopod faunas of Africa and the
West Indies and South America has been noted by previous writers (see
Budde-Lund, 1893, p. 3), though, considering the ease with which
such small creatures may be accidentally transported, it does not seem
that too much significance should be ascribed to the similarity. The
present collection, however, bears out this relationship in the case of
the marine and littoral forms, three of which appear to belong to species
found also on the American side, while three others have allies there
separable only on very slight characters.
In concluding these preliminary remarks, I wish to express my
thanks to the leader of the expedition, Mr. Herbert Lang, for his care
in preserving and labeling the specimens and to him and Dr. J. Bequaert
for information and corrections, particularly those relating to the geog-
raphy of the region; also to Mr. Roy W. Miner of the American Mu-
seum for his help and encouragement in the work of preparing this
article,
DESCRIPTIONS OF SPECIES
When possible, and except when otherwise stated or clearly in-
dicated from the nature of the characters under consideration, the
descriptions and illustrations have been prepared from female speci-
mens, even when the similarity of the two sexes made this apparently a
matter of indifference. Any differences between the sexes that were
eS a ee oe
. .
1920] Van Name, Isopods of the Belgian Congo 49
observed have been noted or illustrated. The specimens of the new
species chiefly used in preparing the descriptions and drawings have
been designated and catalogued as types, although in many cases they
have been badly damaged by the continued handling and the dissection
necessary in making out their characters. To prevent misunderstanding
it may be well to state that in the case of previously described species
the following descriptions and figures are based on Congo Expedition
specimens exclusively, and not partly on the descriptions of other authors.
The small number of groups and species dealt with makes the classi-
fication adopted here a matter of minor importance. That a very con-
servative course in the recognition of families and genera has been fol-
lowed is not due either to reactionary spirit or to any failure to recognize
most of the divisions established by recent authors as natural groups,
but to the belief that the splitting of genera and families has been carried
to an extent that, by magnifying the importance of small differences,
hinders the proper recognition of much more important points of re-
semblance and relationship, and that it has become the chief source of
the instability and inconvenience of our present system of nomenclature.
It has also often resulted in the necessity of defining the genera by such
minute characters or complex and arbitrary combinations of characters
that new species, or even some of the old ones, do not fit into them, and
the alternative is presented of indefinitely multiplying the genera or of
admitting exceptions and qualifications which break down the distinct-
ness of the groups. It seems as if such considerations should lead us to
accept the breaking up of well-established and natural groups with much
caution, and to keep in mind that the mere fact that distinct sections
within a group can be recognized, or that the group is a large one, are in
themselves no reasons for its complete disintegration, with the result of
usually compelling changes in the names of nearly all its members.
Superfamily CyMOTHOIDEA (= Flabellifera)
Cymothoide
Eurydicine (=Cirolanide auct. mult.; Eurydicide, Stebbing, 1905)
Eurydice carangis, new species
Text Figures 1 to 5
Body, as seen from above, of elongate-o-val outline, the widest part somewhat
anterior to the middle. The abdomen is rather short and considerably narrower
than the thorax, but the large epimera of the posterior thoracic segments fill in the
general outline so that it is nearly continuous. Back strongly arched; its surface
smooth. Length of largest specimens (to tip of abdomen) about 10 mm.; ratio of
greatest width to length about 6 to 14.5.
50 Bulletin American Museum of Natural History (Vol. XLIII
Héad narrow, and as seen from above deeply set back into the first thoracic
segment; its anterior border produced at the middle into a short and narrow rostrum,
each side of which the outline of the head is deeply concave to receive the broad,
flattened, basal segments of the first antenne that occupy nearly all the space
between the rostrum and the fore part of the eyes. The latter are large and deeply
pigmented, irregularly oblong, with about 50 ocelli. The rostrum joins, at its for-
Figs. 1 to 3. Eurydice carangis, new species.
1 and 2. Female, x 10.
3. Second pleopod of male, X 14,
ward extremity, the somewhat quadrate, enlarged end of the frontal lamina, that
extends up from below and forms the most anterior part of the head, First antenne
very long, exceeding the second pair and reaching, when well drawn back, beyond the
middle of the fourth thoracic segment. The fist two segments of the peduncle are,
as already stated, very wide and flattened; the first is so short that, as seen from
above, it appears merely as a border to the second, The third segment is narrow,
a fe ee ee
1920] Van Name, Isopods of the Belgian Congo 51
elongate, and cylindrical, and bears at its distal end what appears to be a small
vestigial fourth segment, incompletely united with the third. The flagellum is long
and slender, having twenty-five to twenty-eight articles, of which the proximal one
is usually much longer than any of the others. Second antennze much stouter than
the first, reaching, when well drawn back, to the posterior edge of the third thoracic
segment; the peduncle has four segments, the first of which is composed of two short
segments fused together; the flagellum is stout and tapering, with fifteen to eighteen
articles. The clypeus ends in a free, triangular, downwardly and forwardly extend-
ing apex. The extended lobe of the second segment of the maxilliped bears a single
hook.
Fig. 4. Eurydice carangis, new species. Mouth-parts of female, X 40.
The first, fifth, and sixth thoracic segments are longer than the others; the
epimera of all the thoracic segments are distinct and movably jointed to them
except in the case of the first. Their external surface is smooth and without any
furrow. The epimera end behind in an angle which is very slightly produced back-
ward in the anterior segments but much more so in the posterior ones. The four
last pairs of legs are provided with numerous spines and hairs.
The abdomen has six distinct segments; the first more or less covered by the
last segment of the thorax, and the others successively longer. Only the fourth and
fifth have the lateral angles produced and bent backwards. The terminal segment is
triangular, but rounded off at the extreme apex. A conspicuous depression on its
dorsal surface each side of the median line marks off a raised anterior border from
the comparatively flat posterior part. There is no distinct median carina. The
foliaceous parts of the pleopoda are rather elongate; the uropoda are short, with the
terminal branches somewhat triangular and obliquely truncated. Their inner
branch is wide and has a conspicuous notch on its outer border a little way from the
end; the outer branch is both shorter and narrower. There are moderately long hairs
on the inner and posterior margins of both branches of the uropoda and on the rounded
part of the tip of the abdomen, but none on the outer margins of the branches of the
uropoda.
52 Bulletin American Museum of Natural History [Vol. XLIII
Fig. 5. Eurydice carangis, new species. Legs of right side of female, external aspect, X 16.
Color, in alcobol, yellowish with many small irregularly stellate or branching
spots of black pigment on the upper parts.
The above description is from female specimens, but the sexual differences are
not conspicuous. The males have the peduncular segments of the second antennsw
averaging a little stouter than in the females. The process of the second pleopod of
the male is straight and shaped like a knife blade, tapering to a point from near the
middle of its length; it is shorter than the foliaceous branches of the appendage.
1920] Van Name, Isopods of the Belgian Congo 53
A majority of the numerous examples collected were obtained on
fishes, Caranx hippos (Linné), brought to market at Banana and St.
Antonio. Many of them are young individuals. Males predominate
among the adults. The young differ little in general appearance from
older examples except in size. The specimens in the collection are as
follows:
Cat. No. 3250. St. Antonio, August 1915. 1 specimen. Type.
Cat. No. 3253. St. Antonio, August 1915. About 50 specimens. Cotypes.
Cat. No. 3255. Banana, July 1915. 4 specimens.
Cat. No. 3252. Banana, August 1915. About 40 specimens. ‘‘Taken from dead
sea stars,”
This species, remarkable for the great length of the first antennz,
is one of several that are intermediate in character between Hurydice
Leach, 1815, and Cirolana Leach, 1818} the four-jointed peduncle of
the second antennze would place it in the former genus (the course
adopted here), while the absence of a right-angled bend in the basal
portion of the second antennz and the presence of a hook on the ex-
tended second lobe of the maxilliped would place it in Cirolana. If
placed in Cirolana, it would be a member of the subgenus FEzcirolana
Richardson (1913, Proc. U. 8. Nat. Mus., XLIII, p. 201). It is very
closely related to E. natalensis (Vanhoeffen, 1914, p. 500, fig. 42) from
Natal, but that form has shorter first antennez and lacks the notch on
the inner division of the uropoda. Such species as this render the dis-
tinction between the genera Cirolana and Eurydice difficult to maintain.
Cymothoine (= Cymothoide auct. mult.)
Nerocila cephalotes Schicedte and Meinert
Text Figures 6 to 9
Nerocila cephalotes! ScuimpTe AND Mernert, 1881, Naturhist. Tidsskr., (3) XIII,
pp. 8, 9, and 60, Pl. rv, figs. 16, 18. Srepsrne, 1910, Ann. South African Mus.,
VI, p. 423. Barnarp, 1914, Ann. South African Mus., X, p. 371.
The largest and apparently the oldest specimen of this species has adult female
characters, including a brood pouch; it measures 24.5 mm. long by about 11.4 mm.
wide. Body somewhat asymmetrical, widest at the junction of the sixth and seventh
thoracic segments; back moderately convex, the surface very hard and smooth.
Seven thoracic and six abdominal segments distinct and separate.
1The original description (loc. cit., p. 60) is headed N. cepholotes, but this obvious misprint need
be accorded no standing in nomenclature, as in the key and introductory remarks on preceding pages
(pp. 8 and 9) where some of the characters of this species are also given, the name is correctly spelled
cephalotes, as also in all other parts of the work except p. 60.
(Vol. XLITI
Museum of Natural History
Bulletin American
54
‘i
ee ee ee ee
1920] Van Name, Isopods of the Belgian Congo 55
Head small, gently convex in front and three-lobed behind;. eyes rudimentary,
being reduced to irregular pigmented areas with very imperfect lenses; first antennz
with eight, the second with ten segments, inclusive of the peduncular ones (some other
specimens have eleven segments).
The first, fifth, sixth, and seventh thoracic segments are the longest; the second,
third, and fourth are short. The posterior lateral angles of the first are very slightly
produced backward and rounded off; those of the sixth and seventh are considerably
produced and of angular form. The intermediate ones are not produced at all. The
above refers to the main part of the segments, not to the epimera. The epimera of
all but the first are distinctly separated from the segment; those of the second and
third segments are rounded off behind; the following ones are successively more and
more pointed. The legs have strong hooked claws; the last pair of legs, though long,
are slenderer than the others and have weaker claws.
The abdominal segments, except the last, are short; the first and second have
their lateral ends extended backward into long sharply pointed processes. The
terminal segment of the abdomen is very broad; it is shield-shaped, and ends in a
small, median triangular projection. It is nearly flat with only a slightly thickened
anterior margin and a slight suggestion of a median ridge. The outer ramus of the
uropoda may be described as broadly saber-shaped; it is slightly curved and about
equal to the last segment of the abdomen in actual length, but from its position it
extends beyond the end of the abdomen for about half its own length. The inner
ramus is less than two-thirds the length of the outer, but is broader. Neither the
uropoda nor the last segment of the abdomen are fringed with hairs.
Somewhat younger individuals may equal the specimen just described in length,
but are proportionately narrower, usually exhibit little or no lateral asymmetry, and
have the eyes less reduced and still provided with well-formed lenses.
Very much younger specimens (Figs. 8 and9) are quite different and would not, at
first sight, be regarded as of the same species. The smallest one in the collection
measures about 9 mm. long. The body is symmetrical and much narrower than in
the adult, but the head is proportionately larger and broader and provided with a
pair of very large, evidently perfectly functional eyes; the angles of the posterior
thoracic and anterior abdominal segments, as well as those of the epimera, are little
produced; the abdomen is less abruptly narrower than the thorax and its last seg-
ment is less broad, tapering more gradually to a sharp point at the end. All these
characters give the body as seen from above a smoother outline. The branches of
the uropoda are proportionately shorter and broader, and the inner pair (in very
young individuals also the end of the last abdominal segment) are fringed with hairs.
In some of these young and slender-bodied individuals male characters can be
recognized. The older, wide-bodied individuals, such as the one first described above,
are all females.
All the specimens have the under parts and legs yellowish; the upper parts in
the best preserved specimens are olive with two broad, yellowish, longitudinal stripes
separated by a rather wide, median, olive stripe; but many of them, perhaps because
of fading, are better described as mostly yellowish above with three olive stripes.
Schicedte and Meinert (1881) give the following distributioa for this
species: Gorée, Senegal; Gaboon; Cape of Good Hope; Cape Agulhas
(Cape Colony). Stebbing (1910) also records it from Cape Colony. The
[Vol. XLIII
n of Natural History
Bulletin American Museur
56
ng individuals, X 7.5.
id Meinert,
019. Nerocila cephalotes Schimdte ar
Figs, 8 ar
a a. ee
1920] Van Name, Isopods of the Belgian Congo 57
reader is referred to Schicedte and Meinert’s work for a more particular
description. They had the advantage of having both older (36.5 mm.
long) and younger (4.7 mm. long) specimens for study than those ob-
tained by the Congo Expedition, which are as follows:
Cat. Nos. 3234 and 3239. St. Antonio, August 1915. About 50 specimens.
Cat. No. 3248. Banana. July-August 1915. 5 specimens.
_ All were found on marine fishes (chiefly Caranx) brought to market.
They were adhering to various parts of the body or fins of the fish, some-
times more than one of the isopods on one fish.
Nerocila rhabdota Kcelbel
Text Figures 10 and 11
Nerocila rhabdota Ka@tBex, 1879, Sitzungsber. Akad. Wiss. Wien, (math.-nat. K1.),
LXXVIII, p. 409, Pl. 1, figs. 2a-2c. Scuiapte anp MEINERT, 1881, Naturh.
Tidsskr., (3) XIII, p. 39, Pl. 11, figs. 5,6. Barnarp, 1914, Ann. South African ©
Mus., X, p. 371. ‘
This species is readily distinguished from N. cephalotes by the lateral angles of
_ the main portion of the thoracic segments, which are produced backward (more
conspicuously in old than in younger individuals). Those of the more anterior seg-
ments are only slightly extended but the posterior ones are produced into conspicuous
triangular processes. These processes, and also the more elongate and pointed form
of the epimera, affect somewhat the general outline of the body when seen from above
and are, of course, still more conspicuous when seen from the side. Among other less
striking differences separating the two species are the more complete disappearance
of the eyes in adults of the present species, the larger and less angular median pro-
jection at the tip of the abdomen, the longer processes at the lateral ends of the first
two abdominal segments and the narrower inner branch of the uropoda. The general
resemblance between the two species, including size and color, is so close that it will
be unnecessary to give a more detailed description here.
This species also will be found carefully described and figured in
Schicedte and Meinert’s work. It was described by Keelbel from a
specimen taken from the pectoral fin of the fish Psettus sebe from the
coast of Senegal. Schicedte and Meinert’s description and figure are
based on a re-examination of Keelbel’s type, which measured 25 mm. long
and appears to have been a somewhat older individual than any of those
collected by the Congo Expedition. Schicedte and Meinert do not
appear to have had any other specimen. The species has recently been
reported from South Africa by Barnard (1914). Four individuals
(Cat. No. 3237) of this species were picked out from among the numer-
ous specimens of Nerocila cephalotes taken from market fishes (chiefly
Caranx) at St. Antonio, August 1915: The largest of them (Figs. 10
58
1920] Van Name, Isopods of the Belgian Congo 59
and 11) measures 22 mm. long. It has no brood-pouch but numerous
small, brown eggs are cemented to the posterior thoracic legs and some
of the pleopods and adjacent parts of the ventral regions of the body.
Another example, though not quite so large, has a well-developed brod-
pouch. The other two are not fully adult.
The smaller number of specimens obtained indicates that it is a
much less common species than N. cephalotes.
Nerocila acuminata Schicedte and Meinert, 1881, from the southern
United States and West Indian region, is a closely allied species.
Cymothoa plebeia Schicedte and Meinert
Text Figures 12 and 13
Cymothoa plebeia Scutapre AND Metnert, 1884, Naturhist. Tidsskr., (3) XIV, p.
236, Pl. rx, figs. 1, 2.
A single male, apparently fully fh. was collected by the Expedi-
tion. '
The body as seen from above is oblong, widening somewhat toward the rear,
though the last thoracic segment is a little narrower than the sixth. Abdomen
deeply set into the thorax, narrow in front, widening very rapidly to the large terminal
segment, which is but little narrower than the thorax. Seven thoracic and six ab-
dominal segments distinct and separate.
Head somewhat triangular, broader at the posterior end than it is long, with
sides that converge toward the front, which is rounded and bent downward in the
middle over the bases of the antenne. The latter are cylindrical; the first antennze
are much stouter and a little longer than the second, and arise widely separated from
each other; both pairs have eight segments. Rudimentary eyes are present in the
form of rather large, irregularly quadrilateral spots containing some dark gray or
blackish pigment.
The first segment of the thorax is not emarginate in front; it has no raised or
thickened lateral borders. The processes that extend forward each side of the head
are rather short (scarcely a third the total length of the head) and somewhat trian-
gular, with a rounded apex. No peculiarities were noted in the epimera or thoracic
limbs; the posterior four pairs have the basis keeled.
Last segment of abdomen large and broad; practically smooth above; its
posterior border nearly transverse, with rounded corners and a slight median notch.
The second pleopoda have a straight styliform process as long as, or slightly exceed-
ing, the foliaceous part of the appendage. The branches of the uropoda are small and
narrow and curved inward; they scarcely reach the end of the abdomen.
Size: 19 mm. long; maximum width, 8.3 mm. Color, in alcohol, yellowish, not
pigmented.
The specimen (Cat. No. 3263) was collected at Malela and, accord-
ing to the label, came from the inner side of the gills of a marine fish.
60. Bulletin American Museum of Natural History [Vol. XLIII
Figs. 12 and 13. Cymothea plebeia Schicedte and Meinert, 1884. Male, X 5.6.
Schicedte and Meinert describe this species from a single female from
Cape Verde, the male being unknown. Making the necessary allowance
for sexual difference, the agreement of the present specimen with their
species seems satisfactory.
Ichthyoxenos expansus, new species
Text Figures 14 and 15
The only specimen is a female which is nearly if not quite adult,
as the brood-pouch is large and well distended.
The body is symmetrical and broadly elliptical in outline. Its width slightly
exceeds two-thirds its total length, and the head and abdomen are deeply set into
the thorax; the back is only very moderately arched with a somewhat flattened
1920] .' Van Name, Isopods of the Belgian Congo 61
Fig. 14. Ichthyoxenos expansus new species. Female, X 7.5.
curvature; its surface is smooth and shining in spite of some minute irregular sculp-
turing visible only on magnification, Seven thoracic and six abdominal segments
are distinct and separate.
Head small, so deeply set back in the thorax that it projects but little beyond the
general outline of the body, its dorsal surface so inclined as to be directed forward
rather than upward. Seen from above, it appears triangular with the front rounded
off; from in front it is more nearly square. The eyes are fairly large, laterally placed
elliptical areas of blackish pigment. Antenne cylindrical; the first pair arise well
apart and have but seven segments, of which the basal one is somewhat swollen. The
62 Bulletin American Museum of Natural History [Vol. XLII
Fig. 15. Ichthyoxenos expansus, new species. Female, X 7.5.
second pair are slenderer and a little shorter, but likewise have seven segments.
Mandibular palps long and tapering and composed of three segments. Maxillipeds
without a foliaceous plate; they have palps of two flattened articles,
The thorax comprises by far the greater part of the body; the first segment is
very large, semicircular as seen from above, with its anterior margin hollowed {out
in the median part to receive the head. The lateral ends (directed forward and
slightly outward) are rounded off but not produced into projecting lobes. The re-
maining segments decrease successively in length (when measured on the mediam
line) except the fourth, which is shorter than would be expected from its place in the
series, The seventh is very short on the median line on account of the very deep
ae ee ee
ee ee ee Ce ee i a Tey
1920] Van Name, Isopods of the Belgian Congo 63
excavation of its posterior border to receive the abdomen, but it is quite wide at the
ends, which are directed almost backward and reach near to the anterior margin of
the terminal abdominal segment. Epimera distinct on all the thoracic segments
except the first; they are thick and crescent-shaped, but do not reach quite as far
back as the posterior corners of their segments. The latter are rounded off and not
produced. The legs are all much alike except for a gradual increase in length toward
the posterior end of the body. They are of the usual prehensile type, ending in
fairly large, curved claws. Their basal segments are not conspicuously keeled.
Abdomen short and rather narrow, especially anteriorly, set into the thorax
nearly to the base of its terminal segment, which is large and broad, rounded behind,
and longer than the other five abdominal segments taken together. These latter
increase in length from the first to the last; they are short longitudinally and ex-
tended transversely, their lateral parts being closely crowded together and not pro-
duced. The pleopoda are large and leaf-like, of ovate outline and smooth-edged.
The uropods are small; their branches, which do not reach as far back as the end of
the last segment, are rather narrow and of about equal length and rounded at the
tip. The outer one is slightly curved inward; the inner one is straight.
Total length of specimen, 16 mm.; width, 11 mm. Color, in alcohol, yellowish,
with a few faint and very minute dark pigment-spots on the head and fore part of the
first thoracic segment.
The single specimen (Cat. No. 3261. Type) was taken from the
gills of a characinid fish (Hugnathichthys eetveldii Boulenger, 1898)
about one foot long, collected at Poko on one of the tributaries of the
Uele River, July 1913.
Herklots (1870) described the type species of this genus, a parasite
on a river-fish of Java. Schicedte and Meinert (1884) added a second
species from the Himalayas, and Richardson (1913, Proc. U. 8. Nat.
Mus., XLV, pp. 559-562), a third from a Japanese fresh-water fish.
Another Japanese species has been added by Ishii (1916, Annot. Zool.
Japon., IX, pp. 125-131, 10 figs.). Except that the present form has the
first antennz a little dilated at the base, it agrees well with the generic
characters assigned by Schicedte and Meinert, though its widely separated
habitat may arouse a suspicion that we have here not a natural genus
but a case of convergence of species that have evolved separately from
related marine forms, perhaps those of the genus Livoneca, from which
Ichthyoxenos is distinguished by only very slight characters.
Spheromidee
Spheroma destructor Richardson
Text Figures 16 and 19
Spheroma destructor RicHarDsoN, 1897, Proc. Biol. Soc. Washington, XI, pp. 105-
'. 107; 1900, Amer. Naturalist, XXXIV, p. 223; 1901, Proce. U. S. Nat. Mus.,
XXIII, p. 534; 1905, Bull. U. S. Nat. Mus., No. 54, p. 282, figs. 294-298.
64 Bulletin American Museum of Natural History [Vol. XLIIT
Figs. 16 and 17. Spharoma destructor Richardson, 1897. Female, X 11.5.
(?) Spheroma terebrans Barn, 1866, Ann. Mag. Nat. Hist., (8) XVII, p. 28, Pl. 1, fig. 5.
Spheroma terebrans StessinG, 1904, Spolia Zeylanica, I, part 5, p. 16 (in part).
Hansen, 1905, Quart. Journ. Mier. Sci., XLIX, part 1, p. 116 (in part).
SrepsinG, 1908, Ann. South African Mus., VI, pp. 48, 49, 427 (in part); 1911,
Rec. Indian Mus, Calcutta, VI, pp. 181, 182 (in part).
The body is stout, short, and highly arched; in its general outline and in its
power of rolling into a ball superficially resembling some of the terrestrial isopods.
The dorsal surface is granular, on the posterior half of the body the granules are
coarse and the larger ones bear tufts of minute hairs to which mud adheres. The
thoracic segments, especially the fourth and fifth, have on the dorsal part a well-
roarked transverse ridge, and more or less well-developed paired tubercles are present
on the last one or two of the thoracic and on the abdominal segments, of which there
are but two in this genus. The posterior end of the abdomen is broad and obtuse; its
posterior lateral margins are conspicuously bent upward,
1920] Van Name, Isopods of the Belgian Congo 65
The first and second antenne have flagella with 8 or 9 and 12 or 13 articles
respectively.
The first legs have a short spine on the inner distal end of the propodus. It is
present in both sexes but is wanting on the other legs. All the legs, however, includ-
ing the first, have a short rounded extension of the posterior aspect of the propodus,
which overlaps the base of the dactylus:.
The number of teeth on the outside edge of the external branch of the uropoda
has been used as a distinguishing character in the species of this group. The Congo
specimens indicate that it is not reliable for such a purpose. They have from three
to five well-formed teeth in addition to the apical point. The proximal tooth is often
so reduced as to make it doubtful whether it should be counted or not, or it may
be poorly developed or wanting on one side only.
18 ~
Mm% 4
md MX,
Fig. 18. Spheroma destructor Richardson, 1897. Mouth-parts of female, x 24.
va
The drawings here reproduced are from a female. The males are, however,
closely similar in general form, size, and appearance, but, as a rule, have the tubercles
of the dorsal surface a little more prominent. Often there are fairly well developed
tubercles on the sixth as well as the seventh thoracic segment and one or two pairs of
small tubercles posterior to the central pair on the last abdominal segment. The
males also have the postero-lateral borders of that segment more broadly and con-
spicuously turned up than the females. This gives the posterior end of the body
a slightly narrower outline as seen from above.
A more detailed description can be found in Richardson, 1905.
The largest individuals measure 9.5 mm. to 10 mm. long to the tip of the ab-
domen. Their color varies from yellowish to a fairly dark brown or greenish brown,
the color being due to minute, irregularly branching pigment-spots.
About two hundred specimens (Cat. Nos. 3236 and 3245) were col-
lected in submerged mangrove roots in certain creeks near Banana,
where the water is quite strongly saline. Mr. Lang states that, though
66 Bulletin American Museum of Natural History ]Vol. XLUI
V VI
IV VII
Fig. 19. Sphwroma destructor Richardson, 1897, Legs of right side of female, external aspect,
x Ws.
1920] Van Name, Isopods of the Belgian Congo 67
abundant, its distribution was very local and that it was wanting from
many apparently favorable stations in the vicinity, also that their bur-
rows in the roots extended to, if not actually into, the sound and living
parts of the roots, appearing to cause their progressive death and decay.
I have been unable to find any material differences separating the
Congo Expedition specimens from Richardson’s S. destructor described
from Palatka, Florida, as far as the careful descriptions and figures
given by that authority enable me to judge, in spite of the fact that the
Florida specimens must have been from completely fresh water while
the Congo specimens, as is shown by their boring in mangrove roots,
lived in quite strongly saline water. Stebbing (1904) has reduced S.
destructor to a synonym of S. terebrans Bate, 1866, from Brazil. The
explanation he gives of the peculiarity ia the mandibular palpus in S.
erebransas shown by Bate’s figure is a very plausible one and, with this diffi-
culty removed, the reasons against accepting Stebbing’s view do not seem
at all conclusive. Nevertheless, as Miss Richardson’s species is based
ona full description and a number of careful figures, it would be neither
just nor safe to displace it until more is known about the Brazilian form.
Stebbing in the same article also reduces S. vastator Bate, 1899,
from India, Ceylon, and South Africa (see also Stebbing, 1908, p. 49)
to a synonym of S. terebrans. There are a few South African specimens
in the American Museum collection (Cat. No. 54) from the piling of a
bridge over the Isipingo River, Natal, which probably represent S.
vastator. A comparison of these with the Congo specimens shows the
resemblance to be very close. The tubercles of the back are slightly
more prominent, the last segment of the abdomen is a little narrower
and more angular in outline, and the legs are possibly a little stouter in
the Natal specimens, but the differences are of a very slight and relative
character and, without additional material, I am unable to decide how
much importance should be attributed to them. S. bigranulatum Budde-
Lund (1908, p. 304, Pl. xvm, fig. 55) from Zarizibar can hardly be a
synonym of S. vastator, unless considerable allowance is to be made for
inaccuracy in the figure of it published in that work.
Superfamily BOPYROIDEA (= Epicaridea)
Bopyridee
Pleurocrypta langi, new species
Text Figures 20 to 22
But one specimen of this species was obtained, a female probably
not fully adult, having a large but empty and not completely closed
brood-pouch.
68 Bulletin American Museum of Natural History [Vol. XLIIL
Body as seen from above very widely oval, the widest part far forward; the
anterior end very broadly rounded, the posterior end narrow. Outline conspicuously
asymmetrical, the convexity of the axis being toward the left side. Seven thoracic
and six abdominal segments are distinct and separate. Dorsal surface flat and smooth;
the lateral bosses (present on the four anterior body segments) are rather small.
Length of body to tip of abdomen, 4.8 mm.; width, 4 mm.
Head narrow, but little wider than long, deeply set back into the thorax; the
posterior border quite evenly curved, the anterior obtusely triangular with convex
sides and a minute rounded median projection or rostrum at the apex. Its anterior
border extends conspicuously out beyond the general outline of the front end of the
body. Rudiments of eyes in the form of elongate spots of pigment are present.
Antennz well developed, the first with three, the second with at least five distinguish-
able segments.
Figs. 20 to 22. Pleurocrypta langi, new species.
20 and 21. Dorsal and ventral aspects of female, X 12.
22. First opercular plate of left side, ventral aspect, X 9.
First thoracic segment very short on the median line, longer at the ends; the
third segment is the longest and from that point the thoracic and abdominal segments
diminish fairly regularly in length to the posterior end of the body, The epimera of
the first four body segments are narrow borders, only the fourth is of the full length
of the end of the segment. The epimera of the fifth, sixth, and seventh are succes-
sively wider and are as long as the segments; they end off squarely but have the
anterior and posterior lateral angles projecting a little; the seventh has a small addi-
tional tooth just anterior to the posterior lateral angle, (This description is of the
left side, the epimera of the right side are not well developed.) Legs of the prehensile
type, moderately long for an animal of this group; minute claws appear to be present,
Pe a ee ae
1920 Van Name, Isopods of the Belgian Congo 69
at least on some of them. The basis of these limbs has a dentate crest, and this and
other proximal segments of these limbs have a few irregular papilla-like projections
on the external aspect.
The first five abdominal segments have their lateral ends extended into tapering
flexible processes whose borders are coarsely and simply toothed or deeply sinuate;
these processes or extensions of the segments show evidence of incomplete segmenta-
tion in some of the narrow constrictions caused by the indentations. The pleopoda
of these segments consist of two branches each, their basal portion is a mere projec-
tion on the surface of the abdomen of irregular, somewhat papillated outline; the
branches are both similar in character and size to the above described lateral processes
of the segments. The sixth abdominal segment is very small and its lateral ends
bend directly backward. They are extended into processes similar to those of the
other segments and to the branches of the pleopoda. Uropoda, if not wanting, must
be represented by these extensions of the sixth segment, for all other appendages
appear to belong to the other five segments, though their crowded condition makes
this difficult to demonstrate.
Brood-pouch large, composed of five pairs of plates. Except the specially modi-
fied first pair (see Fig. 22), they end in tapering points which overlap but do not com-
pletely close the pouch. The last two pairs bear short hairs along the edges.
Many parts of the body are studded with small, irregular, rounded projections
or papille. They are present, as already mentioned, on the proximal segments of the
legs and occur also in abundance on the ridges marking the abdominal segments on
the ventral side of the body between the pleopoda, on the bases of the latter, and a
few even on the proximal parts of the exposed surfaces of some of the plates of the
brood-pouch, especially on the fifth pair.
Color, in alcohol, yellowish, without pigment-spots.
The only specimen (Cat. No. 3246) was found in the branchial
chamber of an Upogebia furcata (Aurivillius) collected at Banana, July
1915, one of the same lot in which Pseudione chapini, new species, was
found.
Only the female sex being represented, and that by a single in-
dividual which is perhaps not fully adult, the generic position of this
species is difficult to determine, though it seems to be an undescribed
form. Of the genus Pleurocrypta Hesse, 1865, in which I have included it
provisionally, Bonnier (1900, p. 310) says that the known species are
parasites of Anomala of the genera Galathea and Porcellana, and that two
undescribed species have been reported parasitic on Paguride. Several
members of the family Bopyride have been recorded as branchial
parasites of Upogebia. Three are enumerated by Nobili (1906) and
another is added by Hay (1917). The present collection adds two.
Pseudione chapini, new species
Text Figures 23 to 26
There are two well-preserved females in the collection, both similar
in size and characters.
70 Bulletin American Museum of Natural History [Vol. XLII
Figs. 23 to 26. Pseudione chapini, new species.
23 and 24. Dorsal and ventral aspects of female, X 9,
25 and 26, Dorsal and ventral aspects, of male, X 18.
Body as seen from above broadly oval in outline, narrowing rapidly but evenly
toward the posterior end, which is rounded, Both specimens strongly asymmetrical,
one with the axis convex toward the left, the other with it convex toward the right
side.
Dorsal surface flat and smooth; lateral bosses are developed on the first four
body segments. Seven thoracic segments and six abdominal segments are distinct
and separate.
RT ee ee ee ee ee a a ee
en a a Ne |
1920] Van Name, Isopods of the Belgian Congo 71
Head wider than long, deeply set back in the thorax; its anterior border is
without a raised or thickened edge and is slightly convex in outline, conforming to the
general oval curve of the body. No eyes; antennz very short, the first pair with
three, the second with four segments distinguishable.
First thoracic segment very short in the median part; the fourth is the longest;
the succeeding thoracic and abdominal segments diminish quite regularly in length
toward the posterior end. Lateral parts of the more posterior segments of the body
bent backward, the last two or three very strongly so. Epimera of first four thoracic
segments very distinct but narrow and shorter than the segment; those of the fifth,
sixth, and seventh are wider and as long as the segment, but so fused with it that the
line of union is more or less indistinct. The lateral ends of the thoracic and abdominal
segments and their epimera conform to the general body outline, being slightly ~
rounded or nearly square without produced angles.
Brood-pouch covering the entire lower part of the thorax but leaving the mouth-
parts exposed. Legs small and weak, particularly those of the anterior part of the
body. Basis of limbs not strongly keeled, but one or more of the proximal segments
of each limb bear on the outer aspect a number of small, irregular, papilla-like or
tooth-like projections quite conspicuous under moderate magnification.
The pleopoda each consist of two broad leaf-like divisions; these have the base
rounded, the edges smooth, and they end in an acuminate tip. Uropoda similar to
the pleopoda but consisting of one division only.
Length of largest female 7.1 mm.; width 5.2mm. Color in alcohol valichicialh
without pigment-spots.
A male individual was found clinging to the lateral ventral part of the abdomen
of one of the females. It is only 2.3 mm. long, of elliptical outline when seen from the
dorsal side, and little more than half as wide as it is long. It deviates very little in
shape from the usual isopod type seen in many of the terrestrial Isopoda and, except
that one side of the head is less developed than the other (the two antenne of that
side being wanting), the body is very nearly symmetrical. This peculiarity of the
head is doubtless merely an individual one, caused by some accidental injury.
The head is wide, its posterior margin nearly transverse; it bears on the dorsal
surface near the posterior margin a pair of minute, widely separated eye-spots. The
first antennz are very short and small; three segments are distinguishable in them;
the second antenne are considerably longer and have five segments, the last one end-
ing in a pair of spines or short stiff hairs.
All the seven thoracic segments are separate and do not vary very greatly in
length; the first is, however, somewhat longer and the seventh somewhat shorter
than the others. Five separate abdominal segments are readily distinguishable,
becoming very small! toward the posterior end of the body; their lateral extremities
are rounded and bent backward. This reduced number of abdominal segments
would place the species in the genus Parione, Richardson (1910, p. 39). The tip of
the abdomen is, however, in a more or less abortive condition in this family, and I
cannot regard the exact extent to which this process has gone as affording any evi-
dence of relationship or any secure basis for generic distinctions.
Thoracic limbs all well developed; the anterior ones are short and stout, of the
usual prehensile type and end in fairly long, slightly curved claws; the posterior
ones are similar but less stout, and the claws are much smaller and shorter. The five
pairs of abdominal limbs are reduced to fleshy lobes, which probably serve chiefly
as respiratory organs.
72 Bulletin American Museum of Natural History [Vol. XLIII
Color (in alcohol), yellowish.
Two adult and well-preserved females, to one of which the male was
clinging (Cat. No. of all the specimens 3247; the larger female is the
type) were obtained from the branchial chambers of specimens of Upo-
gebia furcata (Aurivillius), a thalassinid crustacean about one inch long
which burrows among the reots of mangroves, in July 1915, near Banana.
There was but one female in each host. Two more specimens of the host
in the same lot had bopyrid parasites (in one case one on each side of the
body) that were very likely of this same species but that were in too
- advanced a state of disintegration to make out much of their structure.
I have not been able to identify this with any previously described
form. It is closely allied to both the American species P. furcata and P.
curtata Richardson, 1904 (see Richardson, 1905), but is distinguishable
from the former by the absence of a raised margin on the front of the head
and by the wider pleopoda and from the latter by the form of the
abdominal segments in both sexes, also apparently from both by the
presence of papillze on the legs, which Richardson does not mention. But
some of these differences appear to be of slight importance, and the
validity of species so separated cannot be regarded as well established.
P. upogebie, Hay, 1917, from South Carolina, which, as its name shows,
is also parasitic on a Upogebia, differs in having the pleopoda provided
with digitate processes, as well as in other characters. Two other species
of this genus have been described as parasitic on European species of
Callianassa, a genus of burrowing Macrura allied to Upogebia. These
are Pseudione callianasse Kossmann, 1881, and P. dohrni Giard and
Bonnier, 1890. The latter differs in having the pleopoda with thickened
papillated margins; the former has been too incompletely described and
figured to be taken into consideration here.
Superfamily ONISCOIDEA
Ligydidz
Ligyda exotica (Roux)
Text Figures 27 to 30
Ligyda exotica Ricuarpson, 1905, Bull. U. 8. Nat. Museum, No. 54, p. 676, figs.
716-718; 1910, Marine Isop. Philippines, p. 41; 1910, Proc, U. 8. Nat. Mus.,
XXXVI, p. 125.
Ligia exotica Roux, 1828, Crust. Médit., p, 3, Pl. xi, fig. 9. Mitne-Epwarps,
1840, Hist. Nat. Crust., III, p. 157. Buppe-Lunp, 1879, Prospectus gen, spec.
Crust. Isop., p.°8; 1885, Crust, Isop. Terrestr., pp. 266-268. Do.urus, 1890,
Bull. Soc, Etud. Sci, Paris, XII, p. 7 (in part); 1898, Bull. Soc. Zool, France,
ee
1920] Van Name, Isopods of the Belgian Congo 73
XVIII, p. 189; 1893, Feuill. d. jeun. natur., XXIV, p. 24 (in part); 1897, idem,
XXVII, p. 211 (int part); 1898, Bull. Soc. Zool. France, XXIII, p. 126. BuppE-
Lunp, 1898, Deutsch Ost-afrika, IV, part 8, p. 10. Ricnarpson, 1899, Proc.
U.S. Nat. Mus., X XI, p. 866. Doturus, 1899, Proc. 4th Int. Zool. Congress,
Cambridge, pp. 255, 256, 260. Ricuarpson, 1899, Ann. Mag. Nat. Hist., (7)
IV, p. 335; 1900, Amer. Naturalist, XXXIV, p. 306. Sressina, 1900, in Willey,
Zool. Results, part 5, p. 646. Ricuarpson, 1901, Proc. U.S. Nat. Mus., XXIII,
p. 575; 1902, Trans. Conn. Acad. Sci., XI, pp. 306-308, Pl. XL, figs. 62a, 62b.
Goto, 1906, Annot. Zool. Jap., V, pp. 267-281. Do.uirus, 1907, Ergeb. Reise
Niederl. Ost-Indien, IV, p. 381. Buppr-Lunp, 1908, in Voeltzkow, Reise in
Ostafrika, p. 303; 1913, Trans. Linnean Soc. London, Zool., (2) XV, p. 391.
CuriTon, 1916, Mem. Ind. Mus. Calcutta, V, p. 462, figs. 1-22; Terao, 1915,
Dobuts. Zool. Tokyo, XXVII, p 47.
? Ligia grandis Perty, 1830-34, Delectus animalium articulatorum, p. 212, Pl. x1,
fig. 113.
Ligia gaudichaudii Mitne-Epwarps, 1840, Hist. Nat. Crust., III, p. 157. Nicouer,
1849, in Gay, Hist. Chile, III, p. 265. Dana, 1853, U.S. Exploring Exp., Crust.,
p. 741, xurx, figs. 6a—-6h.
? Ligia (italica) coriacea Kocu, 1835-44, Deutschlands Crust., p. 36; Berichtig.,
p. 211.
? Ligia baudiniana Mirrs, 1877, (not Milne-Edwards, 1840), Proc. Zool. Soc. London,
p. 670.
Ligia malleata Prerrer, 1889, Jahrb. Hamburg. Wiss. Anst., VI, p. 36.
This widely distributed form is too well known to require an extended descrip-
tion here. Exclusive of the long slender uropoda, the body may reach a length of
30 mm. but none of the Congo Expedition specimens exceed 24 mm. in length in the
case of male individuals, or 21 mm. in the case of females. In the alcoholic specimens
the ground color is a light greenish yellow, which under moderate magnification is
seen to be thickly dotted with minute, irregularly branched spots of dark pigment.
These, becoming more thickly distributed on the dorsal parts of the body, give the
animal a mottled greenish gray or slate color, which is, however, paler on the lateral
parts of the epimera, so that the body has a lighter border along the sides.
Among the principal specific characters are the form of the body, which widens
rapidly in the anterior part of the thorax and then tapers gradually to the last seg-
ment of the abdomen; the soft integument and the loose articulation and fragility
of the segments of the body and their appendages (few specimens reach the museum
with the uropoda and antennz attached and unbroken); the large epimera separated
from the body of the segment by a distinct suture; the large bulging eyes; very long
antennse and uropoda; and the distinct, though obtuse, angle at the tip of the ab-
domen. The males have on the distal end of the first leg a small flattened lateral
process which overlaps the base of the dactylus on the posterior aspect of the limb.
The inner edges of the carpus and merus of this limb are devoid of spines and are,
roughened with minute file-like oblique ridges. In the female the corresponding
limb is more slender and lacks the lateral process and the ridges, but is provided with
a number of spines in the position of the latter. The dorsal surface of the body is.
more conspicuously granulated than in L. olfersii. ;
1920] Van Name, Isopods of the Belgian Congo 75
A large male specimen, 24 mm. long, has thirty-one articles in the flagellum of the
second antenne. These reach a little way beyond the posterior end of the abdomen
when well drawn back. In another male, 21 mm. long (36.3 mm. long inclusive of the
uropoda), the flagellum of the second antennz has thirty-three articles. The terminal
branches of the uropoda are 9.8 mm. long. In one of the larger females (length, 18
mm.; inclusive of uropoda, 29 mm.) the flagellum of the second antennz has thirty-
three articles. The antennz reach a little beyond the end of the abdomen when well
drawn back, and the terminal branches of the uropoda are 7.5 mm. long. In another
female (length, 17.5 mm.; inclusive of the uropoda, 28 mm.) the terminal branches
of the uropoda are only 6.3 mm. long.
28 29 30
Figs. 28 to 30. Ligyda exotica (Roux), 1828.
28. First leg of right side of female, internal aspect, « 15.
29. Tip of inner division of right maxilliped of male, 15.
30. First leg of right side of male, internal aspect, X 12.
For a more detailed description of this species, with numerous figures
and a discussion of its relationships, see Chilton, 1916.
This isopod is found creeping or running on moist rocks, piles of
wharves, bridges, etc., or hiding in their crevices just above the water on
the coasts of most of the tropical and warmer regions of the world. Its
wide distribution may be in part due to its accidental dispersal through
commerce. It has already been reported from Senegal (Dollfus, 1897)
and from the eastern coast of Africa. Over fifty specimens (Cat. Nos.
3235 and 3238) are contained in the Congo Expedition collection, all
obtained along the shore at Banana from stones and rocks near brackish
water in August 1915.
Bulletin American Museum of N atural’ History
im ey,
ape
Fig. 31. Ligyda olfersii (Brandt), 1833.9 Female , X 6,7.
[Vol. XLITT
1920] Van Name, Isopods of the Belgian Congo 77
Ligyda olfersii (Brandt)
Text Figures 31 to 34 . ;
Ligia olfersii BRANDT, 1833, Bull. Soc. Imp. Nat. Moscou, VI, p. 11. Buppg-Lunp,
1885, Crust. Isop. Terrestr., p. 268. Ricuarpson, 1901, Proc. U. 8. Nat. Mu-
seum, XXIII, p..575. Curiron, 1916, Mem. Ind. Mus. Calcutta, V, p. 466.
Ligia exotica Dotirus, 1893, Feuill. d. jeunes natural., XXIV, p. 25 (in part); 1897,
idem, XXVII, p. 212 (in part).
Ligyda olfersti Ricnarvson, 1905, Bull. U.S. Nat. Mus., No. 54, p. 674, figs. 714, 715.
Resembles L. exotica in color and appearance but is somewhat smaller. The body
is proportionately narrower, especially in its anterior portion, so that the outline
seen from above is more nearly elliptical than oval. The surface is smoother, the
34
32 33 iS
A
D>
Figs. 32 to 34. Ligyda olfersii (Brandt), 1833.
32. First lef of right side of female, internal aspect, X 12.
33. Inner division of right maxilliped of male, X 24.
34. First leg of right side of male, internal aspect, X 14.
granulation being very faint and inconspicuous when the specimen is wet; the eyes
are more elongated and less bulging, and the epimera much more completely fused
with the main parts of the segments, while the body as a whole is firmer, less fragile
and more compactly articulated than in L. exotica. The propodus of the first pair of
legs of the male lacks the lateral process at the distal end that is present in L. exotica,
but in the male the file-like ridges on the merus and carpus, and in the female the
spines, are present as in the corresponding limbs of that species.
Females 17 mm. to 17.5 mm. long when measured to the tip of the abdomen,
have twenty-nine or thirty articles in the flagellum of the second antennx, which,
when drawn well back, reach half-way along the last abdominal segment.
A male 17 mm. long when measured to the tip of the abdomen (29 mm. long to
the ends of the uropoda) has thirty-five articles in the flagellum of the second anten-
nz, which reach, when well drawn back, half-way along the last abdominal segment.
The terminal branches of the uropoda are about 7.5 mm. long.
78 Bulletin American Museum of Natural History [Vol. XLII
This species is credited by Richardson (1905) with the following
distribution: Key West and Puntarasa, Florida; St. Thomas, W. I.;
and Brazil. There are specimens in the American Museum collections
from Andros Id., Bahamas (Cat. No. 3122) and one from Guadeloupe,
W. I. (Cat. No. 3123). With the material available I failed to discover
reasons for considering the Congo Expedition specimens distinct from
the American ones.
The Congo specimens (Cat. Nos. 3240 and 3241), fourteen in number,
were all obtained at Banana in August 1915, with L. exotica, and were
picked out from among the lots of the latter species, from which the
collectors evidently did not differentiate them.
Budde-Lund, 1885, described a species L. gracilipes from Senegal,
which, however, is smaller and has very long uropoda, the latter exceed-
ing the body in length.
Oniscide
Eubeline (= Eubelidse auct. mult.)
The next five species all belong to the typical subgenus of Hubelum.
Eubelum stanleyanum, new species
Text Figures 35 to 45
Body convex, contractile into a ball, oblong in outline when seen from above,
with the sides nearly parallel except for a slight convergence toward the rear; it is of
moderate width, broadly rounded in front, somewhat more narrowly so at the rear.
Surface smooth, with thickly distributed minute pits, visible on magnification, but
without granulation and without more than a slight trace of pubescence. Color of
upper parts varying from gray to brownish with yellowish markings and margins
on the segments. Some of the specimens have the epimeral parts of the segments,
and most of them have the exposed parts of the uropoda, yellowish in more or less
contrast to the adjacent darker parts. Under parts and legs yellowish. Size of large
females about 11 mm. long; males about 9 mm.
Head of moderate width, the epistome with a rather prominent, strongly up-
turned superior margin; the anterior outline of the head seen from above is gently
arched. Eyes rather large, with about seventeen ocelli, Antenne short and rather
stout; their first segment small and very short, their fifth segment not greatly
longer than the fourth; the flagellum is rather indistinctly three-segmented, having
the first and third articles about equal and considerably shorter than the second.
First maxilla with eight long sharp teeth (3-4-5), with an additional, very small,
accessory tooth beside the second tooth on the outer division, and a row of about
eleven plumose processes on the inner division, These processes are so closely
crowded that an exact count is difficult.
Van Name, Isopods of the Belgian Congo
Figs.
35.
35 to 44. Eubelum stanleyanum, new species.
Female, X 8.4.
Pleopoda of female, X 8.4.
Pleopods of male, X 10.
Antenna and outline of anterior segments, ventral view, X 9.6.
. Outline of head, front view, X 9
Tip of inner division of first maxilla, x 100.
. End of abdomen, posterior view, X 16.
Inner division of maxilliped, 36.
First maxilla, < 40.
Second maxilla, X 36.
79
80 Bulletin American Museum of Natural History [Vol. XLII
Posterior lateral angles of first thoracic segment little produced and rather
broadly truncated in an oblique direction, with a very short cleft to receive the
second segment when the body is rolled up. Lateral margin of first segment when seen
from the side very gently curved; it has a wide thick projecting border extending
back to the truncated part of the posterior angle. Along the whole length of this
border there is a narrow sulcus on the external aspect, but so situated that it is visible
in a ventral view also. The whole projecting border is sharply separated from the
main part of the segment by a deep but rather widely open groove. The inner side
of the cleft for receiving the second segment is more produced in an obliquely down-
ward direction than the outer side (though hardly extending as far in a directly
posterior direction), and is narrowly rounded off instead of truncated. The second
and third thoracic segments have a thickening of the anterior border of the epimeral
part, as though caused by an infolding of the edge, but they bear no process. The
lateral ends of the second to fifth (inclusive) segments are somewhat rounded off;
the sixth and seventh are more squarely cut off. Legs fairly long and moderately
stout, with stout but not very numerous spines.
The form of the pleopoda is shown in the accompanying illustrations. In the
female the first two pairs are more conspicuously swollen and tracheate than the
others. The terminal segment of the abdomen has the median portion very wide and
slightly tapering; the end is broadly and almost squarely cut off. The exposed part
of the uropoda is obliquely quadrangular and somewhat elongate, its external surface
only moderately convex. A large, shallow, transversely oval depression occupies
much of the distal half of this surface. In, but near the margin of, this depression
(well removed from both the internal and terminal margin of the basal segment)
the very minute, short, rounded rudiment of the external branch is placed. The
internal branches are rather narrowly tapered and do not reach to the end of the last
abdominal segment.
This species would appear to be common near Stanleyville, as the
following specimens were collected. Females predominate among them;
some have brood-pouches containing, young.
1920] Van Name, Isopods of the Belgian Congo 81
Cat. No. 3267. Stanleyville, Aug. 29, 1909. 6 specimens.
Cat. No. 3257. Stanleyville, March 1915. 2 specimens. Female is type and male
used in drawing figure.
Cat. No. 3244. Stanleyville, March 1915. 31 specimens.
Cat. No. 3258. Stanleyville, Aug. 10, 1909. 7 specimens.
Cat. No. 3300. Stanleyville, Aug. 4, 1909. 1 specimen from the stomach of a toad
(Bufo regularis).
Cat. No. 3265. Bengamisa. Sept. 1914. 1 specimen.
Eubelum stipulatum Budde-Lund
Text Figures 46 to 55
Eubelum stipulatum Buppn-Lunp, 1899, Revision of Crustacea Isop. Terrestr.,
(part 1), p. 71, Pl. 1, figs. 1-16.
Body convex, contractile into a ball; in a dorsal view not greatly different from
E, stanleyanum, though the epimera are rather more squarely and evenly cut off at
' the ends. The body surface is smooth, even, and shining, without granulation but
with scattered extremely minute pits visible only on considerable magnification.
Color grayish brown above with rather obscure yellowish markings and margins on
the segments; basal segments of the uropoda and the under parts and legs yellowish.
Length of specimens: female, about 11 mm.; two males, about 9.5 mm. and 10.6
mm., respectively.
Head with its anterior outline evenly convex as seen from above; the superior
margin of the epistome forms only a very narrow projecting border but this is con-
tinuous across the front. Seen from in front, this border slopes up gradually from
the sides to the middle, where it forms a very obtuse and gently rounded off angle.
Eyes well developed, with 22 to 25 ocelli. Antennz of moderate length and stoutness;
their fourth segment is less than three-fourths the length of the fifth and the flagellum
is distinctly three-segmented, the first article being the shortest, the second slightly
the longest. First maxilla with nine teeth (3+5 with a slender accessory tooth beside
the second) on the outer division, and a row of about eleven closely crowded
plumose processes on the inner division.
Posterior lateral angles of first segment somewhat produced backward, slightly
truncate in an oblique direction at the apex, which is cleft to receive the second
segment when the body is rolled up. The lateral margin of the first segment when
seen from one side is evenly curved, with a wide thick projecting border extending
along its length as far as the truncated part of the posterior angle. This border has a
moderately wide and deep sulcus along its whole length; the sulcus is situated on its
external aspect but is narrowly visible in a ventral view also. The whole projecting
border is separated from the main part of the segment by a very deeply impressed
groove. The inner side of the cleft posterior lateral angle is a little more produced
ventrally (though scarcely more posteriorly) than the outer and, instead of being
truncated, is rather sharply rounded off. Processes wanting on the inner surface of
the lateral ends of the second and following segments; there is scarcely any thicken-
ing of their anterior margin. Legs of moderate length and stoutness, with rather few
spines.
82
Bulletin American Museum of Natural History
49
(Vol. XLIII
Figs, 46 to 54. Bubelum stipulatum Budde-Lund, 1899,
46, Pleopoda of female, X 10.5.
47. Antenna and outline of anterior segments, ventral view, X 10.5.
48. Pleopoda of male, X 13,2.
49, Outline of head, front view, X 9.
50. End of abdomen, posterior view, X 16,
51. Inner division of maxilliped, X 34.
52. Second maxilla, * 34,
53, Tip of inner division of first maxilla, & 85,
54. First maxilla, X 34,
4
1920] Van Name, Isopods of the Belgian Congo 83
The form of the pleopoda is shown in the accompanying figures; in the female
the first two pairs are much more swollen and tracheate than the others. The terminal
segment of the abdomen is broad and tapering, with concave sides; its end is broadly
truncated, with a slightly convex outline and the corners a little rounded off. Uro-
poda with the exposed external surface very conspicuously convex; their outline is
somewhat oval. The small external branch is borne in a notch lying chiefly on the
external surface close to the terminal margin, which is a little indented. There is a
fairly well-marked, shallow depression on the external surface between this notch
and the outer margin. The inner branches do not reach to the end of the terminal
segment of the abdomen.
Specimens collected:
Cat. No. 3264. Stanleyville, February 1915, one female.
Cat. No. 3262. Bengamisa, September 1914, one male.
Cat. No. 3302. Stanleyville, one male, found in the stomach of a toad (Bufo regu-
laris) taken Aug. 4, 1909.
Fig. 55. Eubelum stipulatum Budde-Lund, 1899. Female, X 10.5.
Budde-Lund described this species from Bonge in Cameroon. His
specimens were somewhat larger, 13 mm. to 15 mm. long, and had,
according to his description and figures, more numerous ocelli (twenty-
eight to thirty) and more plumose processes (thirteen) on the inner
division of the first maxilla than the present examples, but otherwise the
correspondence is so close that, in spite of the difference of locality,.I am
unwilling to describe the Congo Expedition specimens as new, especially
as they may not have reached their full growth and size. Possibly they
may eventually prove to be distinct, but the information and material
now available do not appear sufficient to furnish satisfactory distinguish-
ing characters.
This form is closely allied to E. instrenuum Lénnberg and Budde-
Lund, 1912, from British East Africa, but in that species the margin of the
first thoracic segment is described as scarcely sulcated (“vix sulcato’’).
84 Bulletin American Museum of Natural History [Vol. XLITI
Another nearly allied form is E. lubricum Budde-Lund, 1885, from Land-
ana and Chinchoxo (Portuguese Congo), but that species is described as
having the superior outline of the epistome partly effaced in the middle
part of the head. (See also the next species.)
Eubelum propinquum, new species
Text Figures 56 to 62
A rather large male specimen (Cat. No. 3271), 13 mm. long, of a
slaty gray color above with rather conspicuous yellowish markings and
margins on the segments, appears to differ specifically from the form
just described under the name FE. stipulatum, though exceedingly closely
allied to it.
The body is perhaps a little more stout and convex; the surface is exceedingly
smooth, considerable magnification disclosing neither granulation nor pubescence,
but the surface is dotted with minute pits or punctures abundantly and quite evenly
distributed.
The epistome has its upper border complete across the front of the head; it is
but little projecting and is rather gently arched; the forehead is rather low; the eyes
are large and prominent, with about twenty-five ocelli. The antennz are missing in
the specimen here described. (Inaspecimen from Medje, Cat. No. 3249, presumably
of this same species, they are moderately long and rather slender, their fifth segment
rather more than one and one-quarter times the length of the fourth; the flagellum
nearly equal to the fourth segment and composed of three articles, the first the
shortest of the three, the second not much longer than the terminal one.) The outer
division of the first maxilla has nine teeth (3+-5 with an additional slender accessory
tooth beside the second); the inner division rs eleven or more closely crowded
plumose processes.
The first segment of the thorax has its alshion outer angles a trifle more pro-
duced backward than in the case of Z. stipulatum, the outer side of the cleft ends in
sharper posterior angle and the inner side is a little more produced backward than in
that species. In a lateral view the lateral inferior outline of the segment is straighter
(less convex downward); the thickened border is marked off from the main part of
the segment by a narrower groove and is itself more conspicuously narrowed toward
the rear; but these differences are slight and only noticeable on careful comparison,
The legs are also more spiny than in that species; on the inferior aspect of the anterior
pairs of legs the spines are so thickly distributed as to suggest a brush with short
thick bristles.
The abdomen and its appendages are also very similar to those of 2. stipulatum,
In the present form, however, the last segment of the abdomen is longer and narrower,
with the sides of the central extension parallel instead of converging toward the end.
The first pair of pleopoda are proportionately longer. The basal segments of the
uropoda are more elongate and the rudimentary external branches also more elongate
and proportionately longer and more conspicuous, and the oval depression on the
external surface of the basal segment is larger and more elongate,
1920]
Van Name, Isopods of the Belgian Congo 85
The above specimen is labeled ‘‘Thysville, June 2, 1915, from a
mushroom-shaped termite structure.’ The hard integument and bright
colors of this species show that it cannot be an inhabitant of the interior
of termites’ nests and, if the label is correct, it was probably merely
taking refuge in some external crevice of the nest.
Figs. 56 to 62. Eubelum propinquum, new species.
56.
57.
Pleopoda of male, X 9.
End of abdomen, posterior view, X 13.
First maxilla, x 26.
Tip of inner division of same, X 65.
Inner division of maxilliped, X 26.
. Outline of head, front view, X 10.
Side view of head and first segment of body, X 7.8.
Another specimen, alluded to above, (Cat. No. 3249), also a male
but slightly smaller than the one described, labeled Medje, July 1914,
is probably of this same species, but is in a rather poor state of preserva-
tion.
86 Bulletin American Museum of Natural History [Vol. XLIII
Figs. 63 to 70. Eubelum asperius, new species.
63. Pleopoda of female, 12.
64. End of abdomen, posterior view, X 16.
65. First maxilla, < 46. ‘
66. Tip of inner division of same, X 90.
67. Inner division of maxilliped, X 42.
68. Outline of head, front view, X 9.6.
69. Antenna, X 16.
70. Side view of head and first segment of body, X 8.5.
Eubelum asperius, new species
Text Figures 63 to 70
The only specimen obtained is a female, 8.5 mm. long, with an empty
brood-pouch. This species also very closely resembles E. stipulatum,
described above, and the description there givea will apply to this form
also with the following modifications.
Body surface rougher, with small irregularly distributed roughened pits, easily
visible with low magnification, closely distributed over the surface; they are slightly
setose, giving the body surface a somewhat pubescent character. Ground color of
upper parts rather light brown, lower parts and legs pale yellowish brown. There are
the usual light markings and borders on the segments; the markings on the lateral
parte of the back form more regular rows of short bars than they do in Z. stipulatum.
1920] Van Name, Isopods of the Belgian Congo 87
Head narrower and a little more convex in front outline in a dorsal view. Seen
from in front, the anterior or upper margin of the epistome is highest at a point a little
each side of the median line, where it dips down a little and is less well marked. Eyes
proportionately a little larger, with about twenty ocelli. Antenne apparently a little
longer; their flagellum has but two articles,! the first being very short. First maxilla
with nine teeth (3+5 besides a slender accessory tooth beside tooth 2 on the outer
division) and about nine plumose processes on the inner division.
The characters of the first segment of the thorax are closely similar to those of
E. stipulatum, except that the posterior angle is scarcely truncated but merely a little
blunted at the apex. The impressed groove marking off the projecting border from
the body of the segment turns up sharply and conspicuously just behind its anterior
end near the eye. The sulcus on the external-ventral aspect of the border is less
sharply defined. The notch for receiving the second segment when the body is
rolled up is small but with rather widely divergent sides; its inner side is produced
considerably farther back then the outer and is sharply rounded off. The second and
third thoracic segments have the inner side of the anterior margin of the lateral parts
quite conspicuously thickened but they bear no processes. These two segments are
a little more rounded at the ends than in LZ. stipulatum. The spines on the legs are
much worn and broken in this specimen; they appear to have been only moderately
numerous.
Last segment of abdomen wide, its median extension short, with the sides curved
and converging toward the wide, squarely truncated end. The exposed parts of the
basal segments of the uropoda are somewhat triangular, with the external surface
moderately convex without any large depression. The notch for the small short
rudiment of the external branch is on tle posterior margin well removed from the
inner angle, and lies chiefly on the external aspect; seen from the inner or anterior
side the margin is only a little indented by this notch. The inner branches of the
uropoda do not reach to the end of the abdomen.
The only specimen (Cat. No. 3256) is from Zambi, June 1915. Of
the species described by Budde-Lund (1899), it appears most closely
related to E. ignavum from Abyssinia. One of the most conspicuous
differences between the two forms is in the inner lamella of the cleft
rear angle of the first body segment, which in the present species is
considerably extended backward beyond the outer, but is described in the
Abyssinian form as scarcely longer than the outer. The last segment of
the abdomen and the uropoda are also longer and less wide in FE. ignavum.
E. instrenuum Lénnberg and Budde-Lund, 1912, from British East
Africa appears to resemble this species in many characters but has the
flagellum of the antennz composed of three articles.
1Though in some groups the number of articles in the flagellum is a reliable character for generic
or subgeneric distinction, in Eubelum and probably also in some other sections of the Oniscide the char-
acter seems to be of less importance, a two-segmented flagellum having apparently developed independ-
ently in species not closely related by a coalescence of the two terminal articles.
88
73
75
Bulletin American Museum of Natural History
72
74
77
Figs. 71to78. EKubelum garamba, new species.
71. Pleopoda of female, X 18.
72. Antenna and outline of anterior segments, ventral view, X 22.
73. End of abdomen, posterior view, X 25.
74. Outline of head, front view, X 10,
75. First maxilla, * 44,
76. Tip of inner division of same, * 130,
77. Terminal part of second maxilla, X 44.
78. Inner division of maxilliped, * 44.
[Vol. XLIII
1920] Van Name, Isopods of the Belgian Congo 89
Eubelum garambe, new species
Text Figures 71 to 79
This species is represented by but one specimen, a female only about
7 mm. long, with marsupial plates developed. It is closely related to E.
stipulatum and even more closely to E. asperius, but the body, though
very convex, is a little narrower than in either of those species.
Surface even but with minute, not very closely distributed puncte and with
considerable pubescence. The thoracic segments have a considerable degree of in-
dividual convexity in an antero-posterior direction; the part overlapped by the
preceding segment is marked off by a rather conspicuous depressed line or furrow.
Color grayish brown above, with obscure yellowish markings and margins on the
segments; under parts yellow.
Fig. 79. Eubelum garambe, new species. Female, X 15.
Head narrow, superior line of epistome considerably arched and only very slightly
prominent, but continuous across the front of the head. Eyes rather large but with
comparatively few ocelli (about fifteen well-formed ones). Antenne of moderate
length and stoutness, very pubescent. No third article was demonstrated in the
flagellum, which is, however, fairly long and slender. First maxilla with eight teeth
(3+5), in addition to a very small accessory tooth beside tooth 2 on the outer divi-
sion, and about ten plumose processes on the inner division.
Posterior lateral corners of first thoracic segment produced into an angle, which
is not truncated or rounded off and is provided with a small cleft for the reception of
the second segment when the body is rolled up. Sides of this cleft only moderately
divergent, the inner side is rounded off and a little more extended than the, outer.
There is a prominent and thick, but not very wide, projecting lateral border on the
first thoracic segment marked off from the body of the segment by a well-defined
furrow. The border and furrow extend almost to the posterior angle of the segment
but the border becomes narrow and inconspicuous before the angle is reached.
There is a wide and fairly deep sulcus the whole length of the border on its external
ventral aspect, the sulcus widens noticeably in the posterior third of its length. The
lateral parts of the second and third thoracic segments have the anterior border
90 SREP LEE Bulletin American Museum of Natural History
Figs, 80 to 89, Hubelum tenebrarum, new species.
80, Female, x
Sl. Pleopoda of male, * 11,
82. Pleopoda of female, x 11.
84. Antenna and outline of anterior segments, ventral view, 11,
84. Tip of inner division of first maxilla, X 115,
85. Outline of head, front view, * 11,
86. Mandibles, anterior aspect, * 32,
87. First maxilla, X 32,
88. Inner division of maxilliped, * 34,
50. Antenna, X 24,
[Vol. XLIS
:
1920] Van Name, Isopods of the Belgian Congo 91
thickened on the inner surface but bear no processes. Their ends are rather narrowly
and sharply rounded and the ends of the remaining thoracic segments are less abruptly
truncated than in LZ. stipulatum. The legs are of moderate length and rather slender;
their spines are not very numerous but fairly stout. The abdomen is rather narrowly
rounded behind, the ends of the segments bend or flare outward a little. The last
segment is as in EZ, asperius but narrower and longer, and less broadly truncate at the
end. The basal segments of the uropoda have the exposed parts somewhat triangular;
their terminal border is quite oblique, the inner angle being much more produced
than the outer and exceeding a little the tip of the abdomen. The notch for the short
rounded rudiment of the external branch lies in the posterior border but mostly on
the external aspect. Seen from the anterior or inner side, the posterior border is
scarcely indented. The external surface of the basal segment of the uropoda is only
moderately convex and has no large depression. The inner branches of the uropoda
are rather stout; they do not reach to the end of the abdomen.
The only specimen (Cat. No. 3260) is from the extreme north-
eastern part of the Congo region, Garamba, collected in March 1912.
Fig. 90. Eubelum tenebrarum, new species. Female, X 11.
Eubelum tenebrarum, new species
Text Figures 80 to 90
_ Body moderately wide and of oblong outline when seen from above; rather soft
and quite loosely articulated; the lateral ends of the segments slightly extended. The
back is not very convex; its surface is smooth but not shiny; punctate under magni-
fication, but not pubescent. Color in alcohol dull yellowish white (white in life
according to notes taken by the collector). Size of a large female, 8.7 mm. long by
4.3 mm. wide; the males are smaller, a large one measuring 6.8 mm. long by 3.4 mm.
wide.
Head small; its anterior outline nearly straight in the middle, though receding
toward the corners. The superior margin of the epistome forms a projecting border
only near the sides; in the middle part the line of demarkation disappears. Mouth-
parts forming a prominently projecting mass. First maxilla with eight teeth (8+4
with an additional accessory tooth beside the second) on the outer division; the inner
division bears a row of about eight plumose processes. Antenne very long and
slender; the flagellum long, of two distinct articles of which the last is about two and
92 . Bulletin American Museum of Natural History [Vol. XLIII
one-half times the length of the first. In some individuals a very obscure segmen-
tation of the second article into two was demonstrated; in others I could not detect
it. Eyes very small and imperfect, with indistinct ocelli and very little pigment.
First segment of thorax with a moderately thick, but rather narrow, projecting
border separated from the main part of the segment by a narrow deeply impressed
groove. The border, which extends about four-fifths of the length of the margin,
has a fairly well-marked sulcus along its external aspect. This sulcus is slightly
obliquely situated so that it is somewhat visible from a ventral direction also,
especially toward the posterior end. The posterior lateral angle of the segment is
rounded off in a small curve and has a small cleft to receive the second segment;
the outer side of the cleft extends a little farther back but not so far ventrally as the
inner. The front margin of the epimera of the second, third, and fourth segments is
thickened on the inner side but none of them bear any process. The thoracic seg-
ments are somewhat rounded at the ends, especially in the anterior part of the body.
The legs are rather long, quite slender, and rather weak.
The epimera of the third, fourth, and fifth abdominal segments end in rather
sharp, backwardly directed angles. The last segment has a wide but very short base,
the median extension has nearly parallel sides and a very rounded extremity. The
basal segments of the uropoda are deeply notched on the posterior margin; these
notches are occupied by the outer branches which, though small, are elongate and
sharp pointed; they are visible both in a dorsal and ventral view of the body. The
inner branches are long and tapering; they lie beneath the median part of the last
segment, extending to its end.
About sixty specimens (Cat. Nos. 3242 and 3266) of this species
were collected at Thysville, June 2, 1915, “from the shelves of a cave’’
which is described in Bull. Amer. Mus. Nat. Hist., XXXVII, p.
541. A few of the females have a brood-pouch developed, containing a
few large eggs. The rather soft, loosely articulated body, the lack of
pigmentation and the almost functionless eyes are modifications that
would be expected in a form inhabiting such a cave as that in which this
species lives. These modifications are, however, of a very superficial
kind, and in the form and structure of its parts there seems to be no
sufficient reason for placing it elsewhere than in the typical subgenus of
Eubelum. ;
Oniscing (= Oniscide auct. mult.)
Synarmadillo globus Budde-Lund
Text Figures 91 to 100
Synarmadillo globus Buove-Lunp, 1909, in Voeltzkow, Reise in Ostafrika, LI, p.
276, Pl. x1u, figs. 37-44.
Body broad in proportion to its length; back very convex; front outline of head
strongly and evenly convex; rear end of body rather broadly rounded, Articulation
very compact and smooth; body surface unusually smooth, showing under consider-
able magnification only very minute and even granulation and numerous depressed
1920] Van Name, Isopods of the Belgian Congo 93
Figs. 91 to 99. Synarmadillo globus Budde-Lund, 1908.
91. Male, X 6.
92. Pleopoda of male, x 7.2.
93. Antenna and outline of anterior segments, ventral view, X 6.
94. Outline of head, front view, X 9.
95. First maxilla, x 20.
96. Tip of inner division of same, X 50.
97. Terminal part of second maxilla, x 20.
98. Inner division of maxilliped, < 20.
99. End of abdomen, posterior view, X 7.2.
94 Bulletin American Museum of Natural History (Vol. XLIII
dots or puncte. Color dark slaty gray above, the segments marked and bordered
with pale yellow, the under parts and legs also pale yellow. This is the largest and
most conspicuously marked terrestrial isopod collected by the Expedition; the single
specimen obtained, though a male and hence doubtless considerably inferior to adult
females in size, would measure nearly, if not quite, 15 mm. long if straightened cut,
and its stoutness and highly convex back give it unusual bulk foritslength.
Head rather small, the superior border of the epistome forming only a very
narrow projecting border, but this extends completely across the front of the head.
Eyes proportionately small, with about twenty-two ocelli; antenne rather short and
small, the fourth and fifth segments nearly equal in length, the flagellum with only
two articles, the basal about half as long as the terminal one. The mandibles are
more elongate and less crooked than in the next species (Synarmadillo lubilensis).
The first maxilla has nine teeth (4+-5, two of these representing enlarged accessory
Fig. 100, Synarmadillo globus Budde-Lund, 1908, Male, X 7.
teeth beside the main row) on the outer division, and two plumose processes and a
very minute spine on the inner division. Seen from one side, the lateral margin of
the first thoracic segment is curved, especially in the posterior part; its posterior
angle is cleft to receive the second segment when the body rolls up. The outer side of
this cleft is less produced downward and backward than the inner; the latter forms
a rather sharp angle. The whole length of the margin, almost to the extreme rear
angle, is turned outward to form a thick but narrow projecting border of even width.
This border turns sharply out from the main part of the segment, thus forming a
fairly distinct line of demarkation but there is no impressed groove along that line.
The border has a sulcus along its whole length; the sulcus lies on the external inferior
aspect and is rather narrow, widening, however, somewhat rapidly a little way before
the cleft at the rear angle is reached.
Second segment of thorax with a short process on the inside of the lateral part;
third segment with a slight rudiment of a process, The second to fourth thoracic
segments inclusive have the ends somewhat angular, though a little rounded off,
and not produced beyond the general outline of the body; the fifth has the ends
broadly rounded; the sixth and seventh are rather squarely cut off. Legs rather long;
they have few but stout spines.
ee
1920] Van Name, Isopods of the Belgian Congo 95
This species differs from all related forms collected by the Expedi-
tion in having the last segment of the abdomen taper to a slightly
rounded triangular point, which, however, is not quite long enough to
equal the truncated ends of the basal segments of the uropoda that fill
in the outline of the body on either side of it. The external branches of
the uropoda are represented by an exceedingly minute rounded rudiment
borne on the external aspect of the basal segment close to, but a little
above, the terminal margin (which is not indented) and near the inner
corner. Just external to the rudimentary external branch there is a
minute pore or pit filled with a yellow substance. The internal branches
are long and slender and reach a trifle beyond the triangular tip of the
last abdominal segment.
The only specimen (Cat. No. 3243) was collected at Zambi in June
1915. It corresponds well with Budde-Lund’s description and figures.
His specimens were from Cameroon (Bibundi and Bonge), collected by
Dr. Y. Sjéstedt in November 1891. ;
Synarmadillo lubilensis, new species
Text Figures 101 to 110
The single specimen collected is a female, 13 mm. long, apparently
fully adult, but without a brood-pouch.
Body oblong in a dorsal view, contractile into a ball; the head wide, with its
front outline only slightly convex; the abdomen broadly rounded behind. Articula-
tion not very compact; back moderately convex, its surface slightly and very finely
granular under magnification. Color dark brown above with obscure yellowish mark-
ings and margins on the segments; under parts yellowish.
The anterior margin of the epistome forms a moderately prominent projecting
border extending across the front of the head. Eyes rather small, with about twenty
ocelli. Antennz of moderate length, the fifth segment considerably longer than the
fourth, not very stout; the flagellum is rather long and slender with two well-defined
articles; its basal article is over one-third the length of the entire flagellum; the
second article is very obscurely segmented into two parts, the terminal part a little
longer than the proximal. The first maxilla has six teeth (3+-3, with an additional
small accessory tooth beside No. 2) on the outer division, and two plumose processes
and a conspicuous spine on the inner division. The mandibles are short and crooked;
they have (on the right mandible at least) but one small tuft (“penicillus” in the
terminology of Budde-Lund, 1909, p. 54) distal to the large brush-like appendage.
First segment of thorax with its posterior lateral angles slightly produced back-
ward and rounded off; there is a very small cleft to receive the second thoracic seg-
ment when the body is rolled up; the inner side of the eleft is considerably less pro-
duced than the outer. Seen from one side, the lateral margin of the first thoracic
segment is nearly straight: its border is thin, widely projecting and, especially in the
anterior part, considerably turned or rolled upward; there is no groove separating the
96 Bulletin American Museum of Natural History [Vol. XLII
Figs. 101 to 110, Synarmadillo lubilensis, new species,
101, Female, 72
102, Pleopoda of female, X 9.6,
103, End of abdomen, paral view, X 9.6,
104. First maxilla, < 27.
105. Tipofinner division of same, X 66,
106, Second maxilla, 27.
108. Antenna and outline of anterior segments, ventral view, X 7.8,
100, Outline of head, front view, 7.8,
110. Right mandible, anterior aspect, * 27,
cal site at
1920] Van Name, Isopods of the Belgian Congo 97
border from the body of the segment. On the inferior aspect of the edge of the seg-
ment (scarcely visible in a lateral view) there is a broad shallow sulcus (rather deeply
and sharply impressed along its inner margin and poorly defined along its outer
margin) that extends along the whole length of the border of the segment and is con-
tinuous behind with the above-mentioned cleft for the reception of the second seg-
ment. Lateral ends of the second to fourth thoracic segments somewhat rounded off
at the corners; those of the succeeding segments are more squarely cut off. Second
segment with a thickening or infolding of the anterior margin of the epimeral part
of the segment. This thickened part is produced at the end into a very slightly pro-
jecting rudiment of a process. The third segment has the margin a little thickened
but not at all produced. The legs are moderately long and rather slender, their
spines rather few and weak.
Fig. 111. Synarmadiilo lubilensis, new species. Female, X 8.
- Last segment of abdomen T-shaped; its median part is narrow and rather elon-
gate, with straight sides which converge toward the narrow truncated end. The
exposed parts of the basal segments of the uropoda are obliquely quadrangular and
rather narrow and elongate, with straight sides which converge toward the truncated
end. They have a minute notch directly on the posterior margin, close to its inner
corner for the very small and short rudiment of the outer branch.
The inner branches are long and narrow and slightly tapered, they reach almost
to the end of the terminal segment of the abdomen.
The only specimen (Cat. No. 3259) was found in a termite’s nest at
the Lubila River, September 20, 1909.
Cubaris' (Diploexochus) bananz, new species
Text Figures 112 to 117
This and the following species belong, according to Budde-Lund’s
(1909) classification, in the subgenus Diploexochus. While agreeing to
this subdivision of Cubaris as a natural one, the question may be raised
1Armadillo Latreille, 1804, commonly used as the generic name of this group is, as pointed out by
Stebbing (1902, p. 650), antedated by Armadillo Brisson, 1762, syn. of Dasypus, a genus of mammals,
and is therefore absolutely excluded. Cubaris Brandt, 1833, seems to be the next available name and
is here used in a comprehensive sense for the entire genus. If used as a subgeneric name, Cubaris must,
therefore, be employed for the typical Old World section of the genus.
Bulletin American Museum of Natural History [Vol. XLIIT
113
115
Figs, 112 to 117, Cubaria banana, new species,
2, Female, x 16,
3, Pleopoda of female, X 27.
4, Pleopoda of male, X 27,
5. End of abdomen, pomerior view, X 27.
ne
6 Antenna and outline of anterior segments, ventral view, X 18.5.
7. Female, X 16.
1920] Van Name, Isopods of the Belgian Congo 99
as to whether Budde-Lund was correct in using the name Diploexochus
for it, since that was originally applied by Brandt (1833, p. 192) to an
American species (C. echinatus) which may prove to be subgenerically
distinct from the African forms.
Body rather compactly articulated, ovate oblong as seen from above, widest at
the posterior end of the first segment of the thorax. The anterior end of the body is
broadly rounded, the posterior end has the outline of a rather narrow ellipse. Back
highly arched. Surface, when seen under considerable magnification, very slightly
granular and with a trace of pubescence. In addition to this minute granulation the
dorsal surface is raised into low elevations, though these are slight and poorly de-
fined. Along the middle part of the back these are mostly of rounded form and are
very low and inconspicuous; a group of three on the fore part of the first segment,
single median ones on the fourth and fifth abdominal segments, and a pair on the last
abdominal segment being the most prominent. Along the sides of the thorax they
are of oblong form and are more prominent. There are about six of these on each
side of each segment. The lateral ends of the posterior thoracic and of the abdominal
segments turn or flare slightly outward at the extreme edge. Color in the best pre-
served examples gray or grayish brown; the segments with a narrow yellowish or
pale border and yellowish markings, which in many cases correspond more or less
closely in situation and extent with the above described elevations of the surface.
The specimens are all of small size; the largest, could it be straightened out, would
hardly measure over 5.3 mm. long by 2.1 mm. wide. They may not have reached
their full size but do not have the appearance of being young. Except the usual
differences in the pleopoda (see Figs. 113, 114), no sexual differences were made out.
Head proportionately large; its front outline gently arched with a well-developed
projecting border. Antenne short but moderately stout. Their flagella consist of
two quite closely joined articles of which the first is very short. . Eyes large, with
about fifteen ocelli.
Lateral margin of first thoracic segment turned outward to form a fairly wide
projecting border, but this is not separated from the body of the segment by any
distinctly defined groove. The lateral edge is not thickened; on its ventral aspect
_there is a well-marked sulcus on the posterior half; this narrows gradually and closes
completely just in front of the middle of the segment. There is a small cleft (to
receive the anterior margin of the second segment when the body is rolled up) at the
posterior lateral angle of the first thoracic segment, which is conspicuously prolonged
backward and broadly truncated with the corners a little rounded off. The inner
side of the cleft extends backward fully as far as the outer side. The second to fourth
segments inclusive are rather short; their lateral ends are not much extended back-
ward and are rather narrowly rounded off; only the second bears a process on the
inner aspect of the lateral part. This process is, however, quite long, though narrow,
and is directed obliquely posteriorly and ventrally. The last three thoracic segments
are longer than the three preceding ones; the lateral ends of the last two are some-
what squarely truncated. The legs are fairly large and stout and bear few spines.
The first segment of the abdomen is nearly (in some positions of the body en-
tirely) concealed. The last segment is about two-thirds as long as it is wide and is
considerably contracted in the middle part; its truncated end is a little more than
half the width of the upper part of the segment. The inner branches of the uropoda
»
100 Bulletin American Museum of Natural History {[Vol. XLIII
(visible only from below) are short and thick and extend but a little way down the
inner aspect of the terminal abdominal segment; the outer branches are represented by
very minute oval rudiments on the inner margin of the outer aspect of the broad basal
segments of the uropoda, close to the constricted part of the last abdominal segment.
Specimens collected:
Cat. No. 3270. (Type) Banana, July-August 1915.
Cat. No. 3268. (Paratypes) Banana, July-August 1916, 9 specimens.
Cat. No. 3269. Zambi, June 1915. One male.
From two small species of this group which would seem from the
descriptions to be of somewhat similar appearance, C. bituberculatus and
C. nanus, described by Budde-Lund (1910) from the region of Mt. Kili-
manjaro, East Africa, this form is distinguished by many minor
characters—from the former, among other differences, by not having the
sulcus on the inferior margin of the first segment extending the whole
length; from the latter by the outline of the rear margin of the first
segment, which in that species is described as ‘‘subrectus, utrinque ad
angulos laterales levissime incurvus.’”’ In the present species the rear
margin makes a considerable angle above the conspicuously produced
lateral corners. C. liliputanus (Dollfus), 1895, from Pretoria, Transvaal,
is another allied form, but it has a longer sulcus on the first segment, the
outer branches of the uropoda less rudimentary, and the terminal ab-
dominal segment more constricted.
Cubaris (Diploexochus) regulus, new species
Text Figures 118 to 121
In spite of its very different appearance, due to the great development of the
ornamentation of the dorsal surface, this species differs little from that last described
in general form and in the details of its appendages. The largest specimen (both of
the two good specimens available are females) slightly exceeds in size any of those
of C. banana, although if it could be fully straightened out it would hardly measure
over 6 mm. long. Such differences in the general form of the body as exist are due
chiefly to the lateral ends of the segments being more squarely cut off and turning or
flaring outward in a horizontal direction very much more than in C. bananw. Antenne
short and small, their flagellum with two articles. Eyes with about fourteen ocelli.
The outline of each side of the first thoracic segment is nearly straight when seen in a
dorsal view. The sulcus on the ventral aspect of the lateral border of this segment is
searcely at all developed. There is only a small cleft to receive the second segment
when the body is rolled up; the outer side of this cleft is extended backward consider-
ably more than the inner and it is rather widely truncated in an oblique direction.
Second thoracic segment with a backwardly directed, tapering process on the inner
side of the lateral part; the third segment has no process, The legs have some of their
segments, especially the carpus, somewhat swollen or expanded dorsally. Color in
alcohol grayish above, with the summits of the tubercles and borders of the segments,
as well as the legs and under parts, light yellowish.
———— eee >
NO0O0.0_0_000
(0 0.00 0M
00.0 0.0
102 Bulletin American Museum of Natural History [Vol. XLITI
Dorsal surface ornamented with large, highly elevated tubercles regularly
arranged. They are mostly of more or less conical form with an oval base and rounded
summit. In addition there is a transverse ridge at the posterior margin of the head,
and near the lateral ends of the second to seventh thoracic segments there is on each
side a large oblong ridge from which a curved elevation, less raised, extends down on
the epimeron. The above tubercles are in addition to a conspicuous granulation of
the surface; these minute granules are slightly setose. The tubercles are distributed
on the head chiefly in two transverse rows; on the first thoracic segment in four
rows; on the remaining thoracic segments in two rows, the posterior row having an
odd number (seven or nine) and hence a median tubercle, the anterior row eight, none
being median. The abdominal segments bear tubercles in one transverse row as
follows: the first two, none; the second and third, six each; the fourth, four; the
last segment, two. The pleopoda and uropoda differ scarcely at all from those of
the last described species, though the rudiment of the external branch of the uropoda
is larger.
Two females, Cat. No. 3251 (the larger one the type) were collected
at Stanleyville in March 1915, with specimens of Eubelum propinquum.
Another specimen (Cat. No. 3301), in poor condition, was found in the
stomach of a toad (Bufo regularis) collected at Avakubi, Oct. 2, 1909.
This form is related to C. formicarum (Budde-Lund), 1909, p. 57, Pl.
v, figs. 44-56, from South Africa, but the figures show, among other
differences, that the terminal segment of the abdomen and the uropoda
are much more elongated in Budde-Lund’s species than in the present one.
Niambia squamata (Budde-Lund)
Text Figures 122 to 126
Porcellio (Leptotrichus) sgquamatus Buppr-Lunp, 1885, Crust. Isop. Terrestr., p. 196.
? Leptotrichus squamatus Douurvus, 1898, Bull. Soc. Zool. France, XXIII, p. 125.
Niambia squamata Buppve-Lunp, 1904, Revision Crust. Isop. Terrestr., p. 37; 1908,
in Voeltzkow, Reise in Ostafrika, II, p. 280, 295; 1909, in Schultze, Ergebn.
Forschungsr. Siidafrika, Il, p. 60, Pl. vi, figs. 1-3. Srepsina, 1910, Ann.
South African Mus., VI, p. 441.
Body as seen from above very wide; the general outline broadly rounded in
front and behind; it is fairly compactly articulated; both the head and abdomen are
short and moderately set into the thorax. The back is quite convex; the surface
smooth and even, but thickly covered with small evenly distributed setose granules.
Color slaty gray above, with inconspicuous yellowish markings on the lateral areas of
the thorax; legs and lower parts yellowish. The animal does not appear to be able
to roll up into a ball, In the single female specimen the body broadens posterior to
the middle, reaching its greatest width about the end of the fifth segment; in the two
male specimens the outline seen from above is more oblong, with sides more nearly
parallel, The female also has the back more strongly convex, Size of female (not
fully adult), 5.6 mm. long by 3 mm, wide; largest male, 7.2 mm. long by 4.1 mm,
wide.
1920] _ Van Name, Isopods of the Belgian Congo 103
Head small; as seen from above its posterior border is sinuous, while its anterior
outline appears simply convex; two small but very prominent lobes are, however,
present at the anterior lateral corners but extend so directly downward (ventrally)
that they are barely if at all visible in a dorsal view. The superior border of the epi-
stome is marked by an impressed line. The eyes are small, with few ocelli, and are
situated just above the small lateral lobes; they are likewise but slightly visible in a
dorsal view. Antenne short but fairly stout; their flagella have two articles, of which
the last is about twice as long as the first.
Figs. 122 and 123. - Niambia squamata (Budde-Lund), 1885. Female, x 15.
The first thoracic segment is considerably the longest; the fourth and seventh
are noticeably shorter than the others. Their lateral ends are cut off almost squarely,
but the corners of the first two are rounded off and the posterior corners of the last
four or five are produced a little backward, the posterior ones to the greatest extent.
Legs moderately long and stout (the anterior ones shorter), with sharp, moderately
long claws and numerous rather short, stout spines at the joints and on the ventral
side, especially on the merus and carpus of each limb.
104 Bulletin American Museum of Natural History [Vol. XLITI
Figs. 124 to 126. Niambia squamata (Budde-Lund), 1885.
129. Male, X 12.
125. Pleopoda of male, X 12.
126. Pleopoda of female, x 14.
Abdomen with all six segments separate and exposed, The lateral parts of the
first and second are covered by the thorax; the following three have the ends extended
and tapering to a point, and bent almost directly backward. The terminal segment
has the median part of its dorsal surface depressed or concave; the outline of the
segment is triangular, with concave sides tapering behind to a rather sharp triangular
point. The basal joints of the uropoda are slightly tapering; they do not reach as far
as the tip of the last segment. The external branches are large, rather wide at the
base, and tapering gradually to a point. The inner branches are elongate and tapering
and lie along the ventral aspect of the last segment of the abdomen but do not quite
reach its tip.
Three specimens(Cat. No. 3254) were obtained at Zambi, June 1915.
The species has previously been recorded from the Portuguese Congo
(Landana, Chinchoxo) and doubtfully (Dollfus, 1898) from Senegal.
1920] Van Name, Isopods of the Belgian Congo 105
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652-680, Pls. 32-35.
Leacn, W. E. 1815. A tabular view of the external Characters of Four Classes of
Animals which Linné arranged under Insecta. Trans. Linnean Soc.
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1818. Cymothoadées, in Dictionnaire des Sciences naturelles, XII, pp. 338-
354,
Lénwnpera, E. anp Buppe-Lunp, G. 1912. Crustacea collected by the Swedish
Zoological Expedition to British Hast Africa, 1911, Ark, Zool. k.
Svensk. Vetenskpsakad., VII, (No. 26), pp. 1-9, 5 figs.
Mrers, E.J, 1881, Ona Collection of Crustacea made by Baron Hermann Maltzam
on Gorée Island, Senegambia. Ann. Mag. Nat, Hist., (5) VIII, pp. 204—
220, 259-281, 364-377, Pls. xur—xvi, (Isopoda, pp, 369-371 and 376.)
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1920] | Van Name, Isopods of the Belgian Congo 107
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1 Pl.-
1906. Spedizione al Ruwenzori di S$. A. R. Luigi Amedeo di Savoia Duca
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544, pp. 1-2.
1909. Crostacei. Il Ruwenzori, Parte Scientif., I, pp. 357-358.
Perty, M. 1830-1834. Delectus animalium articulatorum que in itinere per
Braziliam annis 1817-1820 collegerunt J. B. de Spix et C. V. Ph. de
Martius. Monachii.
Ricuwarpson, H. 1897. Description of a new species of Spheroma. Proce. Biol.
Soc. Washington, XI, pp. 105-107.
1900. Synopses of North American Invertebrates. VIII. The Isopoda.
Amer. Naturalist, XXXIV, pp. 207-230, 295-309, figs. 1-16.
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59.57,8 (67.5)
Article VI— LEPIDOPTERA OF THE CONGO, BEING A SYSTE-
MATIC LIST OF THE BUTTERFLIES AND MOTHS COL-
LECTED BY THE AMERICAN MUSEUM OF NATURAL
HISTORY CONGO EXPEDITION, TOGETHER
WITH DESCRIPTIONS OF SOME HITHER-
TO UNDESCRIBED SPECIES!
By W. J. HoLuanp
Piates VI tro XIV anv 9 Text FiGuRES
About twenty-five years ago I published a number of papers upon
the Lepidoptera of tropical Africa, in which I described numerous forms,
which appeared to me to be new to science. These papers appeared in
various journals.? I am pleased to observe that in most cases the
correctness of my judgment with the lapse of time has been con-
firmed, and but few of the species named in these publications have
been relegated to the synonymy. In the case of the moths my industri-
ous and learned friend, Sir George F. Hampson, in his monumental
work upon the moths of the world, has in some instances changed the
generic references, but has accepted most of the new genera of Hetero-
cera which I proposed, and has allowed my specific names tostand. At
the time to which I refer I had in contemplation the preparation of a
comprehensive work upon the butterflies of Africa. As a preliminary
to this large undertaking I published in the Proceedings of the Zoo-
logical Society of London, 1896, pp. 1-104, a ‘Synonymic Catalogue
of the Hesperiide of Africa and the Adjacent Islands.’ Shortly after
this had appeared I became aware through correspondence that my
friend, Dr. Christopher Aurivillius, the Secretary of the Royal Academy
of Science in Stockholm, was about to publish a work upon the same
subject. It presently appeared under the title ‘Rhopalocera Aithiopica.’
To my astonishment I discovered that the author had paid me the
compliment of omitting from his treatise the great family of the Hes-
periide, for the reason, as he states in his introduction, that this family
had been so thoroughly covered in my recently published paper, that
he did not deem it necessary to retraverse the ground. With the appear-
ance of the great work of Dr. Aurivillius the motive to further prosecute
my self-imposed labors vanished to a great degree, but my interest in
1Scientific Results of the American Museum Congo Expedition. Entomology, No. 6.
2Entomological News, Psyche, Annals and Magazine of Natural History, Canadian Entomolo-
gist, Proceedings of the U. 8. National Museum, and Proceedings of the Zoological Society of London.
109
110 Bulletin American Museum of Natural History [Vol.. XLIIIL
the subject did not altogether cease. Though busy with many other
things, and becoming more and more absorbed in paleontological re-
searches, I still cherished a fondness for that particular faunal nook in
which I had passed so much time with pleasure, and kept on from time
to time adding to my collection, and latterly to that of the Carnegie
Museum, as opportunities presented themselves. The result has been
the gradual accumulation of a collection of the insects of Africa, and
particularly of the Lepidoptera, which is one of the largest in the world.
It is indeed far from complete, but, nevertheless, contains many thou-
sands of species and a vast assemblage of specimens.
At this point I am tempted to narrate a little incident, which I have
never recorded, but which is recalled to me as I am writing these pre-
liminary notes. In the summer of 1903 it became my duty to repair to
Brussels in order to supervise the removal from that city to Pittsburgh
of the great paleontological collection which had been brought together
by Baron Ernst Bayet, and which Mr. Andrew Carnegie had instructed
me to secure for the museum of which I have the honor to be the
Director. His Majesty, King Leopold, onthe day set apart to celebrate
Belgian Independence, sent me an invitation to lunch with him at the
Royal Palace. The Court was in attendance, because at half-past two
o’clock in the afternoon the king, attended by his ministers, was to re-
pair in state to the church of St. Gudule to join in the Te Deum.
After having been presented to the ladies and gentlemen of the Court
who were in waiting, I was introduced to Sir Ernst Cassel, the only
other guest, and then the king was announced. He entered the room
with the Princess Clementine upon his arm, came forward and greeted
Sir Ernst and me, turned the Princess Clementine over to me to escort
to table, and then with Cassel at his side bade the assembled company
follow on into the apartment where refreshments were provided. I
found myself seated between the King and the Princess. Conversation
went on merrily, and finally a mischievous notion occurred to me, and,
turning to the king, I said ‘“‘ Your Majesty, I have a favor to ask of you,
which I am sure you will be willing to grant, and the propriety of asking
for which I am sure you will recognize.’’ He looked me in the face and
answered, ‘“‘My dear Doctor, I shall be happy to serve you, if I can,
but what is the favor you wish?” I replied, ‘Your Majesty, I wish a
concession in the Congo Free State.”’ ‘A concession in the Congo
Free State!’ he exclaimed, and I saw a look of astonishment pass over
the faces of a number of those at the table. “Yes, your Majesty. But
I must explain myself fully. For many years I have been interested in
1920] Holland, Lepidoptera of the Congo 111
African entomology, and have named and described many species of
butterflies and moths from the region over which you in part bear rule.
I wish to go on with my work, and I therefore wish you to issue a decree
that the insect tribes of the Congo shall at a given date depute two of
each species, a male and a female, to surrender themselves as hostages
of science to whomsoever you may appoint to receive them, they then
_ to be turned over to me that I may go on with the good work of finish-
ing the task, which Adam left incomplete, of naming the living
things on the globe.’”’ The king laughed, and answered, “‘ Doctor, you
are as great a flatterer as the man who told King Canute to order the
waves of the sea to retire before him.”’ “What?” I replied, ‘‘is it pos-
sible that the royal prerogative does not extend so far?”’ “T am afraid
it does not,” he answered, ‘‘but I will tell you what to do. Go to the
Congo Museum and tell the Director that it is my wish that he shall
turn over to you all the butterflies which he has not yet named, so that
your wish may be gratified.” ‘“‘ Your Majesty,” I answered, “I know the
Director of the Congo Museum, and unless you give me a note to the
effect just stated, I am sure I shall not get a single butterfly into my
hands.” The king laughed and retorted, ‘‘ Aha! I discover that he is a
faithful servant of mine.”” We both laughed, and os was the end of this
particular bit of our conversation.
About two years ago I happened to visit The American Museum
of Natural History and, among other things, was shown the large col-
lection of Lepidoptera which had been brought back from the interior of
the Congo State by Messrs. Lang and Chapin. The gentlemen in charge
requested me to undertake the task of arranging and classifying the
Lepidoptera. In amoment of weakness I acceded to the request, but
with the proviso that they must not expect from me an immediate
report, in view of the fact that my duties were already very numerous
and that the work would have to be done at odd moments of time.
The collections were sent to me, the butterflies being, I think, more in
number than were in the Congo Museum in 1903. The result is the
list herewith presented, which represents work done in the midst of
incessant interruptions, or at times given to me for rest and vacation,
or when others have been in bed and asleep. It has been a labor of love.
I hope that to some extent it may help students of the future.
It is hardly necessary for me to enter into a lengthy and detailed
discussion of the relationship of the Ethiopian faunal region with those
of other parts of the globe. It suffices tosay that, while the lepidopterous
fauna of those parts of Africa which border immediately upon the
112 Bulletin American Museum of Natural History [Vol. XLIII
Mediterranean is distinctly palearctic, the fauna of the lands lying
south of the Sahara and traversed by the great river systems of the con-
tinent is more nearly related to the Indo-Malaysian fauna, but pos-
sesses a number of genera and species which occur nowhere else upon the
globe. Beginning in the southern part of Senegal, in latitude 12 ° N.,
and extending eastward and southward to the headwaters of the various
affluents of the Congo and the Coanza, there is a more or less densely
forested region, thoughout which the flora and fauna with slight modi-
fications are practically the same. Southern and eastern Africa are char-
acterized by the presence of great expanses of grass-land, save along the
watercourses. This territory, in which there is more or less aridity,
extends northwest from the region of Uganda and thence west about the
headwaters of the various rivers flowing into the Atlantic south of the
Sahara, forming a selvage between the hot densely forested jungle-
lands to the south and the dry desert-lands to the north. Here and
there the forested country is interrupted, asin Angola and various points
along the western coast, by smaller tracts where the forests are less
luxuriant and open grass-lands occur. The lepidopterous fauna of the
grass-lands, which until the end of the last century have been the home
of vast herds of ruminant animals, reveals the predominance of certain
genera, such as Teracolus, which are characteristic also of Abyssinia,
Arabia, and southern India. The humid jungles along the Coanza, the
Congo and its tributaries, the Ogové, the lower Niger, and the various
rivers emptying into the Atlantic from Lagos to Dakar are the home of a
fauna which by common consent is known as West African. Here is the
metropolis of the African Nymphalide, of various mimetic forms of
Lycenide belonging to the genera of the subfamily Liptenine and of
various genera of the Hesperiide, which are found nowhere else upon
the globe. Here and there this great forest region is invaded on its
eastern and northern borders by inwardly projecting stretches of the
surrounding grass-lands, and there is thus noted a transition on its
periphery from the West African fauna to the South African or East
African fauna. In fact, the East African fauna and South African fauna
reappear on the north and northwest of the irregular territory in which
the West African fauna occurs. At the very mouth of the Congo there
occurs in the region of Matadi an intrusion into the West African region
of some forms which may properly be considered as characteristic of
the South and East African fauna.
The largest portion of the collection returned by the American
Museum Congo Expedition was obtained at Medje, a point near the
Oe
ee
See? Sa
1920] Holland, Lepidoptera of the Congo 113
Nepoko River in the very heart of the forest. The collections at Medje
were principally made from April until September, 1910. A number of
specimens are recorded as taken at points not far distant from Medje,
such as Gamangui, Bafwabaka, and Avakubi. In the fall of the year
1910 and thereafter during the year 1912 considerable collecting was
done at Niangara and Faradje, the former on the Uelle River, the latter
upon the Dungu, an eastern affluent of the Uelle. The collections from
Niangara and Faradje reveal the fact that thosé localities, while still
within the limits of the West African subregion, are nevertheless not
far from the line of contact with the East African, or Abyssinian, region
which sends a long, narrow, westward projection south of the Sahara
toward the mouth of the Senegal. Specimens were occasionally taken in
the course of the journeyings of the members of the expedition at
various points throughout the region from Angola eastward, and the
labels attached to the insects reveal captures made at such points as
Matadi, Basoko, Stanleyville, and Bafwaboli. Regular and systematic
collecting, however, seems to have been confined largely to the three
points already indicated, Medje, Niangara, and Faradje, and more than
nine-tenths of the specimens brought back bear these locality labels.
The collection as a whole has a distinctly West African facies.
The collection is one of the largest which has been made in recent
years in that region. I am given to understand by the gentlemen who
made it that to a considerable extent they employed the assistance of
natives. It is particularly rich in the larger and showier species, es-
pecially of the Nymphalide, which are characteristic of the territory
visited. But little attention was paid to the moths, which is much to be
regretted, as it is among these that the greatest number of novelties
might have been expected to occur. Mr. Lang tells me that little col-
ecting was done at night. The smaller diurnal lepidoptera are also but
scantily represented. This is particularly true of the Lycznide and the
Hesperiide. Of course I understand the principal aim of the expedition
was to collect vertebrates. The making of entomological collections
was more or less a subsidiary purpose. Nevertheless, it is upon the whole
an extensive collection, containing not far from nine thousand specimens,
representing more than seven hundred and twenty-five species and
varieties. Most of the specimens are in good condition, and, while not
yielding as many species new to science as I had hoped might be the
case, it gives the American Museum a fine nucleus upon which to build
in coming years.
114 Bulletin American Museum of Natural History [Vol. XLIII
In preparing the following list I have acted upon the suggestion of
Mr. Lang to give a reference to the most easily accessible illustration of
each species. In a few cases no illustration has as yet been published,
in other cases the only illustration is in a recondite corner of the litera-
ture, and I have not in some such instances taken the pains to cite the
figure, as the specialist interested in the subject will know as well as I
how to find such illustrations. I have constantly referred, in the case
of the butterflies, to the illustrations given by Aurivillius in Vol. XIII
of Seitz’s ‘Die- Gross-Schmetterlinge der Erde.’ This volume gives, for
the most part, very good figures of the Rhopalocera of the Ethiopian
region. The copy in my possession unfortunately is incomplete, its
publication apparently having been interrupted at the outbreak of the
war. At all events, since the fall of 1914 no parts of the book which was
being issued at Leipzig have come to hand.!
In no instance have I endeavored to give a complete synonymy
where a species has been frequently mentioned in the literature of the
subject. I have, however, endeavored in all cases to cite the original
description or figure of the species and havé followed this by citation
of one of the latest references to that species, or of the subspecies, in
ease a subspecifie reference is called for. The student who desires to
explore the synonymy may consult among other works the ‘Rhopalo-
cera Aithiopica’ of Dr. Aurivillius and the revisions published by Roth-
schild and Jordan in the ‘Novitates Zoologice’ of the genera Charazes,
Papilio, and the family Sphingide. The synonymy of the Hesperiide
published up to the year 1895 is given quite completely in my ‘Synony-
mic Catalogue of the Hesperiide of Africa,’ and so forth (ef. Proc.
Zool. Soc. London, 1896). In studying the moths, reference should be
made to Sir George F. Hampson’s ‘Catalogue of the Phalsenz’ so far as
published. Assistance may be derived from Kirby’s ‘Catalogue of the
Lepidoptera-Heterocera,’ but this work must be used with caution for,
although references to the literature are correct, many species have in
recent years been assigned to other genera than those under which Kirby
listed them. In studying the Pyraustids and allied groups the writings
of Sir George F. Hampson must be consulted, and in studying the
Geometridae it is necessary to consult various papers published in recent
years by Warren, a number of which appeared in the ‘Novitates Zoologi-
Ce, 4
'The edition I have is that published in the German lang e between which and that published
in the English language there may be a few slight discrepancies in the pagination,
1920] Holland, Lepidoptera of the Congo 115
I have not attempted to give in connection with this paper a com-
plete bibliography of the subject, as such an undertaking seems un-
necessary in the case of any student who has access to the works men-
tioned above and to the ‘Zoological Record.’ A complete bibliography
would constitute a considerable volume in itself.
Before closing these brief introductory notes I desire to express my
sincere gratitude to Dr. F. E. Lutz and his amiable associates in the
Department of Invertebrate Zoology in The American Museum of
Natural History, and to the Director of the Museum, Dr. F. A. Lucas,
who kindly granted me the privilege of reserving for our collections in
Pittsburgh a small series of duplicates of each species in cases where
there were duplicate specimens in sufficient number to permit the writer
to retain a few. For this generous permission, as well as for the uni-
formly kind indulgence shown me by the officials of The American Mu-
seum of Natural History, I desire to express my thanks. To Mr.
Herbert Lang, the leader of the expedition, I am indebted for a number
of kind letters written to me in answer to inquiries made during the
progress of my work.
APPROXIMATE LOCATION OF PLACES MENTIONED IN THIS PAPER
Avakubi.—1° 20’ N., 27° 40’ E. Isiro.—2° 50’ N., 27° 50’ E.
Bafwabaka.—2° 10’ N., 27° 50’ E. Ja R.—2° to 3° 30’ N., 12° 25’ to 15° E.
Bafwaboli.—0° 40’ N., 26° 10’ E. Kangvé.—0° 45’ S., 9° E.
Bafwasende.—1° 10’ N., 27° 15’ E. Kwamouth.—3° 20’ 8., 16° 10’ E.
Banalia.—1° 30’ N., 25° 40’ E. Leopoldville—4° 25’ S., 15° 20’ E.
Banana.—6° §., 12° 20’ E. Lolodorf.—3° 15’ N., 10° 40’ E.
Banza Manteka.—5° 30’ S., 13° 50’ E. Lubila R.—1° N., 26° 30’ E.
Barumbu.—1° 10’ N., 23° 20’ E. Lukolela.—1° 10’ 8., 17° 10’ E.
Basoko.—1° 20’ N., 23° 35’ E.’ Malela.—6° S., 12° 40’ E.
Batama.—1° N., 26° 40’ E. ; Matadi.—5° 50’ 8., 13° 35’ E.
Benito.—1° 35’ N., 9° 35’ E. Medje.—2° 25’ N., 27° 30’ E.
Bolengi.—0° 5’ S., 18° 10’ E. Munie Katoto.— 0° 35’ N., 26° 5’ E.
Boma.—5° 50’S., 13° 10’ E. . Ngayu.—1° 40’ N., 27° 40’ E.
Boyulu.—1° N., 27° E. Niangara.—3° 40’ N., 27° 50’ E.
Bumba.—2° 10’ N., 22° 30’ E. Noki.—5° 50’ S., 13° 30’ E.
Duala.—4° N., 9° 40’ E. Nouvelle Anvers.—1° 40’ N., 19° 10’ E.
Efulen.—2° 40’ N., 10° 45’ E. Ogové R.—1° S., 10° E.
Faradje.—3° 40’ N., 29° 40’ E. Panga.—1° 45’ N., 26° 15’ E.
Fernando Po.—3° 30’ N., 8° 30’ E. Pawa.—2° 25’ N., 27° 50’ E.
Freetown.—8° 30’ N., 13° 15’ W. Poko.—3° 10’ N., 26° 50’ E.
Gamangui.—2° 10’ N., 27° 20’ E. Risimu.—1° N., 26° 45’ E.
Isangi.—0° 50’ N., 24° 15’ E. Stanleyville—0° 30’ N., 25° 15’ E.
Ukaturaka.—2° N., 20° 30’ E.
116 Bulletin American Museum of Natural History [Vol. XLIII
New GENERA, WITH THEIR TYPE SPECIES
PAGE
Kallimula; type, K. osborni, new species. .... 2... 0... cece ee teens 149
Neptidomima; type, Neptis exaleuca Karsch................0 00 eee eens 164
New SPECIES AND VARIETIES, WITH THEIR TYPE LOCALITIES
PAGE
Bicyclus medontias var. obsoletus. Medije............... 00. cece cee cence 132
Mycalesis langi.. .MOd3e 5 0:05.5 10:5 Urs om. ead bis Me ate oa eee ee De 139
st Chapin. | Niangerae * so. cao Se ah reas Gs tees pF Neneh 140
Precis stygia ‘ver: fusctta: Mae os eS ahi oe be eke Se teat 148
Kallinwula osborné...” Mele 00565 SCRE eT Oe ea 150
Hypolimnas bartteloti var. obliterata. Medje................. 00.00 e eee ee 156
Buphadro ct var. noteta.: Medios ces" eta elas. I ee ae 173
6 ubotmidte.\ MOU uid ps cn ewinn Re ae 174
op om 06" fuloojasciata, Media. io. es os eke nee 174
iy " latajascinka. = Medeor er sy Oe as 175
" se! a epaiablier:: "RAGS, 6 60 5 Sethe ols cute oe eee 175
13 inandides. -Medje 3). 5) OSE RS BRS a ee ee ee 175
- VOMOMCR WARE i sis ease eis 5s bi EOS le a eee 177
. sypete var; miima. °° Media. o3 3 oo 36s ya os <0 deine 178
* Cpeemee. DENG 5.0 oo oe soo Sg ea cette ack einai ae 178
ad meodon ‘var. innotata; . Miedie» 60526 eR es 179
Buryphone lucast.: : Madios: ..5\55 6650S a DE Ce ls a ee 183 |
Dieslogyna kahli. : Medi... 5 6. T es ea Rage eee OE ES eA 191
. ohndale: THOU ie 55 is she bce a EER A EON a ees 191
Cymnethor herminia var. poénsis. Fernando Po...............0.e eee e eens 196
amngt. ** DEOGRE . 6 os wn ic Re a re b Ca 6 ceo 197
rf éapellides, MOAI). EOL NE Tne ey cs Ch ae 198
* - ‘@nte fone Fubida,”: Media oS. F5 a hot hae ce Cas eae omen 199
n Reginw-Biisabetha:: Meage.. |. aciaci tes tek Cas sac acci he ramen 201
" Og0da War. TURORCENG. TAOBVE oo Oiien wi ca css 0 os ss cuaemoeane 203
Charazes protoclea var. marginepunctata, Gaboon............ 6660 c eee eens 206
Totigna rothtoides. ~ MOG. 035 6 PSF ae epee is sae oS UE 214
S) pgdpetials, DACRE S i PEED HERR he He vey se aN TRAE ees 214
Bolioia tanigh.- Methes.5. oR ERROR. ee ee 217
Deudorta batthélides,: ‘Niaheara. os 2075s eae vas 6 oe 00k ae eee 221
Spindaste Ghiiptnd:, Niangaen. oc. i ccc cae sin's.+ s+ o's 00.0 oh acpi ane 225
Tridkema Wutel.:° MaGie io. Se EF ae ee es nk cov ee OS Vv sckt Pe 229
Oboronla ornate Var. flava, BOG Ere ee ia ks bc a a's ER Ps 235
Mylothris spica form donovani, Niangara,.........6. 0. 0cc ee eee vere eeenns 236
Papilio ridleyanus var. fumosus. Bafwasende.... 0.0.0.0... 000000 e eee eee 247
Abantls rubrac “Medias 5 iis RPP CE as CSc 0 se a RE CED es 253 :
Leplalina niangarensts. Niangara... ice eesec sec esavevdcceevesbovers 256
Myopoychs langl. “Me O56 55 CDEC SEE bs Cv in BV eee Te 262
Metarclla chaptnk.* Mayes: i558 hE cer ee tees ves os se eg ees 264
Nola bananés.. Banage: . o5 hice oa ae OL ROOT Sa ie nr Ae 266
1920] Holland, Lepidoptera of the Congo 117
Seer GNOMOLE, > Benito... 6255 Gis wihsda ts cael eal anemaeee es | 268
maunonmuomerys medjensis. Medje... .... 0... ....5000nscevevawtovewees 271
NINE SOCSBOTAGD 256s SD Pe cha hc eek oe ee 274
pease wariqoiuis. ~ Lolodorf . : «)(5,5 2idee8kh. Oe A eee eee 275
er eemnenes..) Dkattiraka...:. 5 .2'.5 .taucacss wes ot clit ee eens 277
Pere aeeemen. -AVAKUD o.oo)... bec civinde tip kaa ae aa eee 278
IEE EMO oP ee ee 280
Sphingomorpha aliena.. Banza Manteka................. 0.00. c eee e ee cees 285
ENE. AVORUD I i es LS cin puke eR oe aa ee le Rae 287
* eS OMIOMPOOR fF. 5 5.5. 50d ok ois Salk we ele eee 287
Ampnigonia hyalinata. Stanleyville....... os ccs’ topo cece ane eew end 288
srmermemmtt rwmeas, < MGIC... .. ... 6. oes cn chaeeeet ne se nah awe hee 289
DONT. GCC... chin eC kvnvcwsont undoes ek eemeael 291
mt ae Wat, polidior. Ukaturaka.. 000 Oe See 292
at NH MAOCIO 225.65 5h AE 6 dae VS SEA See ah a ee Re 292
- NNR. DAO 4 6 sis oa keinale cosas Chaban ool oe eae 292
af A OCT SE NORM RE Cg eer e28 293
ve CE REG 9, ahis c 0d oes hale c Ah MiSw eee Shae ee 294
ONTO Dik bs cla a's c's wlawo es ba ane sae bares 295
I BRBEE OMI OTOOD 55. 2) 68, d's ols ae SAE a Solis oe 296
a Scline «ope dle Ug ci Oe Sep a ae ae 298
a esa oh u's 0, ca: die, suse be # poate woh peasant lar dae ee 299
EE SS EEDSER CT RE ME He et es a een OP. 304
Cnr TL mmmsemete. MGGIC, . os sv ois cancun vos caccas ou bus im eee 305
meusere (1) weertanria... Medje..i.......5 6 0 og ee oe ee 307
eee edb. yaa, cae, Se a 310
Gomeememeariseneeria. Medje....... 22.6 .600. 00s as a ee 311
I UIIIEE DAOUIG oo. oii ons 8 sow eda e eo wdeb mete ee 312
Seen tT) MRMMEOOPOEG, DOI... ws. 2... +. oa ieie «ose stake akg bee Mien Ouie 313
Pseudoterpna (?) chapinaria. Belgian Congo..................0.-02000005 314
Eudemonia brachyura var. minor. Sierra Leone................6.00.00 00 316
I ES PORTE oe. oe backs Oda s oy # UR. Gaels bE Re Oe 318
Meee NERE, -MULUMIORT i. 5 gk Wales CRN Obs pnb wb elena oT 320
Rpmmemenenee: WOME, NIGTIGATA. (68. 6.5 is. cate as Ss » igi od > oe aid vk oe OE 322
Saliunca rubriventris. Stanleyville...................0-0000 eee Pewee nies 324
Proterozeuxis (?) medjensis. Medje......... Bo ae a oe eae aera a aa 325
er nnASnCES.- SOUGINES 2 3c. oa oa Bo a a Sia Ns oe eka pa ate ati eles 328
The following new names are proposed in this paper:
Mylothris spica form donovani Holland, p. 236, 9 of M. spica (Meeschler) =
9 Papilio rhodope Donovan, not 2 of P. rhodope Fabricius.
Anua hampsoni Holland, p. 280, for Anua producta Hampson, 1913, not A.
producta Holland, 1894.
118 Bulletin American Museum of Natural History [Vol. XLIIT
I. RHOPALOCERA
Danaide
Danais! Latreille
(1) |. Danais chrysippus (Linnzeus)
Papilio chrysippus LiInn2&vs, 1758, Syst. Nat., L0th Ed., p. 471.
Danaida chrysippus Avurtviutvs, 1911, Seitz, Gross-Schmett., XIII, p. 71.
In the entire collection, consisting of one hundred and ninety-seven
specimens of this species, there are only thirteen individuals which can
be referred to the typical form, D. chrystppus, and even these show traces
on the hind wings of the white squamation, which is characteristic of
the varietal form D. alcippus. As in the ease of the latter form, they fall
into two categories, those in which the upper side of the wings is bright
rufous and those in which it is dark chestnut or maroon. Noki, Kwa-
mouth, Medje, Niangara, and Faradje.
(2) la. Danais chrysippus alcippus (Cramer)
Papilio alcippus Cramer, 1777, Pap. Exot., II, p. 45, Pl. cxxvn, figs. Z, F.
Danaida alcippus AurRtviLui0s, 1910, Seitz, Gross-Schmett., XIII, p. 72.
One hundred and eighty-four specimens are referable to this varietal
form. There is great variation in the shading of the wings. Some have
the color of the fore wings bright rufous, others dark chestnut, while one
aberrant male has both the fore and the hind wings prevalently pale
argillaceous. The majority of the specimens have the hind wings
broadly white, which is the typical form, but there are many specimens
in which the white is reduced in extent until in some they nearly approach
the condition which prevails in typical D. chrysippus, in which the white
color is totally lacking. I find it utterly impossible by associating the
specimens under the dates of capture, or according to the localities
given upon the labels, to divide them into seasonal or local races.
According to the labels, captures took place in January and February,
then again in June, July, and August, and still later in November.
From this it appears probable that there are at least two annual broods,
and the insect may be on the wing throughout the year. Isangi, Avakubi,
Gamangui, Risimu, Niangara, Medje, Bafwabaka, and Faradje. |
‘As every student knows, the word Danaus, first introduced by Linnwus into entomological
nomenclature, was not by him and cannot be by us employed in a strictly generic sense, :
Aurivilliue has resuscitated the name Danaida originally employed by Latreille, but for which he
latter substituted Danaus and finally Danais. An author surely has a right to correct himself and
amend his nomenclature. Latreille did this and, as Danais has been universally employed for a cen-
tury, it seeme to the writer an excess of obedience to ‘the law of priority’’ to fish up Latreille’s lon g
abandoned and forgotten name and apply it again,
1920] Holland, Lepidoptera of the Congo 119
(3) 2. Danais petiverana Doubleday and Hewitson
Danais petiverana DouBLEDAY AND Hewitson, 1857, Gen. Diurn. Lep., I, p. 93, PI.
xu, fig. 1.
Danaida petiverana, AuriVituivs, 1911, Seitz, Gross-Schmett., XIII, p. 72, Pl. xxima.
This is the African form of D. limniace (Cramer). The collection
contains thirty-four males, taken mostly at Medje in June, July, and
August, though there are a number captured at Niangara in November,
and several from other localities: Bafwasende, Bafwaboli, Risimu,
Faradje.
Amauvris Hiibner
(4) 1. . Amauris niavius (Linnzus)
Papilio niavius Linnzvs, 1758, Syst. Nat., 10th Ed., p. 470.
Amauris niavius AuRIVILLIUsS, 1911, Seitz, Gross-Schmett., XIII, p. 74, Pl. xxmid.
There are twenty-five specimens, of which three (two of them not
expanded) are females. The most of them were taken at Medje from
July to September, though a few are labelled as taken at Niangara in
November.
(5) 2. Amauris damocles (Palisot de Beauvois)
Papilio damocles Pauisor pr Bravvois, 1805-1821, Ins. Rec. en Afrique et Amérique,
Lép., p. 239, Pl. VI, figs. 3a, 3b.
Amauris damocles AurtviLutus, 1911, Seitz, Gross-Schmett., XIII, p. 74.
This common species is represented in the collection by twenty-
eight males and two females. A few are labelled as having been cap-
tured at Gamangui in February, most of them were taken at Medje
about the middle of the year, and a few were captured at Niangara in
November.
(6) 3. Amauris psyttalea Ploctz
Amauris psyttalea Puarz, 1880, Stett. Ent. Zeit., XLI, p. 189. Aurtvitirus, 1911,
Seitz, Gross-Schmett., XIII, p. 74, Pl. xxva.
This species is discriminated from the preceding by the fact that
the white basal area of the hind wing reaches the tip or goes a little
beyond the tip of the cell, while in A. damocles it does not, and by the
further fact that the outer angle of the hind wing is adorned on the
upper side by a continuous band of marginal spots.
There are twenty-four males in the collection. They were taken
at various localities and at different dates from July to December.
120 Bulletin American Museum of Natural History (Vol. XLIII
(7) 4. Amauris damoclides Staudinger
Amauris damoclides STAUDINGER, 1896, Iris, VIII, p. 367, Pl. vu, fig. 3. AuRIVIL-
Lius, 1911, Seitz, Gross-Schmett., XIII, p. 74, Pl. xxvb.
There are two specimens of this species, one taken at Medje in
July, the other at Avakubi in November.
(8) 5. Amauris tartarea Mabille
Amauris tartarea MABILLE, 1876, Bull. Soe. Zool. France, I, p. 199. AuRIvILLIUs,
1911, Seitz, Gross-Schmett., XIII, p. 74, Pl. xxva.
Six males, five taken at Medje in June, July, and August, and one at
Niangara in November.
(9) 6. Amauris hecate Butler
Amauris hecate BUTLER, 1866, Proc. Zool. Soc. London, p. 44. Aurrvitirus, 1911,
Seitz, Gross-Schmett., XIII, p. 75, Pl. xx1vb.
Fifteen males, one taken at Stanleyville in August 1909, one at
Gamangui in February, two at Medje in September, six at Avakubi in
November, and five at Niangara in the same month.
(10) 7. Amauris hyalites Butler
Amauris hyalites Burumr, 1874, Cist. Ent., I, p. 209. AurtviLuirus, 1911, Seitz, Gross-
Schmett., XIII, p. 76, Pl. xxrva.
Of this species the collection contains forty-six males and five
females. Most of the specimens were taken at Niangara in November,
but quite a number are labelled as from Medje and other localities and
were taken in the midsummer months.
Acreidee ’
PitanemA Doubleday and Hewitson
(11) 1. Planema epea (Cramer)
Papilio epwa Cramer, 1779, Pap. Exot., III, p. 64, Pl. ccxxx, figs. B, C.
Planema epwa Avrivitrus, 1913, Seitz, Gross-Schmett., XIII, p. 245, Pl. viria.
The collection contains five males and seventeen females, all of
which were taken at Medje from July to September, except one female
which was collected at Faradje, “1911-1912.”
(12) 2. Planema tellus Aurivillius
Planema tellus Avnivitit0s, 1893, Ent. Tidskr., XIV, p. 280, fig. 7; 1913, Seitz,
Gross-Schmett., XIII, p. 245, Pl. pvuta,
Two males taken at Medje, one on June 6, the other on July 30,
1910.
a ae
i ee Le
1920] Holland, Lepidoptera of the Congo 121
(13) 3. Planema epiprotea Butler
Planema epiprotea Butier, 1874, Cist. Ent., I, p. 210. Aurtviniius, 1913, Seitz,
Gross-Schmett., XIII, p. 245, Pl. uvimb.
Twelve females taken at Medje from May to September.
(14) 4. Planema consanguinea Aurivillius
Planema consanguinea AvRIVILLIUS, 1893, Ent. Tidskr., XIV, p. 282, fig. 8; 1913,
Seitz, Gross-Schmett., XIII, p. 244, Pl. rvmb.
Two males caught at Medje, one in July, the other in September.
(15) 5. Planema elongata Butler
Planema elongata ButiER, 1874, Cist. Ent., I, p. 212. Aurtvinxius, 1913, Seitz,
Gross-Schmett., XIII, p. 244, Pl. rv1ub, c.
A single female taken at Medje, August 1, 1910.
(16) 6. Planema pseudeuryta Godman and Salvin
Planema pseudeuryta GODMAN AND SALvIn, 1890, Story of the Rear Column, p. 429.
Planema dewitzi SrauDINGER, 1896, Iris, [X, p. 209, Pl. 11, fig. 5.
Planema pseudeuryta AuRIvILLIvS, 1913, Seitz, Gross-Schmett., XIII, p. 243, Pl. rvmte.
I refer to this species a single male taken at Medje, May 6, 1910.
(17) 7. Planema nelsoni (Smith and Kirby)
. Plate VI: Figure 1, #; Figure 2, ?
Acrea nelsoni Smith AND KirsBy, 1892, Rhop. Exot., I, Acrea, Pl. 11, figs. 9, 10.
I refer to this species a male captured at Niangara, November 26,
and a female taken at Medje, August 24, 1910. As the female of P.
nelsoni has never been described or figured, I give on Plate VI arepresen-
tation of this female, and also of the male, for purposes of comparison.
The insect has been erroneously identified with P. poggei Dewitz.
(18) 8. Planema alcinoe (Felder)
Acrea alcinoé FEtpER, 1865-1867, Reise Novara, Lep., III, p. 368, Pl. xuvu, figs.
12, 13.
Planema alcinoé Aurtviuuius, 1913, Seitz, Gross-Schmett., XIII, p. 241, Pl. vu.
I assign to this species seven females, six taken at Medje at dates
ranging from May 6 to September 1, and another female taken at
Niangara, November 26, 1910. They agree very closely with specimens
in my collection determined as P. alcinoé by the late Dr. Otto Staudinger,
except that the discal area of the hind wings is pure white and not
creamy white, as is the case in the specimens sent me from Dresden. It
is possible that they may belong to a closely allied species, but there is
122 Bulletin American Museum of Natural History [Vol. XLII
no way of certainly determining this. The association of the sexes in
some species of this genus without the help of correct data obtained in
the field is a matter of conjecture, as the females of related species are
very much alike.
(19) 9. Planema macaria (Fabricius)
Papilio macaria Fasrictus, 1793, Ent. Syst., III, part 1, p. 174.
Planema macaria AuRIvILuivs, 1913, Seitz, Gross-Schmett., XIII, p. 240, Pl. tvuf.
One male caught at Medje, August 1, 1910.
Acrz#a Fabricius
(20) l. Acrea alciope Hewitson
Acrea alciope Hewitson, 1852, Exot. Butt., I, Acr@a, Pl. 1, figs. 4, 5.
Acrea fumida ELTRINGHAM, 1912, Trans. Ent. Soc. London, p. 325, @.
Acrea bakossua StranpD, 1912, Archiv f. Naturg., LX XVII, part 1, Suppl. 4,
. 114, 9.
Pk macarina BuTuER, 1868, Proc. Zool. Soc. London, p. 221, Pl. xvu, fig. 6, 2.
Acrea aurivillii StraAuDINGER, 1896, Iris, [X, p. 209, Pl. m1, fig. 2, 9.
Acrea alciope AuRIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 248, Pl. pve, o.
The collection contains sixty-five males and forty-one females of
this species and its varieties. The males are all quite alike, except seven
which do not have the ground-color of the wings a bright’ pale orange-
yellow, which is the normal color, but are dull brownish, and the darker
markings are not deep black but grayish black. There is also a single
female which, in these respects, is like the males I am describing and
agrees perfectly with the description of a female to which Eltringham
applied the subspecific name of fumida (cf. Eltringham, loc. cit.). The
name used by Eltringham is,in my judgment, also to be applied to these
males, and the aberration is evidently not in this case confined merely
to the female sex. Most of the aberrant female forms described by
authors are represented in the batch of specimens before me. There are
several specimens referable to the form dubbed macarina by Dr. Butler,
in which the margin of the hind wings is not marked with a dark band;
of the form named aurivillii by Dr. Staudinger, in which the hind wings
are crossed on the middle by a band of white of varying degrees of in-
tensity; and of the variety named bakossua by Strand, in which the
costal third of the transverse median band of the fore wings is whitish.
. And there are some intergrading forms which the writer, if he were a
professed ‘‘species-maker”’ and not engaged in other and more important
matters, might be tempted to describe and tag with so-called subspecific
names. These forms are mimetic, and some interesting observations
concerning them are contained in Eltringham’s work which is cited above.
1920] . Holland, Lepidoptera of the Congo 123
The vast majority of the specimens were taken at Medje, and mainly
in the months of June and July. There are about half a score of speci-
mens which bear other locality labels, but they give no hint whatever
of the existence of “local races.’ All of the varietal forms mentioned
above occurred at Medje and were taken at the same dates. Speci-
mens labelled as from Bafwaboli (Munie Katoto), Bafwasende,
Batama, Lubila, and Risimu are dated as captured in September; one
from Avakubi is dated October, another from Niangara bears the date
of November, and one from Gamangui was taken in February. The
writer has specimens in his collection taken in Cameroon and the
French Congo in March, April, and May, and it is probable that the
insect may be found upon the wing more or less commonly all the year
round in the regions which it frequents.
(21) 2. Acrea jodutta (Fabricius)
Papilio jodutta Fasricius, 1793, Ent. Syst., III, part 1, p. 175.
Acrea jodutta AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 249, Pl. xvite.
There are fifteen males and four females, most of them taken at
Medje from June to September. Of the females, two are of the typical
white form and two of the form named dorothee by Miss Emile Sharpe
(figured as A. metaprotea Butler, in Seitz, XIII, Pl. tvmd) taken at
Medje, June 20, 1910.
(22) 3. Acrea lycoa Godart
Acrea lycoa Gopart, 1819, Enc. Méth., IX, p. 239. Aurtvititus, 1913, Seitz, Gross-
Schmett., XIIT, p. 250.
There are five males and thirty.seven females in the collection.
The specimens were mostly taken at Medje from June to September,
but there are two females from Batama, two others from Risimu, and a
fifth from Bafwaboli(Munie Katoto), which are all dated as captured in |
September.
(23) 4. Acrea semivitrea Aurivillius
Acrea semivitrea Aurivituius, 1895, Ent. Tidskr., XVI, p. 111; 1913, Seitz,
Gross-Schmett., XIII, p. 253, Pl. rviib.
Three males taken at Medje, one in June, the other two in August.
(24) 5. Acrea servona Godart
Acrea servona Gopart, 1819, Enc. Méth., IX, p. 239. Aurivitirus, 1913, Seitz,
Gross-Schmett., XIII, p. 253, Pl. uvib.
Twenty-six males and one female, all taken at Medje (June-August),
except one male captured at Lubila, September 20, 1909, and the single
female caught at Gamangui, June 7, 1910. ;
124 Bulletin American Museum of Natural History [Vol. XLII
(25) 6. Acrea penelope Staudinger
Acrea penelope StauDINGER, 1896, Iris, IX, p. 195. AuRIVILLIUS, 1913, Seitz, Gross-
Schmett., XIII, p. 254, Pl. rvmb.
Fifteen males: three from Gamangui, two taken in February and
one in June; seven from Medje caught June to August; two from Risi-
mu and three from Munie Katoto taken in September.
(26) 7. Acrea peneleos Ward
Acrea peneleos Warp, 1871, Ent. Mo. Mag., VIII, p. 60. seine Mee i 1913, Seitz,
Gross-Schmett., XIII, p. 255, Pl. vib (first figure).
Twenty-nine males mostly taken at Medje, alehodeh there are
others taken at the same localities and in the same months as those
recorded for the preceding species. Lubila, Munie Katoto, Gamangui,
Ngayu, Risimu, and Boyulu.
(27) 7a. Acrea peneleos pelasgius (Grose-Smith)
Acrea pelasgius Grose-Smitu, 1900, Nov. Zool., VII, p. 545. Smiva anp Kirsy,
1901, Rhop. Exot., III, Acrea, Pl. vu, figs. 9, 10.
Eighteen specimens, all from Medje (April to August), except one
caught at Niangara in November.
(28) 8. Acrea parrhasia (Fabricius)
Papilio parrhasia Fasricius, 1793, Ent. Syst., III, part 1, p. 175. -
Acrea parrhasia AurIvLuius, 1913, Seitz, Gross-Schmett., XIII, p. 255, Pls. uvria,
LIXe.
One male, Munie Katoto, September 10, 1909.
(29) 9. Acrea orina Hewitson
Acrea orina Hewirson, 1874, Ent. Mo. Mag., XI, p. 180; 1875, Exot. Butt., V,
Acrea, P|. vu, figs. 43, 48.
There are thirty-one specimens, the majority of which were taken
at Medje at dates ranging from May to September. Two are from
Gamangui, one caught in February, the other in June; one from Stan-
leyville, August 1, 1909; one from Isangi, August 11, 1909; one from
each of the localities known as Pawa, Munie Katoto, and Avakubi,
captured in the month of October.
(30) 10. Acrwa orestia Hewitson
Acraa orestia Hewrrson, 1874, Ent. Mo, Mag., XI, p, 131; 1875, Exot. Butt., V,
Acraa, P|, vi, fig. 47.
One male caught at Medje in September 1910,
ye ee ee
1920] Holland, Lepidoptera of the Congo — 125
(31) 11. Acrza quirinalis Grose-Smith
Acrea quirinalis Grosr-Smiru, 1900, Nov. Zool., VII, p. 544. Smrra anp Kirey,
1901, Rhop. Exot., III, Acrea, Pl. vn, figs. 5, 6.
Two males and three females captured at Medje (June to August).
(32) 12. Acrea vesperalis Grose-Smith
Acrea vesperalis GrosE-Smrru, 1890, Proc. Zool. Soc. London, p. 466. Smrru AND
Kirpy, 1892, Rhop. Exot., I, Acrea, Pl. 11, figs. 1, 2.
_ One female taken in the first week of August 1910, at Medje.
(33) 13. Acrea pentapolis Ward
Acrea pentapolis Warp, 1871, Ent. Mo. Mag., VIII, p. 60; 1872, Afr. Lep., p. 7,
Pl. vi, fig. 2. Aurtvitiius, 1913, Seitz, Gross-Schmett., XIII, p. 257, Pl. ivte.
Two captured at Medje in June. This species is ordinarily not
common in collections, but we have recently received a very large series,
which Mr. A. I. Good informs me were part of a huge swarm which
passed a couple of years ago over Efulen in Cameroon, flying from the
north toward the south, in such numbers as to call for comment even
on the part of the natives.
(34) 13a. Acrea pentapolis thelestis (Oberthiir)
Acrea thelestis OperTHiR, 1893, Etudes d’Entomologie, XVII, p. 17, Pl. m, fig. 33.’
AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 257, Pl. uvif.
’ This is a form of the preceding species in which the white patch on
the hind wing of A. pentapolis is replaced by red. Breeding experi-
ments made at Lagos by Lamborn have shown that the two forms emerge
in about equal numbers from the chrysalids reared from the same batch
of eggs. The collection contains one specimen taken at Medje, Sep-
tember 1, 1910.
(35) 14. Acrea encedon (Linnzus)
Papilio encedon Linnxus, 1758, Syst. Nat., 10th Ed., p. 488.
Acrea encedon AvuRIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 258, Pl. tvte.
One male labelled ‘‘ Pawa, Oct. 20’; two females taken at Niangara,
one labelled “Noy. 20, 1910,” the other 1911-1912”; and a third
female taken at Faradje “1911-1912.”
(36) 14a. Acrea encedon lycia (Fabricius)
Papilio lycia Fasricius, 1775, Syst. Ent., p. 464.
Acrea encedon form lycia AurtviLuius, 1913, Seitz, Gross-Schmett., XIII, p. 258,
Pl. tvte.
One female labelled “‘ Niangara, Nov. 14-16, 1910.”
126 Bulletin American Museum of Natural History [Vol. XLIII
(37) 14b. Acrea encedon fumosa Aurivillius —
Acrea encedon aberration fumosa AuRIvILLIus, 1913, Seitz, Gross-Schmett., XIII,
p. 258.
One male, Avakubi, November 20, 1909.
(38) 14c. Acrea encedon infuscata (Staudinger)
Acrea infuscata STAUDINGER, 1885, Exot. Schmett., p. 83.
A male and a female taken at Gamangui in February, and a male
taken at Medje in March.
(39) 14d. Acrea encedon alcippina Aurivillius
Acrea encedon var. alcippina AuRIvILLIvs, 1898, Rhop. Athiop., p. 111.
Three females, one caught at Niangara in November, and two at
Faradje, “1911-1912.”
(40) 15. Acrea pharsalus Ward
Acrea pharsalus Warp, 1871, Ent. Mo. Mag., VIII, p. 81. Aurivitiius, 1913,
Seitz, Gross-Schmett., XIII, p. 258, Pl. tvid.
Eighteen males and four females, mostly taken at Medje in the
middle months of the year, but a couple were caught at Gamangui in
February and several at Niangara in November.
(41) 16. Acrea salambo Grose-Smith
Acrea salambo Grosre-Smitu, 1887, Ann. Mag. Nat. Hist., (5) XIX, p. 62. Smrrx
AND Krrpy, 1889, Rhop. Exot., I, Acrea, Pl. 11, figs. 3, 4.
Thirty-two males, mostly taken at Medje in the middle months
of the year, although a few were caught at Gamangui in February and
some at Avakubi in October.
(42) 17. Acrewa rogersi Hewitson
Acrea rogersi HewitTson, 1873, Ent. Mo. Mag., X, p. 57.
Acrea ehmckei Dewrrz, 1889, Ent. Nachr., XV, PI. 1, figs. 6-8.
There are eleven males, the localities and dates of capture being as
follows: two, Kwamouth, July 1909; one, Stanleyville, August 1909;
one, Lubila, September 1909; one, Gamangui, February 1910; three,
Medje, August 1910; three, Niangara, November 1910.
The test of breeding alone can decide whether salambo is a mere
race or form of this species. In the shape and location of the dark spots
they agree very closely. In the coloration of the upper side of the wings
A. rogersi very nearly resembles A. egina, with which, however, it has
nothing to do. The resemblance to the latter is purely superficial.
wee
1920] Holland, Lepidoptera of the Congo 127
(43) 18. Acrea althoffi rubrofasciata Aurivillius
Acrea althoffi Dewrrz, 1889, Ent. Nachr., XV, p. 102, Pl. 1, fig. 5.
Acrea althoffi var. rubrofasciata Aurtvituius, 1895, Ent. Tidskr., XVI, p. 111.
Eight males and one female taken at Medje from July to September,
and a male caught at Niangara in November.
(44) 19. Acrea oberthiiri Butler
Acrea oberthiiri Butter, 1895, Ann. Mag. Nat. Hist., (6) XVI, p. 271. AurRivILiius:
1913, Seitz, Gross-Schmett., XIII, p. 262, Pl. tvie, d.
Seven specimens taken at Medje from May to September.
(45) 19a. Acrea oberthiiri confluens Suffert
Acrea oberthiiri confluens Surrert, 1904, Iris, XVII, p. 33. Aurivititius, 1913,
Seitz, Gross-Schmett., XIII, p. 262.
Three males, one taken in each of the months of May, August,
and September at Medje.
(46) | 20. Acrea viviana Staudinger
Acrea viviana STaupINGER, 1896, Iris, LX, p. 204. Aurrvitiius, 1913, Seitz, Gross- ,
Schmett., XIII, p. 262, Pl. tvie.
Ten males and seven females, mostly from Medje, taken in the
middle of the year, but one is recorded as captured at Gamangui in
February, and several were caught at Niangara in November.
(47) 21. Acrza bonasia (Fabricius)
Papilio bonasia Fasricius, 1775, Syst. Ent., p. 464.
Acrea bonasia AuRIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 263, Pl. uv1b.
Seventy-three specimens taken mostly at Medje and Niangara at
the times when collections were made at those places, but there are also
others from various scattered localities: Faradje, Lubila, Risimu.
This species is one of the most widely diffused and commonest in the
forest-lands of central tropical Africa.
(48) 2la. Acrea bonasia supponina (Staudinger)
Acrea supponina StaupincER, 1896, Iris, IX, p. 204. Aurtvinirus, 1913, Seitz,
Gross-Schmett., XIII, p. 264, Pl. uv1b.
Of this trifling variety, which has been dignified by a name, there
are ten specimens from various scattered localities: Niangara, Faradje,
Gamangui, Medje, and Pawa.
128 Bulletin American Museum of Natural History [Vol. XLII
(49) 22. Acrea vinidia Hewitson
Acrea vinidia Hewitson, 1874, Ent. Mo. Mag., XI, p. 130. Aurrvituirus, 1913,
Seitz, Gross-Schmett., XIII, p. 264, Pl. tvia.
Eighty specimens, representing almost all the localities at which
collections were made. Forty-seven were taken at Gamangui in Febru-
ary ; thirteen at Medje (June—September) ; five at Bafwabaka in January;
four at Niangara in November; others at Bafwaboli, Batama, Stanley-
ville, Faradje, Pawa, Kwamouth, and Isangi.
Some of the specimens have dots in the discal area of the fore wings,
others do not. Here is an opportunity for a “species-maker’’to signalize
his acumen by separating the forms and counting the spots, which are
variable in number and size. The case might be referred to certain
of my friends in Berlin who used, before being otherwise employed, to
occupy themselves in making ‘‘new species” and advertising them for
sale to ardent collectors. When I was younger I often “took the hook,”
but now, as I look back, I can fancy Dame Nature smiling at the per-
formance.
(50) 23. Acrea terpsichore (Linneus)
_ Papilio terpsichore Linnxvus, 1758, Syst. Nat., 10th Ed., p. 466.
Acrea terpsichore AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 264.
There are forty-three males and eleven females in the collection.
Of this number twenty-three were taken at Medje or at localities in the
forested lands of the western and central parts of the country, Stanley-
ville, Avakubi, Munie Katoto, Bafwabaka, Kwamouth, Batama,
Gamangui. They all conform more or less exactly to the typical form
of the western coast in which the subapical black bar is complete, fully
enclosing the light subapical spot. Thirty-one were taken at Niangara
and Faradje to the northeast in more open country, and show a tendency
toward the obliteration of this band, and two specimens from Faradje,
a male and a female, are without it, the subapical spot being confluent
with the lighter area of the disk, they being therefore referable to
the subspecies buxtoni, which is the prevalent form on the eastern coast
and the grass-lands of the southeastern parts of the continent.
(51) 24. Acrwa pseudegina abadima (Ribbe)
Acraa abadima Rise, 1889, Iris, II, p. 182, Pl. rv, fig. 2.
This insect is represented in the collection by sixteen males and two
females. One was taken at Avakubi, three males and a female at Kwa-
mouth, seven males at Medje, five males and a female at Niangara. It
was originally described from the Niam-Niam country, but evidently
1920] Holland, Lepidoptera of the Congo 129
ranges far to the westward. The specimens from Kwamouth tend in the
darker coloration of the fore wings in the direction of typical A. pseude-
gina.
(52) 25. Acrea atergatis Westwood
Acrea atergatis Westwoop, 1881, Oates’ Matabeleland, p. 342, Pl. r, figs. 1, 2.
AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 268, Pl. ivf.
A single male caught at Risimu, September 6, 1909.
(53) 26. Acrea cecilia (Fabricius)
Papilio cecilia Fasricius, 1781, Spec. Ins., II, p. 34.
Acrea cecilia AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 268.
Four males and three females. Three males and two females were
caught at Niangara in November; a male and a female at Faradje
1911-1912.”
(54) 27. Acre#a acontias Westwood
Acrea acontias Wrestwoop, 1881, Oates’ Matabeleland, p. 343, Pl. Fr, figs. 7, 8.
AvRIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 274, Pl. ive (as atolmis).
Two males, taken at Kwamouth, July, 1909.
(55) 28. Acrea cepheus (Linnzeus)
Papilio cepheus Linnus, 1758, Syst. Nat., 10th Ed., p. 487.
Acrea cepheus AuURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 278, PL. tivf.
Twenty-six males and eleven females, taken principally at Medje,
but other localities, as Kwamouth, Bafwaboli, Avakubi, Niangara, and
Gamangui are represented. Captures are recorded in February and
- from June to November.
(56) 29. Acrea abdera Hewitson
Acrea abdera Hewitson, 1852, Exot. Butt., I, Acraa, Pl. 1, figs. 1,2. AuRIvILLIUs,
1913, Seitz, Gross-Schmett., XIII, p. 278, Pl. ivf.
Six males and one female, all taken at Niangara in November,
except one male, which was caught at Bafwaboli in September.
(57) 30. Acrwa perenna Doubleday and Hewitson
Acrea perenna DouBLEDAY AND Hewitson, 1848, Gen. Diurn. Lep.., I, Pl. xrx, fig. 4.
AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 279, Pl. tive.
Seventy-five males and four females, principally from Medje, but
most of the other localities mentioned in this paper are also represented;
Niangara, Bafwaboli, Gamangui, Boyulu, Lubila, and Avakubi. The
sexes are very much alike, the females being a little duller in color than
the males, and the fore wings not quite as arcuate.
130 Bulletin American Museum of Natural History {Vol. XLII
(58) 31. Acrea egina (Cramer)
Papilio egina CRAMER, 1775, Pap. Exot., I, p. 64, Pl. xxx1u, figs. F, G.
Acrea egina AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 279, Pl. trvd.
Eight males and one female. The female was captured at Gamangui
in February. The males were taken at Kwamouth, Avakubi, Munie
Katoto, Medje, and Niangara. The dates of capture run from July to
November.
(59) 32. Acrea zetes (Linnzeus)
Papilio zetes LINN=vS, 1750, Syst. Nat., 10th Ed., p. 487.
Acrea zetes AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 280. .
There are six males and two females from various localities. One
female was taken at Gamangui in February; three specimens were
caught at Medje from July to November; and three at Niangara in
November. Typical A. zetes (Linnseus) with the fore wings almost solidly
black is represented by a male taken at Risimu, September 8, 1909.
Such specimens are most often found in material from South Africa.
All the other specimens in the present collection belong to the varietal
forms A. menippe (Drury) and A. jalema Godart, which are connected
by intergrading forms.
I have in my possession a long series of specimens bred at Kangvé
on the Ogové River by the late Dr. A. C. Good. -He found the larvee
feeding gregariously, and, as specimens sent me reveal, these insects
in no respect differed from each other either in the larval or pupal stages.
The brood consisting of males and females, represented every varietal
form running from typical Acrea zetes (Linnzeus) through Acrwa menippe
(Drury) to Acrea jalema Godart. The latter, as has been pointed out .
by Professor Aurivillius, seems to constitute a connecting link between
the West African races and the East African form named Acrea acara
Hewitson. Some of the specimens of A. jalema Godart contained in
this brood very closely approximate males of Acrea acara, of which I
have many from Mombasa, with the difference that no specimens from
the French Congo show the white squamation in the middle of the hind
wing, which is characteristic of all the males of A. acara, and the sub-
apical band of the fore wing is invariably narrower than in the Kast.
African form. There is, however, a decided difference between the
females from the west coast and the east coast. The females of A. acara,
of which I possess a considerable number, are smaller in size and not as
brilliantly colored as the West African females, and the subapical white
band of the fore wings is much broader and more diffuse than in the
West African form.
1920] Holland, Lepidoptera of the Congo 131
(60) 33. Acrea insignis siginna Suffert
Acrea insignis siginna Surrert, 1904, Iris, XVII, p. 19. Aurrvitxrus, 1913, Seitz,
Gross-Schmett., XIII, p. 284, Pl. trva.
One female, Niangara, November.
(61) 34. Acrea neobule Doubleday and Hewitson
Acrea neobule DouBLEDAY AND Hewirson, 1848, Gen. Diurn. Lep., I, Pl. xrx, fig.
3. AvRIvILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 285, Pl. uurf.
Two males, Niangara, November.
(62) 35. Acrea leucographa Ribbe
Acrea leucographa Risse, 1889, Iris, II, p. 181, Pl. rv, fig. 1. Aurrviirius, 1913,
Seitz, Gross-Schmett., XIII, p. 285, Pl. ite.
A solitary male, taken at Niangara in November.
Satyride
Elymniinze
Etymniopsis Fruhstorfer
(63) 1. Elymniopsis phegea (Fabricius)
Papilio phegea Fasricivus, 1793, Ent. Syst., III, part 1, p. 132.
Elymniopsis phegea Aurivitiius, 1911, Seitz, Gross-Schmett., XIII, p. 82, Pl.
XXVIa.
One male taken at Medje, August 4, and another at Niangara
about the beginning of November 1910.
(64) 2. ivmantodats bammakoo (Westwood)
Melanitis bammakoo Wes1woop, 1851, Gen. Diurn. Lep., IT, p. 405, Pl. uxvimi, fig. 3.
Elymniopsis bammakoo Avurivituius, 1911, Seitz, Gross-Schmett., XIII, p. 82,
Pl. xxvia,
Seven specimens, six males and one female, taken at Medje from
August 5 to November 1910.
Satyrinz
Metanitis Fabricius
(65) 1. Melanitis leda africana (Fruhstorfer)
Melanitis africana FruustTorFer, 1908, Ent. Zeitschr., XXII, p. 84. AURIVILLIUS
1911, Seitz, Gross-Schmett., XIII, p. 82.
One female taken at Faradje. The only information given by the ©
label is that it was captured in “‘1911-1912.”” It belongs to the rainy
season form. The separation of this race from typical M. leda of India
is a rather meticulous refinement.
132 Bulletin American Museum of Natural History [Vol. XLIII
(66) 2. Melanitis ansorgei Rothschild
Melanitis ansorgei Roruscuitp, 1904, Nov. Zool., XI, p. 451. Aurtvinuius, 1911,
Seitz, Gross-Schmett, XIII, p. 83.
Two males taken at Medje in the first week of September 1910.
GnopHopes Westwood
(67) «1. +~Gmnophodes parmeno Doubleday and Hewitson
Gnophodes parmeno DouBLEDAY AND Hewitson, 1851, Gen. Diurn. Lep., IT, p. 363,
Pl. uxt, fig. 2, 7. AurRrtvi~irus, 1911, Seitz, Gross-Schmett., XIII, p. 83, Pl.
xxvib.
One damaged male taken at Stanleyville on September 16 and a
better preserved female (lacking antennz) taken at Medje in the early
part of September 1910.
(68) 2. Gmnophodes chelys (Fabricius)
Papilio chelys Fasrictus, 1793, Ent. Syst., III, part 1, p. 80.
Gnophodes chelys AurtviLutus, 1911, Seitz, Gross-Schmett., XIII, p. 83, Pl. xxvie.
Two males and five females all captured at Medje, the dates ran-
ging from July 19 to the first week of September 1910.
Bicyctus Kirby
(69) 1. Bicyclus hewitsoni nanodes (Grose-Smith)
Idiomorphus nanodes Grose-Smitu, 1890, Proc. Zool. Soc. London, p. 472.
Mycalesis hewitsoni var. nanodes AurtviLLIus, 1911, Seitz, Gross-Schmett., XIII,
p. 84, Pl. xxvid.
Structurally not capable of being separated from B. hewitsoni.
(Doumet) (cf. Rev. Zool., 1861, (2) XIII, p.175) but on the upper side the
submarginal ocelli are more or less obsolete, and the outer pale area of
the wings is laved with violaceous. The latter also holds true of many
specimens of typical B. hewitsoni, which are not all pale brown on this
part of the wings. The line of demarcation between the dark basal
area of the hind wing and the light outer area is not quite straight as in
the typical form but is bent slightly inwardly basad at a point just
beyond the end of the cell.
Ten males, all taken at Medje, except one which is recorded as
from Faradje, but without any indication of the day or month. The
specimens from Medje, with the exception of one taken on April 4,
were captured at dates ranging from July 7 to September 24, 1910.
(70) 2. Bicyclus medontias obgoletus, new variety
Plate VII, Figure 1 ,o@*
On the upper side hardly to be distinguished from typical B. medontias (Hewitson)
(ef. Exot, Butt., 1874, Mycalesis, Pl. rx, figs. 56, 57), except that the ground-color
1920] Holland, Lepidoptera of the Congo 133
in both sexes is a little paler. On the under side the median line of both wings is
perfectly straight, and not at all curved, as it is in most specimens of B. medontias
and only half, or even less than half, as wide as it is in the typical form, being reduced
to a very narrow, sharp, creamy white line, contrasting strongly against the darker
ground-color. All the ocelli are greatly reduced in size, some of them being occasion-
ally obsolete. This may be a wet seasonal form of B. medontias.
Types, o' and 9, in The American Museum of Natural History; paratypes in
the Holland Collection, Carnegie Muesum.
The collection contains twenty-five males and eight females, all
taken at Medje, the dates of capture ranging from July 17 to the middle
of September 1910.
(71) 3. Bicyclus iccius (Hewitson)
Idiomorphus iccius Hewrrson, 1865, Exot. Butt., III, [diomorphus, Pl. 1, figs. 4% 5.
Mycalesis iccius AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 85, Pl. xxvid.
Seven males and one female, the latter taken at Medje, July 21,
1910. Of the males three were taken in April, three in August, and one
in September.
(72) , 4. Bicyclus sebetus (Hewitson)
Idiomorphus sebetus Hewirson, 1877, Exot. Butt., V, Pl. Mycalesis and Idiomorphus,
figs. 6, 7.
Mycalesis sebetus AuRIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 85.
Four males and three females taken at Medje, the dates of capture
ranging from June 6 to August 6, 1910.
Mycatesis Hiibner
(73) 1. Mycalesis xeneas Hewitson
Mycalesis xeneas Hewitson, 1865, Exot. Butt., III, Mycalesis, Pl. vu, fig. 48.
Avrivituivs, 1911, Seitz, Gross-Schmett., XIII, p. 86, Pl. xxvira.
The figure given by Hewitson is very poor; that given in Seitz is
much better.
The collection contains five males and six females taken at Medje
from July 9 to the first week in September and one female taken at
Gamangui, June 17, 1910.
(74) 2. Mycalesis analis Aurivillius
Mycalesis analis Aurtvituius, 1895, Ent. Aiea, XVI, p. 113, fig. 1; 1911, Seitz,
Gross-Schmett., XIII, p. 87.
There are fourteen males and three females, all taken at Medje
from July 7 to August 24, 1910.
134 Bulletin American Museum of Natural History [Vol. XLIII
(75) 3. Mycalesis tenias Hewitson
Mycalesis tenias Hewitson, 1877, Exot. Butt., V, Pl. Mycalesis and Idiomorphus,
fig. 66. AurriviLiius, 1911, Seitz, Gross-Schmett., XIII, p.87, Pl. xxvub.
Represented by a single male specimen taken at Niangara and
belonging to the lot of insects collected at that place between November
26 and December 2, 1910.
(76) 4. Mycalesis ignobilis Butler
Mycalesis ignobilis Butter, 1870, Trans. Ent. Soc. London, p. 124; 1871, Lep.
Exot., p. 55, Pl. xxi, fig. 4. Aurtvitiius, 1911, Seitz, Gross-Schmett., XIII,
p. 87, Pl. xxvub. ;
This species, which is quite common in the Ogové valley, is re-
presented in the present collection by five males and a female which
were taken at Medje in September 1910.
(77) 5. Mycalesis asochis Hewitson
Mycalesis asochis Hewitson, 1865, Exot. Butt., III, Mycalesis, Pl. vu, figs. 46, 47.
AuRIVILLIvs, 1911, Seitz, Gross-Schmett., XIII, p. 88, Pl. xxvuic.
Of this not uncommon species there are four males and one female.
One male was captured at Gamangui, June 19, two were taken at
Medje in July and August, the female was taken at the same place on
August 6, and one male is ticketted as from ‘“Niangara, November
to December 4, 1910.” From this record it appears that the insect
must have at least two broods annually.
(78) 6. Mycalesis sambulos Hewitson
Mycalesis sambulos Hewrtson, 1877, Exot. Butt., V, Pl. Mycalesis and Idiomorphus,
figs. 63, 64. Avurivitiius, 1911, Seitz, Gross-Schmett., XIII, p. 89, Pl. xxvue.
There are two males in the collection, both taken at Medje, one on
July 12, the other on August 24, 1910.
(79) 7. Mycalesis mandanes Hewitson
Mycalesis mandanes Hewrrson, 1873, Exot. Butt., V, Mycalesis, Pl. rx, figs. 61, 62.
Avurivit.ies, 1911, Seitz, Gross-Schmett., XIII, p. 89, Pl. xxvite.
Professor Aurivillius in his ‘Rhopalocera A®thiopica,’ p. 53, sank
this species as a synonym of M. auricruda Butler, but in Seitz, ‘Die
Gross-Schmetterlinge des Afrikanischen Faunengebietes,’ p. 89, he
restores it to its rightful place as a distinct species. The outline of the
hind wings, which are much more elongate than in M. auricruda, is
enough, without the many differences in the markings of the under side
of the wings, to reveal its distinctness.
Two males taken at Medje in July 1910.
1920] Holland, Lepidoptera of the Congo 135
(80) 8. Mycalesis auricruda Butler
- Mycalesis auricruda Butimr, 1868, Cat. Satyr., p. 131, Pl. m, fig. 6. AuRIvILLIUs,
1911, Seitz, Gross-Schmett., XTIT, p. 89, Pl. xxviie.
Six males and four females taken at Medje at dates ranging from
July 8 to September 6, 1910.
(81) 9. Mycalesis uniformis Bethune-Baker (?)
Mycalesis uniformis BeTHUNE-BAkER, 1908, Ann. Mag. Nat. Hist., (8) II, p. 470.
AvrRrivituivs, 1911, Seitz, Gross-Schmett., XTII., p. 90.
I refer to this species two males and three females before me, which
agree better with the description of this species given by Bethune-
Baker than with any other known to me. I was at first tempted to
describe them as new to science but, after considerable study, have
finally decided that they belong here, but cannot be quite positive, as I
have not had the opportunity to examine the type of the species, which
isamale. The females in the collection agree absolutely with the males
in the markings of both the upper and under sides and are only differen-
tiated from them, aside from their sexual characters, by having a greater
expanse of wing.
The specimens were taken at Medje, four in August, and one female
in September.
(82) 10. Mycalesis sandace Hewitson
Mycalesis sandace Hewitson, 1877, Exot. Butt., V, Pl. Mycalesis and Idiomorphus,
‘fig. 65. Aurrtvitiius, 1911, Seitz, Gross-Schmett., XIII, p. 89, Pl. xxvud.
There is a series of ten males, nine taken at Medje from June 6 to
September 20, and one at Gamangui, February 5; and three females,
one caught on April 6, the others about the middle of August 1910.
None of the specimens are in good condition, all being more or less
rubbed and defective.
(83) : 11. Mycalesis miriam (Fabricius)
Papilio miriam Fasrictivus, 1793, Ent. Syst., III, part 1, p. 242.
Mycalesis miriam AurRtviLuius, 1911, Seitz, Gross-Schmett., XIII, p. 90, Pl. xxviid.
One female caught at Medje, September 1910, and nine males and
another female taken at Faradje, five labelled as taken in December, the |
rest ticketted 1911-1912.”
(84) 12. Mycalesis melusina (Fabricius)
Papilio melusina Fasrictus, 1787, Mantissa Ins., II, p. 43.
Mycalesis melusina AuRIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 90, Pl. xxviid.
Eleven males and one female caught at Medje, two in April, the
rest from July to September 1910.
136 Bulletin American Museum of Natural History [Vol. XLIII
(85) 13. Mycalesis sophrosyne Plotz
Mycalesis sophrosyne Pucerz, 1880, Stett. Ent. Zeit., XLI, p. 196. AuRtviuuivs,
1911, Seitz, Gross-Schmett., XITI, p. 90.
Three males taken at Medje, one in June, the others in July.
(86) 14. Mycalesis mollitia Karsch
Mycalesis mollitia Karscu, 1895, Ent. Nachr., XXI, p. 281. Aurtviturus, 1911,
Seitz, Gross-Schmett., XIII, p. 90, Pl. xxvure.
; Karsch described the male; the female was described by Wichgraf
in the Deutsche Entomologische Zeitschrift, 1914, p. 345. It is some-
what larger than the male, paler in color throughout, with the sub-
apical light area of the primaries enlarged and extending along the
costa as a light line to a point about opposite the end of the cell. On the
under side it does not differ materially in its color and markings from the
male.
The collection contains a set of seven males and five females, taken
at Medje, one male and three females in April, the rest in July and
August.
(87) 15. Mycalesis obscura Aurivillius
Mycalesis obscura AuRIvILLIUvs, 1901, Ent. Tidskr., XXII, p. 114; 1911, Seitz, Gross-
Schmett., XIII, p. 91, Pl. xxvina.
I refer to this species a long series of males and several females
the general resemblance of which on the under side of the wings to M.
martius and M. golo recalls the remarks of the author of the species,
who says that M. obscura is very like the two which have been mentioned.
There are, however, structural differences which clearly separate this
species from M. martius. One of these is the absence in M. obscura of
the tuft of androconia, which is always found in M. martius between
veins 2 and 3 near the point of their origin on the fore wing, and
another is the fact that the hind wing is not produced or angulated at the
extremity of vein 4 as is the case in M. martius. From M. golo they
may at once be discriminated by their more uniformly dark color on
the under side of the secondaries. The females which I associate with
the males have broader wings and are paler in color than the males,
but, like the males, strongly recall in the pattern of the markings the
corresponding sex of M. martius.
Of M. martius I have before me, as I write, an enormous series of
males and females taken at various localities, ranging from the valley
of the Ogové River to Lolodorf in Cameroon and, while superficially
the two species are very much alike, so that an untrained investigator
1920] Holland, Lepidoptera of the Congo 137
might at first glance easily confound the two, the structural differences
which I have pointed out enable them to be easily separated. M.
martius is not represented by a single specimen in the collection upon
which I am reporting.
The seventeen males and six females in the collection were all taken
at Medje, a couple in April, the rest from July to September.
(88) 16. Mycalesis dubia Aurivillius
Mycalesis dubia Aurivittrus, 1893, Ent. Tidskr., XIV, p. 270, fig. 4; 1911, Seitz,
Gross-Schmett., XIII, p. 92, Pl. xxvub.
There are two males of this species, one taken at Medje in April
and another at the same place in September.
(89) 17. Mycalesis golo Aurivillius
Mycalesis golo AuRIvVILLIUS, 1893, Ent. Tidskr., XIV, p. 267, fig. 2.
Mycalesis golo var violascens AURIVILLIUS, 1898, Rhop. A®thiop., p. 55; 1911, Seitz,
Gross-Schmett., XXIII, p. 91, Pl. xxvite.
Mycalesis golo was originally described by Professor Aurivillius
from Cameroon. I have in my possession a large series of specimens
from that region, which agree exactly with his description and figure.
The series taken by the American Museum Congo Expedition are not
typical, but may in part be referred to the varietal form violascens, in
which the limbal area of the wings is more or less suffused with violet
and the transverse line of the hind wing is obscured by the outward ex-
tension of the dark color of the basal and discal area of the wing. Some
of the specimens taken at Medje approach more nearly to the typical
form but these differ, nevertheless, by having the upper surface of the
wings distinctly brown and not dark blackish brown like the insect
from Cameroon. This difference, while it holds good of all the specimens
not referable to the variety violascens, hardly seems to me to be of
sufficient importance to justify the erection of a subspecies.
Of M. golo violascens there are in the collection seventeen males,
one taken at Niangara in November, the rest caught at Medje from
July to September. Of the reddish form there are eight males taken at
Medje in August and September.
(90) 18. Mycalesis baumanni Karsch (?)
Mycalesis baumanni Karscu, 1894, Ent. Nachr., XX, p. 230. Aurtvitiius, 1911,
Seitz, Gross-Schmett., p. 91, Pl. xxvue.
With a good deal of hesitation I refer to this species five males which
are before me. They belong to the Fourth Subgroup of the classifica-
tion proposed by Professor Aurivillius (cf. Seitz, ‘Die Gross-Schmetter-
138 Bulletin American Museum of Natural History [Vol. XLIII
linge des Afrikanischen Faunengebietes,’ p. 91), but they differ from M.
baumannii in not having the transverse bands on the under side of the
fore and hind wings strongly produced on vein 4. This band has its
outer margin more or less irregular, and its course recalls that of the
band on the under side of M. sandace Hewitson, with which: the speci-
mens otherwise have nothing to do, as is shown by the sexual brands
and the entire absence of ocelli on the upper side of the primaries.
The specimens are not, for the most part, very well preserved, some
of them being rather badly rubbed, making the determination of the
exact pattern of the markings a little difficult to determine, but I do
not feel like hazarding their description as a new species, though eventu-
ally they may turn out to be undescribed.
The specimens were taken at Medje, one in April, the others ~
dates ranging from July to September.
(91) 19. Mycalesis milyas Hewitson (?)
Mycalesis milyas Hewitson, 1864, Exot. Butt., III, Mycalesis, Pl. v1, fig. 34.
AuvRIvILuivs, 1911, Seitz, Gross-Schmett., XIII, p. 92, Pl. xxvug.
I refer with great doubt a rubbed male and a rather poorly preserved
female to this species. The male was taken at Medje in September
1910, the female in July.
The description and figure of M. milyas given by Hewitson
(loc. cit.) leave much to be desired. The figure given by Aurivillius on
Plate xxvir of “Die Gross-Schmetterlinge des Afrikanischen Faunen-
gebietes’ more nearly represents the under side of the specimens under
consideration, but his description does not agree either with the figure
he gives or the specimens before me. I have in my possession a male
taken at Efulen by A. C. Good, and there are in the Carnegie Museum
some specimens taken by A. C. Good at Lolodorf, which are identical
with the insect taken at Medje. I have assigned all of them to M,
milyas with a double interrogation mark, as they do not perfectly agree
either with the descriptions or figures hitherto given by authors, al-
though they come nearer to that species than to any other.
(92) 20. Mycalesis pavonis Butler
Mycalesia pavonis Butter, 1876, Ann. Mag. Nat. Hist., (4) XVIII, p. 481.
Avmiviniies, 1911, Seitz, Gross-Schmett., XIII, p. 92.
There are seven males and three females of this well-marked species;
all taken at Faradje in November and December.
1920[ Holland, Lepidoptera of the Congo 139
(93) 21. Mycalesis desolata Butler
Mycalesis desolata Butter, 1876, Ann. Mag. Nat. Hist., (4) XVIII, p. 480.
Mycalesis leptoglena Karscu, 1893, Berl. Ent. Zeit., XX XVIII, p. 208, PI. v, fig. 7.
Mycalesis desolata AuRiviL.LIus, 1911, Seitz, Gross-Schmett., XIII, p. 93.
Of this species there are five males, four of which agree absolutely
with the published descriptions and figure, but the fifth has the sub-
marginal ocelli much larger, with round black centers pupilled with
white, contrasting with the dull ground-color of the wings in a very
striking manner. In the typical form the ocelli are more or less obsolete.
The specimens were all taken at Niangara in November.
(94) 22. Mycalesis safitza ethiops Rothschild and Jordan ie
Mycalesis safitza ethiops RotHSCHILD AND JORDAN, 1905, Nov. Zool.,XII, p. 175.
M. safitza was described and figured by Hewitson (cf. Gen. Diurn.
Lep., 1851, II, p. 394, note, Pl. uxv1, fig. 3). Of the form ethiops there
are fifteen males and two females, all of which were taken at Faradje
and Niangara in November and December. There are also two males
and seven females taken at Medje in the months of July and August,
which are so much like the others in their markings that it is impossible
to separate them, though they are somewhat larger in size.
(95) 22a. Mycalesis safitza evenus Scary 9
Mycalesis evenus Hoprrer, 1855, Monatsber. Akad. Wiss. Berlin, p. 641; 1862,
Peters, Reise n. Mossambique, Ins. ,p 394, Pl. xxv, figs. 5,6. AuRIvILiius, 1911,
Seitz, Gross-Schmett., p. 93, Pl. xxvmie.
Of this, the dry-season form of M. safitza, there are two males and
two females. One of the males was taken at Medje in August, the other
at Faradje in November. Both of the females were captured at Niangara
in November.
(96) 23. Mycalesis langi, new species
Plate X, Figure 10, @
3&. The fore wing with a small, but distinct, sexual brand at the middle of
vein 1, which at this point is slightly bent costad. The hind wing with a pale yelfowish
brown pencil of hairs on the upper margin of the cell about the middle and beyond it with
a black tuft of hairs on the sixth interspace. The upper side of both wings is totally
devoid of all ocelli. The prevalent color is mouse-gray, the discal area of the fore
wings being black and velvety, this black area covering the end of the cell and the
origin of the submedian nervules, extending to the inner margin from near the outer
angle to within one-third of the distance from the base. The hind wings also are
black, or very dark brown, except on the outer and inner margins. Both wings have a
very fine black marginal line, paralleled inwardly by a similar fine submarginal
line, separated from the outer line by a space less than half a millimeter in diameter.
140 Bulletin American Museum of Natural History [Vol. XLIII
The fringes are obscurely checkered at the ends of the nervules by darker scales,
contrasting with the pale mouse-gray color of the rest of the fringes. On the under
side the markings recall those of M. safitza, but the ground-color is darker. Expanse,
37mm. (The expanse of the smallest specimen of M. safitza before the writer—there
are many scores of them—is 45 mm.)
Type from Medje; paratypes, from Faradje. Type and two paratypes in The
American Museum of Natural History, New York; two paratypes in the Holland
Collection in the Carnegie Museum.
This may prove to be a local or seasonal form of M. safitza, but it
differs so markedly from all the varieties of that species known to the
writer that he does not hesitate to describe it as new. The entire absence
of any trace of ocelli on the upper side of the wings, the black velvety
color of the central area of the fore wings, the checkered fringes, and the
uniformly small size of the specimens, all indicate that we are dealing
with what is at all events a marked variety, separable at a glance from
M. safitza and all the varieties of that species hitherto described.
I have the honor of naming the species after Mr. Herbert Lang, the
capable head of the Expedition, to whose enthusiasm we are indebted
for the large entomological collections which were returned.
(97) 24. Mycalesis chapini, new species
Plate VII, Figure 9, @
o. On the upper side having a general resemblance to the male of M. langi
Holland, but considerably larger in size, and having the fringes of the wings more
decidedly checkered with dark at the ends of the nervules. On the under side wholly
different from M. langi and in the disposition of the markings, but not their color,
recalling the under side of M. bawmanni Karsch. The ground-color of the under
side is pale sienna sprinkled with minute dark transverse lines and dots, the basal
half of both wings and the limbal area of the primaries near the outer angle broadly
clouded with purplish brown. A sub-basal and median transverse line, curving
outwardly from the costa of the fore wing to the inner margin of the hind wing about
its middle, run approximately parallel to each other. The median line is produced
somewhat sharply at the origin of vein 4 on each wing, and is more or less waved on
the interspaces, the curves bending outwardly. The space between the sub-basal
and median lines is a trifle darker than the rest of the wings, partly due to the increase
in this field of the minute strigule with which the wings are strewn. The ocelli are,
for tht most part, entirely obsolete, at most being represented by minute white dots,
except the two nearest the anal angle of the secondaries, which in the type are exceed-
ingly minute black circles, under the glass seen to be pupilled with white and ringed
about with pale ochreous, Expanse, 42 mm.
Niangara. Type unique, in The American Museum of Natural History.
This insect is so totally unlike any other in the group to which it
belongs that I do not hesitate to describe it as new. I take pleasure in
naming it in honor of Mr. J. P. Chapin, one of the leaders of the Expe-
dition.
1920] Holland, Lepidoptera of the Congo 141
(98) 25. Mycalesis vulgaris Butler
Mycalesis vulgaris Butuer, 1868, Cat. Satyr., p. 130, Pl. m, fig. 8. AuRIVILLIUs,
1911, Seitz, Gross-Schmett., XIII, p. 95, Pl. xxviig.
A single male taken at Niangara, November 26, 1910.
(99) 25a. Mycalesis vulgaris tolosa (Platz)
Mycalesis tolosa Puarz, 1880, Stett. Ent. Zeit., XLI, p. 197.
This is the dry-season form of this species, and is characterized by
having the ocelli well developed and not reduced to mere points, as is
the case in typical M. vulgaris. It is the prevailing form in the region
of the Ogové River, in Cameroon, and in the Belgian Congo. In spite
of its name, typical M. vulgaris is not very common in collections, so far
as my observations show. ye
Of the variety tolosa (Ploetz) the collection upon which I am report-
ing contains three males and eight females. One female was taken at
Medje in July, one at Lubila in September, three at Niangara in Nov-
ember, and.three at Faradje in December. One of the males was caught
at Niangara in November, and the other two at Faradje in December.
(100) 26. Mycalesis nebulosa Felder
Mycalesis nebulosa FuupEerR, 1867, Reise Novara, Lep., III, p. 502. Aurivriiius,
1911, Seitz, Gross-Schmett., XIII, p. 95, Pl. xxvuf.
There are two poorly preserved males and one female of this
species, which were taken at Niangara in November.
(101) 27. Mycalesis agraphis Karsch
Mycalesis agraphis Karscu, 1893, Berl. Ent. Zeit., XX XVIII, p. 207. Aurrviturus,
1911, Seitz, Gross-Schmett., XIII, p. 96, Pl. xxvurf.
A single male of this common West African species caught at
Avakubi in November.
Henoresia Butler
(102) 1. Henotesia perspicua (Trimen)
Mycalesis perspicua TRIMEN, 1873, Trans. Ent. Soc. London, p. 104, PI. 1, fig. 3.
Henotesia perspicua AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 97, Pl. xxvimia,
Five males: one was taken at Risimu in September; two at Niangara
in November; and two at Faradje, one in December, the other without
date.
(103) 2. Henotesia phea (Karsch)
Mycalesis phea Karscu, 1894, Ent. Nachr., XX, p. 232.
Henotesia phea Aurtvi1Luivs, 1911, Seitz, Gross-Schmett., p. 98, Pl. xxvimb.
Four males taken at Faradje, one on December 12, 1912, the other
three ticketted 1911-1912.”
142 Bulletin American Museum of Natural History [Vol. XLII
(104) 3. Henotesia eliasis (Hewitson)
Mycalesis eliasis HEwitson, 1866, Exot. Butt., III, Mycalesis, Pl. vu, figs. 44, 45.
Henotesia eliasis AuRtv1Lutvs, 1911, Seitz, Gross-Schmett., XIII, p. 99, Pl. xxvinid.
A single damaged male labelled ‘“ Kwamouth, July 14, 1909.”
(105) 4. Henotesia peitho (Pletz)
Mycalesis peitho Pucerz, 1880, Stett. Ent. Zeit., XLI, p. 197.
Henotesia peitho AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 99, Pl. xxvima.
Four males taken at Medje, one in April, two in August, and one in
September.
Yrruma Hiibner
(106) 1. Ypthima asterope (Klug)
Hipparchia asterope Kiva, 1832, Symb. Phys., Pl. xxrx, figs. 11-14.
Y pthima asterope AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 115, Pl. xxrxc.
One male taken at Niangara, November 25, 1910.
(107) 2. Ypthima simplicia Butler
Ypthima simplicia BuTuER, 1876, Ann. Mag. Nat. Hist., (4) XVIII, p. 481.
AuRIvILuivs, 1911, Seitz, Gross-Schmett., XIII, p. 115.
A poorly preserved male caught at Faradje and labelled “1911-1912.”
(108) 3. Ypthima doleta Kirby
Ypthima doleta Kirpy, 1880, Proc. Roy. Dublin Soe., (2) II, p. 336. AurtviLiius,
1911, Seitz, Gross-Schmett., XIII, p. 115, Pl. xxrxe.
Of the fourteen males one is labelled as from Bafwasende, September
23, 1910; all the rest were taken at Medje, two in April, the rest from
July to September. Of the two females one was caught at Medje in
August, the other at Niangara in November.
(109) 4. Ypthima itonia Hewitson
Y pthima itonia Hewrrson, 1864, Trans. Ent. Soc. London, p. 287, Pl. xvi, fig. 138.
AuRIVILLIvs, 1911, Seitz, Gross-Schmett., XIII, p. 116.
Six males taken at Faradje in November.
Nymphalidez
Argynnidinsz
LacunoprTera Doubleday
(110) 1. Lachnoptera iole (Iabricius)
Papilio iole Fannicivs, 1781, Spec. Ins., II, p. 78.
Lachnoptera iole Auniviti108, 1913, Seitz, Gross-Schmett., XIII, p. 280, Pl. xan.
This species is represented in the collection by twelve males and one
female of the dimorphic form named L. hecatwa by Hewitson. All of the
_ specimens are from Medje (June-September) except one male specimen
captured at Gamangui in June.
1920] Holland, Lepidoptera of the Congo 143
Atretta Doubleday
(111) 1. Atella columbina (Cramer)
Papilio columbina Cramer, 1779, Pap. Exot., III, p. 76, Pl. ccxxxvmt, figs. A, B.
Atella columbina AuriviLuivs, 1913, Seitz, Gross-Schmett., XIII, p. 230, Pl. ua.
Twenty-six specimens; six taken at Gamangui in February; one
at Lubila in September 1909; eight at Niangara in November; and the
rest at Medje, one in April, the others from July to September 1910.
(112) 2. Atella phalantha xthiopica Rothschild and Jordan
Atella phalantha form ethiopica RotuscuiLp AND JORDAN, 1903, Nov. Zool., X,
p. 505.
Twenty-eight specimens, two of which were taken at Medje in
July, the rest at Niangara in November. The species was originally
described from the East Indies, and is figured by Drury (1773, Ill.
Exot. Ent., I, p. 41, Pl. xx1, figs. 1, 2). Its range covers the tropical
regions of the Eastern Hemisphere and it is one of the most widely
distributed insects of the Old World. The separation of the race found
in Africa from the oriental varieties is a recent refinement in classifica-
tion which is justifiable, although founded on differences which are
recognizable, but very slight.
Vanessine
AnTanarRTi1A Rothschild and Jordan
(113) 1. Antanartia delius (Drury)
Papilio delius Drury, 1782, Ill. Exot. Ent., ITI, p. 18, Pl. xrv, figs. 5, 6.
Hypanartia delius AuRIvILLIUS, 1898, Rhop. Aithiop., p. 130.
Antanartia delius Roruscuizp AnD JorDAN, 1903, Nov. Zool., X, p. 508.
AuRIVILLIus, 1913, Seitz, Gross-Schmett., XIII,p. 229, Pl. x1c.
Fifty-three specimens, mostly males, principally taken at Medje
from June to September, but there are a couple caught at Niangara in
November and one captured at Munie Katoto in September.
Pyrametis Hiibner
(114) 1. Pyrameis cardui (Linnzus)
Papilio cardui Linnxus, 1758, Syst. Nat., 10th Ed., p. 475.
Pyrameis cardui AuRIVILLIUs, 1913, Seitz, Gross-Schmett., XIII, p. 227.
This cosmopolitan species is represented in the collection by three
examples, two taken at Niangara, and one at Faradje, 1911-1912.”
It is worthy of note that no specimens were taken at Medje or other
localities in the hot, humid tropical forest-lands. The insect does not
144 Bulletin American Museum of Natural History [Vol. XLITI
appear to turn up in the dense jungle of the hottest parts of Africa,
but seems to be confined to the more open lands, where thistles grow.
In the more than forty years in which I have been receiving collections
from tropical Africa I never have obtained specimens of ‘‘The Painted
Lady” from such places as the Ogové Valley, or the swampy palm-clad
savannas along the big rivers; but, on the other hand, it has often been
received from the sandy coastal ridges, and the higher grass-lands of both
the western, the eastern, and the southern parts of the continent.
VANEssULA Dewitz
(115) 1. Vanessula milca (Hewitson)
Liptena milca Hewitson, 1873, Exot. Butt., V, Pentila and Liptena, Pl. 1, fig. 17.
Vanessula milca AURIVILLIUS, 1913, Seitz, Gross-Schmett.,/XIII, p. 227, Pl. Lie.
Nine specimens captured at Medje from ‘July to September.
This pretty little insect is very common in the ‘valley of the Ogové,
and apparently swarms in southern Cameroon.
Junonia Hiibner
I deem it more natural to associate under the term Junonia a
number of species which for many years past have been placed in this
genus, but which Dr. Aurivillius in his recent writings has incorporated
into the genus Precis.
(116) 1. Junonia orithya madagascariensis Guenée
Junonia orithya \ar. madagascariensis GuentE, 1865, Vinson’s Voy. Madgr., Lep.,
. a7.
Beds orithya AuRIvILLIvs, 1913, Seitz, Gross-Schmett., XIII, p. 226.
This is the African form of the insect originally named from the
East Indies by Linnzus (cf. Mus. Ulr., 1764, p. 278). The collection
contains thirteen males and one female, all taken at Faradje and
Niangara in November 1910, or else labelled “1911-1912.”
(117) 2. Junonia clelia (Cramer)
Papilio clelia Cramer, 1775, Pap. Exot., I, p. 23, Pl. xx1, figs. 2, F.
Precis clelia AuRiViLLi108, 1913, Seitz, Gross-Schmett., XIII, p. 226, Pl. 1a.
Twenty-three males and fifteen females were taken. Two males and
one female were caught at Kwamouth in July 1909. Two males and
one female were captured at Pawa, October 1910. Eight males and
four females are labelled as taken at Medje from April to August 1910.
Eleven males and nine females were caught at Niangara in November of
the latter year.
1920] Holland, Lepidoptera of the Congo 145
(118) 3. Junonia cebrene Trimen
Junonia cebrene TRIMEN, 1870, Trans. Ent. Soc. London, p. 353.
Precis cebrene AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 226, Pl. xia.
This is the African race of J. enone (Linnzeus) originally described
from Asiatic specimens, which do not materially differ from the African
form, except that in the latter the large yellow area of the primaries is
somewhat more restricted, and the blue spot on the hind wing is rounder,
and not oval as in the Asiatic specimens, a long suite of which is before
me as I write. Eighteen males, six females, all captured at Niangara
and Faradje in November 1910.
(119) 4. Junonia westermanni Westwood
Junonia westermanni Westwoop, 1870, Ent. Mo. Mag., VI, p. 278; 1874, Thes.
Ent. Oxon., p. 182, Pl. xxxrv, fig. 7, o’, fig. 8, @.
Precis westermanni AuRIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 225, Pl. xia.
Of this lovely insect fifty-three males and one female are contained
in the collection. Most of them were caught at Medje, the dates of cap-
ture ranging from April to August. There are two males taken at
Gamangui in February and two others taken at the same place in June.
There is one male captured at Ngayu in April. The extreme rarity of
the female in collections is illustrated in the present case.
(120) 5. Junonia sophia (Fabricius)
Papilio sophia Fasricrus, 1793, Ent. Syst., III, part 1, p. 248.
Precis sophia Auriviuutus, 1913, Seitz, Gross-Schmett., XIII, p. 225, Pl. i1b.
Though very dissimilar in its style of markings from most species
of the genus, sophia is a true Junonia, as is shown by the neuration,
the outline of the wings, and the structure of the antenne and palpi.
Of this insect, the smallest of the genus, which superficially on the
upper side bears a likeness to the female of the preceding species, there
are nine males and five females. One male and the five females were
taken at Medje, from June to August 1910. Seven males are ticketted
as taken at Niangara and Faradje in November 1910, or else ‘‘1911—
1912.”
Precis Hiibner
Dr. Aurivillius in his latest account of the butterflies belonging to
this group (ef. Seitz, ‘Gross-Schmetterlinge der Erde,’ XIII, pp. 218-
227), has placed in the genus Precis a number of forms which for many
years past have been by most writers classified in the genus Junonia
Hiibner. The type of the genus Precis is P. octavia (Cramer). The type
146 Bulletin American Museum of Natural History (Vol. XLIII
of the genus Junonia is J. lavinia (Cramer). It requires no effort to
detect the fact that the species included by Aurivillius in his “Fifth
Group” of the genus Precis, except P. octavia and allies, viz., clelia
(Cramer), orithya (Linneeus), westermanni Westwood, and their allies are in
structure, form, and markings more nearly related to J. lavinia than to P.
octavia, the type of the genus, and that they are widely different from
the Kallima-like insects which, especially in their wet-season forms, show
on the under side of their wings close resemblance to dried leaves.
In the present paper I have restored the species of the clelia-awnone
Group to the genus Junonia. I also separate from the genus Precis the
long-tailed Kallima-like insects, which on the under side of the wings’
show mimetic resemblance to dried leaves, and in which the outer
border of the hind wing is evenly curved opposite the end of the cell,
not angulated or toothed, and greatly elongated, or tailed at the anal
angle. For this group of insects I propose the generic name Kallimula.
As restricted in the present paper, the genus Precis may be divided
into groups as follows.
A. Hind wings greatly expanded opposite end of cell, not greatly produced at
anal angle.
1. Outer margin of hind wing crenulated: P. octavia (type of genus)
and allies; P. archesia and allies.
2. Outer margin of hind wings entire, evenly rounded, not crenulated:
P. artaxia and allies.
B. Hind wings angulated and toothed opposite the end of the cell.
1. Anal angle not greatly produced: P. chorimene and allies; P. terea
and allies; P. ceryne and allies; P. rhadama and varieties.
2. Anal angle greatly produced: P. andremiaja and allies.
(121) : 1. Precis octavia (Cramer)
Papilio octavia Cramer, 1777, Pap. Exot., I, p. 60, Pl. cxxxv, figs. B, C.
Precis octavia Aurivit.ivs, 1913, Seitz, Gross-Schmett., XIII, p. 224, Pl. rib.
Thirty-six males and thirty-five females. One male was caught at
Kwamouth, July 14, 1909; three females were taken at Medje in August;
and all the rest were obtained at Niangara in November 1910, except a
solitary individual, which is labelled as caught at Faradje in the same
month and year.
This is the rainy-season form of the species in the northwestern part
of the range, and the capture of these specimens shows that the region
about the headwaters of the Uelle River in the Niam-niam country is
faunistically allied to the country of the upper waters of the Niger.
1920] Holland, Lepidoptera of the Congo 147
(122) la. Precis octavia amestris (Drury)!
Papilio amestris Drury, 1782, Ill. Exot. Ent., III, p. 26, Pl. xx, figs. 3, 4.
Precis amestris AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 224.
There are twenty males and eleven females of this, the dry-season
form of P. octavia. They were all taken at Niangara in November
1910, except two of the males, which were caught at Medje in August.
In the southern and eastern parts of the range of this insect amestris
is replaced by the form sesamus Trimen, the upper side of the wings of
which are much bluer than in this form, and do not have the red spots in
the cell of the fore wings, which are always found in the form under con-
sideration. We have specimens of P. octavia sesamus from Natal and
Mozambique.
(123) 2. Precis chorimene (Guérin)
Vanessa chorimene Guirin, 1844, Icon. Régne Anim., Ins., p. 476.
Salamis ethyra Fris. HAMEL, 1850, Ann. Soc. Ent. France, (2) VIII, p. 250.
Precis chorimene AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 220, Pl. xub.
Four males and two females. One male was taken at Faradje,
“1911-1912”; three at Niangara in November 1910; one female was
caught at Pawa, October 19, the other at Niangara on November 25,
1910. Ena
(124) 3. Precis stygia Aurivillius
Precis ethyra StauDINGER, 1883, Exot. Schmett., I, p. 102, Pl. xxxvim (non ethyra
Feisthamel = chorimene Guérin),
Precis stygia Aurtvitiius, 1894, Ent. Tidskr., XV, p. 275; 1898, Rhop. Atihiop.,
p. 142; 1913, Seitz, Gross-Schmett., XIII, p. 220, Pl. ub.
This species is represented in the collection by eighty-seven speci-
mens. We have several hundreds of other specimens in the Carnegie
Museum. It evidently is common where it occurs. The female differs
from the male in being lighter in color and broader of wing. The females
vary in their markings like the males, of which there are three varieties:
_ P. stygia stygia Aurivillius, the typical form; P. stygia gregorii Butler;
and a third, which I believe has not hitherto been described or figured,
and to which I apply the name of P. stygia fuscata, new variety.
1P. simia Wallengren and P. trimeni are near allies of P. octavia (Cram.) and differ from P. antilope
pe igoel The latter species appears to me to be represented by Aurivillius reese Gross-Schmett.,
III, Pl. tic) by a female of P. trimeni, to which there is a ny ape ye resemblance upon the H spe
side, but which differs greatly from P. antilope on the lower side of the wings. I am strongly inclined
to think that an unintentional error was committed by the author of the plate to which I make
reference. P. antilope (Feisthamel) falls into the group which I designate as Kallimula.
148 Bulletin American Museum of Natural History [Vol. XLIII
(125) 3a. Precis stygia stygia Aurivillius (typical)
Plate VII, Figure 3, 2
Accepting the figure given by Aurivillius in Seitz, XIII, Pl. vib, as
adequately representing the species, and comparing it wit his diaccte
tion of the same, it is plain that this form is characterized by the pre-
sence of light areas on the under side of the wing, particularly at the
apex and outer margin of the primaries and around the outer border of
the secondaries. It is also larger in size than the other two forms, though
this difference is not so marked in some specimens of P. stygia gregorii, .
many of which approximate P. stygia stygia in size.
This is no doubt a dry-season form. Niangara, Medje, ae Munie
Katoto.
(126) 3b. Precis stygia gregorii (Butler)
Plate VII, Figure 4, 7
Precis gregorii BUTLER, 1895, Proc. Zool. Soc. London, p. 726, Pl. x11, figs. 7, 8.
This form may or may not be marked at the apex of the fore wing
and on the outer margin of both wings with the light maculation which |
characterizes typical P. stygia, but it is invariably discriminated from
the former and from the third form by the presence in the male of at
least one brilliantly white or very light yellow spot on the costa of the
secondaries on the lower side, which spot is exactly at the termination
of the dark band which in all the three forms runs from the anal angle to
about the outer fourth of the upper margin of the wing. In the females
this same spot also occurs, but it is frequently accompanied by one or
more similar spots on the interspaces lower down on the wing, which
spots usually are situated either in the middle of the dark medial band,
or slightly beyond it toward the outer border.
This is probably, like the preceding, a dry-season form. Niangara
and Medje.
(127) 3c. Precis stygia fuscata, new variety
Plate VII, Figure 5, @
This form, which is by far the commonest of the three, is, on the average, appreci-
ably smaller in expanse of wing than the two preceding, lacks almost altogether any
light cloudings or spots or marks on the under side of the wings, the only traces of such
being the almost microscopic dots which indicate the location of the vanished ocelli,
which are more or less visible on the outer third of the wings of P. stygia stygia and
P. stygia gregorti. The prevalent color of the lower side of the wings is brown marked
with darker brown or black spots, the most prominent of all these being the transverse
band on the secondaries.
This is the wet-season form and is the commonest one on the western
coast. The type locality is Medje; also at Munie Katoto.
1920] Holland, Lepidoptera of the Congo 149
That the distinction between these forms may be clear I have given
figures of them on Plate VII. The females, so far as I can ascertain,
have never before been depicted.
Of P. stygia stygia, the collection contains six males taken at
Niangara in November; two males and one female caught at Medje in
July and August; and one female captured at Bafwaboli, September 10,
1909. Of P. stygia gregorti, there are eight males caught at Medje, one
in April, the rest in July and August; and two males taken at Niangara
in November 1910. Of P. stygia fuscata, there are sixty-one males and
six females, all taken at Medje, except one male captured at Bafwaboli,
September 10, 1909; those taken at Medje were caught in July and
August, except a few taken in April.
(128) 4. Precis terea (Drury)
Papilio terea Drury, 1773, Ill. Exot. Ent., I, p. 32, Pl. xvmz, figs. 3, 4.
Precis terea AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 220, Pl. x1.
The collection contains seventy-two males and twenty females of this
species. Two of the females were taken at Leopoldville, two at Medje,
and sixteen at Niangara. One male was taken at Bafwaboli, September
1909, and eleven at Medje, June to August 1910. The remaining sixty
males were taken mostly at Niangara, in November 1910, though a few
specimens are labelled as taken at Faradje, in the same month and year,
and one or two at Medje. The females are somewhat greater in
expanse of wing and lighter in color than the males, but there is no
great variation between the sexes.
(129) 5. Precis ceryne (Boisduval)
Salamis ceryne Boispuvau, 1847, Delegorgue, Voy. Afr. Australe, II, p. 592.
Junonia ceryne TRIMEN, 1862, Rhop. Afr. Austr., I, p. 131; 1866, II, Pl. 11, fig. 4.
Precis ceryne AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 223, Pl. urd.
Four males taken at Faradje and labelled “1911-1912.” They are
all of the wet-season form of the species.
KALLIMULA, hew genus
Allied to Precis Hiibner and Junonia Hiibner, but differing in having the fore
wings strongly falcate at the apex and deeply excavated on the outer margin below
vein 6; the hind wings either strongly produced or evenly rounded opposite the end
of the cell, the margins more or less entire and strongly produced, or tailed, at the
anal angle; both wings crossed near the middle by light bands, generally some shade
of red, pink, or ochreous, though in one form (K. harpyia) these bands are shot with
blue over the red.
Type, Kallimula osborni, new species.
150 Bulletin American Museum of Natural History [Vol. XLII
_In separating from the genus Precis, as constituted by Aurivillius,
the Kallima-like forms which he has brought together in part in his
fourth group, I think I am not doing violence, in spite of the fact that
in my arrangement I am compelled to introduce as one of the members
of this group the species named pelarga, which in its dry-season form
shows a resemblance to the species which I have included under Precis.
However, K. pelarga, in its dry-season form, always has the hind wings
more or less elongated at the anal angle and thus may be discriminated
from Precis octavia and its allies, which do not have the anal angle
greatly produced. K. pelarga marks the transition between the genus
Precis and the genus Kallimula.
The species which I refer to the new genus are the following: K.
osborni, new species; eurodoce (Westwood); milonia (Felder); sinuata
(Ploetz); celestina (Dewitz); actia (Distant); antilope (Feisthamel);
antilope, var. cuama (Hewitson); pelarga (Fabricius); pelarga, var.
leodice (Cramer); pelarga, var. harpyia (Fabricius).
(130) 1. Kallimula osborni, new species
Plate VII: Figure 7, o; Figure 8, 9
@. Palpi black above, brilliantly white below. Upper side of head, thorax, and
abdomen black. Pectus white, like the lower side of the palpi. Legs pale yellowish,
dorsally shaded with fuscous. Lower side of abdomen pale fulvous. Fore wings with
costa strongly arcuate; outer margin at apex very falcate, strongly produced at the
end of vain 6; deeply excised below this and then gently rounded to the lower angle.
The hind margin of the fore wing is nearly straight. Hind wings with the costal
margin gently (flatly) curved from the base to the upper angle; the outer margin
regularly curved outwardly to a point opposite the lower angle of the cell and then
inwardly toward the anal angle, the curve running to the tip of the long tail-like pro-
jection of the anal angle. On the inner margin the hind wings are strongly curved
inwardly from the abdomen until a little after the middle, and then the curve is reversed
to the tip of the anal projection. The wings are very dark brown on the upper side
interrupted on the discal area with a bright rufous mesial band. This band on the
primaries is widest at the inner margin, and sweeps upward toward the costa termina-
ting near the origin of vein 6, being externally delimited by an almost regular curve,
running from the inner margin from a point about three millimeters behind the outer
angle to the origin of vein 6; internally the band is more irregular, being deeply
invaded by an outward projection of the dark basal tract on interspace 4. The band
has the outline of an inverted cornucopia. There isa sharply defined, small, white
spot not far from the apex between veins 5 and 6; and three black spots in the transverse
rufous band, one on each of the interspaces 2, 3, and 4, nearer the outer than the
inner margin of the band. The transverse band of the hind wing is continuous with
the band on the fore wing, narrower at the costa than on the inner margin. It is
evenly curved externally, except on interspaces 6 and 7, where the dark outer border
invades it by step-like inner projections. The outer band is wider on the secondaries
than on the primaries. The inner margin of the light band is somewhat irregular,
1920] Holland, Lepidoptera of the Congo 151
where it crosses the middle of the cell, being invaded by a sharp tooth-like projection
of the dark basal area of the wing.. There is a regular curved series of black circular
spots beginning between veins 2 and 3 and running upward to the costa, the spots
gradually increasing in size toward the costa, and being located nearer the outer
margin of the band than its inner margin. On the lower side of the wings there is a
dark narrow line of deep brown or black which runs from the tip of the tail-like pro-
jection of the anal angle of the hind wing to about the middle of the costa of the fore
wing, simulating the appearance of the mid-rib of a leaf. On either side of this
central transverse line the wings are more or less rufous, irregularly mottled with
darker brown, and being heavily clouded with dark brown at the apex of the primaries
and the anal angle of the secondaries. The fore wings near’ the apex and the hind
wings near the anal angle are marked with lighter marginal and submarginal narrow
lines and more or less frosted with light-colored scales. The series of spots which are
conspicuous on the upper side reappear on the lower side in the interspaces, but on
the lower side are pupilled with white, in most, but not in all cases.
Q. The female is not unlike the male, but the wings are broader, and at the
apex of the primaries not so strongly falcate, and the transverse band of the second-
aries is more or less creamy white, especially toward the anal angle. Expanse, <,
45-50 mm.; 9, 50-55 mm.
The male type (Medje) is in The American Museum of Natural History, the
female allotype (Cameroon) is in the Holland Collection in the Carnegie Museum.
Four males were taken by the Lang-Chapin Expedition at Medje, the dates
of capture ranging from July to September.
I have long had this species in my collection, where it is represented
by specimens collected for me many years ago in southern Cameroon
by the late Dr. A. C. Good. I take pleasure in naming it in honor of my
friend, Prof. Henry Fairfield Osborn of The American Museum of
Natural History, with whom I have spent many pleasant hours both at
home and abroad in the pursuit of pleasure and knowledge.
(131): 2. Kallimula milonia (Felder)
Precis milonia Fe.pEer, 1867, Reise Novara, Lep., III p. 403. Avurrivitiius, 1913,
Seitz, Gross-Schmett., XIII, p. 223.
Precis kowara Warp, 1871, Ent. Mo. Mag., VIII, p. 22; 1873, Afr. Lep., p. 6, Pl. v,
figs. 5, 6.
The collection upon which I am reporting contains six males and
one female. Four of the males were taken at Medje in June and July,
the other two at Niangara in November. The female was captured at
Medje in September. The transverse band in the female is much lighter
than in the males, inclining to yellowish.
(132) 3. Kallimula sinuata (Platz)
Precis sinuata Piarz, 1880, Stett. Ent. Zeit., XLI, p. 477. Aurtvitiius, 1913,
Seitz, Gross-Schmett., XIII, p. 222, Pl. tie.
Two males, one caught at Medje in August, the other at Niangara
in November. ;
152 Bulletin American Museum of Natural History [Vol. XLII
(133) 4. Kallimula pelarga (Fabricius)
Papilio pelarga Fasrictus, 1775, Syst. Ent., p. 513.
Precis pelarga AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 223.
Of this well-known species there are twenty-three specimens in the
collection. One male was caught at Gamangui in February; two males
were taken at Medje in August and a female at the same place in July;
eight males and eight females were captured at Niangara in November;
and there are two males and one female labelled as taken at Faradje
1911-1912.”
(134) 5. Kallimula leodice (Cramer)
Papilio leodice Cramer, 1777, Pap. Exot., I, p. 64, Pl. cxxxviut, figs. G, H.
Precis leodice AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 223.
There are eighty-eight specimens of this species or form, nine females
and seventy-nine males; all were caught at Niangara in November,
except one female, which was taken at Medje in August. Aurivillius
regards it as being the dry-season form of pelarga (Fabricius).
(135) 6. Kallimula harpyia (Fabricius)
Papilio harpyia Fasrictus, 1781, Spec. Ins., II, p. 104.
Precis harpyia AuRIvItuivs, 1913, Seitz, Gross-Schmett., XIII, p. 223, Pl. 1d.
The collection contains sixty specimens, forty-four males and six-
teen females, all of which were captured at Niangara in November,
except two males which were caught at Medje in August. According to
Aurivillius this is an aberrant dry-season form of pelarga.
CATACROPTERA Karsch
(136) 1. Catacroptera cloanthe (Cramer)
Papilio cloanthe Cramer, 1781, Pap. Exot., IV, p. 93, Pl. cccxxxvim, figs. A, B.
Catacroptera cloanthe AurtviLuivs, 1913, Seitz, Gross-Schmett., XIII, p. 218, Pl.uue.
This insect, formerly included in the genus Precis, is indeed very
closely related to the latter, but may easily be distinguished by its larger
size, its robust form, and somewhat hirsute appearance, not to mention
certain other minute anatomical differences which exist in both the imago
and the larva. There is but one species, C. cloanthe (Cramer), of which
two local races have been recognized by authors, one which is spread
over the eastern and southern portions of the continent, to which the
name given by Cramer strictly applies, the other C. cloanthe ligata
Rothschild and Jordan, which according to these authorities is the north-
western race and extends from Senegal to Togo. The American Mu-
seum Congo Expedition brought back a small series of specimens which
*
1920] Holland, Lepidoptera of the Congo : 153
show that the two races commingle about the headwaters of the Uelle
River and indicate a great eastward extension of the race originally de-
scribed by Rothschild and Jordan from the northwestern areas
of the Ethiopian subregion. This is, however, what might have been
expected. C.cloanthe (typical) has no dark marginal line on the primaries;
C. cloanthe ligata has such a line. Both forms have “wet-season”’ and
“dry-season”’ subvarieties. The ‘dry seasonal’’ form in each. case is
light under the wings, and the “‘ wet seasonal”’ form is dark.
The specimens, with the solitary exception of the female captured
at Medje on August 1, 1910, were taken at Faradje and Niangara in
November of that year. Three males and seven females, including the
female taken at Medje in August, belong to the form C. cloanthe ligata
Rothschild and Jordan; five males and six females belong to the “wet-
season’’ form to which Staudinger applied the varietal name C. ob-
scurior, in which the under side of both wings is quite dark brown.
SaLamis Boisduval
(137) 1. Salamis parhassus ethiops (Palisot de Beauvois)
Papilio ethiops Patisor de Brauvois, 1805, Ins. Ree. en’ Afrique et Amérique,
Lép., p. 22, Pl. m1.
Salamis parhassus AuRIvILLIvs, 1913, Seitz, Gross-Schmett., XIII, p. 217.
Sixteen specimens. One was taken at Gamangui in February and
another at the same place in June. Four were caught at Niangara in
November and all the rest were captured at Medje from June to
September.
(138) 2. Salamis cacta (Fabricius)
Papilio cacta Fasricrus, 1793, Ent. Syst., III, part 1, p. 116.
Salamis cacta AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 218, Pl. te.
On the under side of the wings there is displayed a remarkable dis-
similarity in color and maculation. No two specimens are exactly alike.
This is true also of the very large series which we have from the French
Congo and southern Cameroon. It is singular that some “species-
maker”’ has not found pleasant recreation in naming and describing the
different individuals which exist in various collections, each of which
might give opportunity for the exercise of descriptive powers, especially
if use were made of a binocular microscope, which has become such a
fashionable adjunct in entomological and ichthyological research in
recent years.
Twenty-five specimens, mostly males. One was taken at Baf-
wabaka in January, several at Niangara in November, but by far the
largest number at Medje from May to September.
154 Bulletin American Museum of Natural History [Vol. XLIIT
Katuima (Westwood)
(139) 1. Kallima rattrayi Em. Sharpe
Kallima rattrayi Em. SHarpe, 1904, Entomologist, p. 182. AurrIvILiius, 1913, Seitz,
Gross-Schmett., XIII, p. 211.
The male is very much like that sex in Kallima rumia, of which this
is probably a local race, but considerably smaller in size. The female is
different in that the hind wings are concolorous and entirely without
the broad whitish band which is always found in K. rumia; and in
addition has a conspicuous white mark near the apex of the fore wing
on the costal margin. The subapical transverse band is narrower than
in K. rumia, and is not white, as in the female of that species, but bright
straw-yellow.
Three males and a female caught at Medje (July-August), and a
male captured at Niangara in November.
(140) 2. Kallima ansorgei Rothschild
Kallima ansorgei Rotuscuitp, 1899, in Ansorge, ‘Under the African Sun,’ p. 319,
Pl., fig. 1. AvrtviLiius, 1913, Seitz, Gross-Schmett., XIII, p. 211.
This seems to be quite distinct from K. cymodoce (Cramer), al-
though it has the same form of the wings. It occurs in the eastern parts
of Cameroon also, where K. cymodoce is likewise found, and there have
been no intergrading varieties discovered thus far, to the best of my
knowledge and belief. The species is represented in the collection by
seven males taken at Medje.
Hypotimmnas Hiibner
(141) ' 1. Hypolimnas misippus (Linneus)
Papilio misippus Linnavs, 1764, Mus. Lud. Ulr., p. 264.
Hypolimnas misippus Aurivitutus, 1913, Seitz, Gross-Schmett., XIII, p. 213.
There are eleven males and seven females of this common and widely
distributed species. Three of the females belong to the varietal form
H. inaria (Cramer). The specimens were all taken at Niangara in
November 1910, except one female captured at Medje, August 11, and
one male, which was taken at Faradje, without any indication on the
label of the day and month.
(142) 2. Hypolimnas salmacis (Drury)
Papilio salmacis Drnvuny, 1773, Il, Exot, Ent., I, p. 14, Pl. vin, figs. 1-2,
Hypolimnas salmacis Avnivitii0s, 1913, Seitz, Gross-Schmett., p, 214, Pl. xiviub.
The collection contains one hundred and sixty-two males and
two females of this species, all taken at Medje. Of this number a few
=
1920] Holland, Lepidoptera of the Congo 155
are labelled as having been captured in April, the rest were taken at
dates ranging from July 8 to September 24, 1910. If collecting went
on continuously at Medje this fact would indicate that there may be
two broods annually.
There is some variation in the coloring of the upper side of the wings,
a few specimens having a purplish tint contrasting with the purer blue
of the markings of most of the specimens, and the white spots beyond
the end of the cell of the fore wing in an occasional specimen are suffused
with blue almost throughout their extent.
The two females in the collection belong to the form to which
Suffert has applied the name H. cissalma, characterized by having the
white of the paler spots replaced by yellow. The collection contains
no males corresponding to the form cissalma in that sex, which Suffert
describes as having all the lighter markings on the upper side, except
the subapical spots, blue throughout.
(143) 3. Hypolimnas monteironis (Druce)
Diadema monteironis Druce, 1874, Cist. Ent., I, p. 286.
Hypolimnas monteironis Auriviuuius, 1913, Seitz, Gross-Schmett., XIII, p. 214, Pl.
xivub.
This species, which has been by some authors regarded as a varietal
form of the preceding but which is quite distinct, is represented in the
collection by twenty-one males and one female. They all were taken at
Medyje, except one male which is ticketted ‘‘Niangara, November 14—
16, 1910.” Three of the males taken at Medje were captured in April,
all of the other specimens bear dates ranging from July 8 to September
27, 1910. Like the preceding species, this probably has at least two
annual broods.
(144) 4. Hypolimnas mechowi (Dewitz)
Diadema mechowi Dewirz, 1884, Berl. Ent. Zeit., XXVIII, p. 187, Pl. 1, fig. 2.
Hypolimnas mechowi Aurivituius, 1913, Seitz, Gross-Schmett., XIII, p. 214, Pl.
xivud.
This species is represented in the collection by a single male, which
does not differ at all from specimens in the collection of the writer from
the southern parts of Cameroon. It was taken at Bafwaboli, Nov-
ember 11, 1909.
(145) 5. Hypolimnas bartteloti Grose-Smith
Hypolimnas bartteloti GRose-Smr14, 1890, Proc. Zool. Soc. London, p. 468; 1891, Rhop.
Exot., I, Hypolimnas, Pl.1, figs. 1-2. AvuriviLuius, 1913, Seitz, Gross-Schmett.,
XIII, p. 214.
156 Bulletin American Museum of Natural History [Vol. XLIII
Of this species and its varieties there are twenty-two examples, all
of which are males. One specimen was taken at Avakubi, October 1,
1909; one at Lubila, September 20, 1909; all the rest at Medje, one on
April 6, the remainder at dates ranging from July 8 to September 27,
1910.
There is considerable variability in the specimens due to the ob-
solescence of the subapical and submarginal spots. In four of the speci-
mens belonging to this collection, as well as in numerous other speci-
mens taken in Cameroon and which are before me as I write, these spots
are wholly obliterated and I propose for this extreme fore the varietal
name obliterata.
(146) 5a. Hypolimnas bartteloti obliterata, new variety
Plate VI, Figure 5, #7
Distinguished from the typical form by the complete obliteration of the sub-
apical and submarginal spots on the upper side of both the fore and hind wings.
Traces of the uppermost subapical and the lowermost submarginal spot are to be
found on the lower side of the fore wing in some specimens, but not in all. In typical
H. bartteloti the submarginal white spots form a continuous series, one in each inter-
space, though they may vary in size, in some specimens being quite large, in others
being reduced to mere points.
The type of H. bartteloti obliterata is in The American Aronson of Natural History;
paratypes are in the same collection and in the Holland Collection in the Carnegie
Museum. Medje and Cameroon.
(147) - 6. Hypolimnas dubia (Palisot de Beauvois)
Papilio dubius Patisor pg Bravvois, 1805, Ins. Rec. en Afrique et Amérique,
p. 238, Lep., Pl. v1, figs. 2a, 2b.
Hypolimnas dubia Aurtvituivs, 1913, Seitz, Gross-Schmett., XIII, p. 215, Pl. xivirta.
There are ten males and two females of this species. One male was
taken at Bafwabaka, December 9, 1909; two males were captured at
Niangara in November 1910; the remaining nine specimens were taken
at Medje, the dates of capture ranging from June 8 to September 27,
1910. The two females were taken in the first week in August.
There is some variation in the maculation of the specimens,
particularly in the extent of the large white discal area of the hind wings,
which in one specimen is rather broadly suffused with pale ochreous.
This variability is characteristic of the species, as shown by numerous
specimens coming from various parts of the continent in the collection
of the writer.
1920] Holland, Lepidoptera of the Congo 157
(148) 7. Hypolimnas anthedon (Doubleday)
Diadema anthedon Dovsiepay, 1845, Ann. Mag. Nat. Hist., (1) XVI, p. 181; 1850,
Gen. Diurn. Lep., II, p. 281, Pl. xxxvu, fig. 2.
Hypolimnas anthedon Avurivituius, 1913, Seitz, Gross-Schmett., XIII, p. 215, Pl.
XLVIIC.
There are twenty-two males and one female in the collection.
The latter was taken at Medje in September 1910. One male is ticketted
as from Bafwaboli, September 11, 1909; another as from Bafwasende,
September 25, 1909; a third from Ngayu, December 11, 1909; a fourth
from Niangara, November 9-10, 1910. The remaining specimens were
taken at Medje, one on April 6, 1910, the rest at dates ranging from
June 6 to October 4, 1910.
There is great uniformity in the specimens, and they differ in no
respect from hundreds of others which have passed under the eye of
the writer and which have come from all parts of tropical western
Africa.
Euryteline
EuryYTELA Boisduval
(149) 1. Eurytela hiarbas (Drury)
Papilio hiarbas Drury, 1782, Ill. Exot. Ent., III, p. 17, Pl. x1v, figs. 1, 2.
Eurytela hiarbas Aurivituivs, 1913, Seitz, Gross-Schmett., XIII, p. 209, Pl. xurxe.
Four males and two females, Medje (July-August).
(150) 2. Eurytela dryope (Cramer)
Papilio dryope Cramer, 1775, Pap. Exot., I, p. 125, Pl. uxxvim, figs. Z, F.
Eurytela dryope AurtviL.Ius, 1913, Seitz, Gross-Schmett., XIII, p. 210.
One male taken at Niangara, November 14, 1910.
(151) 3. Eurytela alinda Mabille
Eurytela alinda Masi.ue, 1893, Ann. Soc. Ent. Belgique, XX XVII, p. 50. Aurt-
VILLIuS, 1913; Seitz, Gross-Schmett., XIII, p. 210, Pl. xirxe.
Six males caught at Medje (July-August).
Neptipopsis Aurivillius
(152) 1. Neptidopsis ophione (Cramer)
Papilio ophione CraMER, 1777, Pap. Exot., II, p. 27, Pl. extv, figs. Z, F.
Neptidopsis ophione AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 209, Pl. xurxd.
Eleven specimens (two not expanded) taken at Medje, the dates of
capture running from early in July to early in September.
158 Bulletin American Museum of Natural History [Vol. XLIII
Ercotis Westwood
(153) 1. Ergolis enotrea (Cramer)
Papilio enotrea CRAMER, 1779, Pap. Exot., III, p. 73, Pl. ccxxxv1, figs. A, B.
Ergolis enotrea AURIVILLIUsS, 1913, Seitz, Gross-Schmett., XIII, p. 208, Pl. xirxe.
Twenty-five specimens, nineteen males and six females, are included
in the collection. They were all taken at Medje about the middle of the
year, except two specimens which were captured at Niangara in Noy-
ember.
(154) 2. Ergolis pagenstecheri Suffert
Ergolis pagenstecheri Surrert, 1904, Iris, XVII, p. 125. AURIVILLIUS, 1913, Seitz,
Gross-Schmett., XIII, p. 208, Pl. xurxe.
Ergolis murina BARTEL, 1905, Nov. Zool., XII, p. 135.
Three male specimens taken at Medje in July.
(155) 3. Ergolis actisanes Hewitson
Ergolis actisanes Hewitson, 1875, Ent. Mo. Mag., XI, p. 183. Aurtvituivus, 1913;
Gross-Schmett., XIII, p. 208, Pl. xurxe.
This species is represented by five male specimens, four from Medje,
two taken in July, and one in each of the months, August and September,
the fifth being labelled as captured at Niangara in November.
Mesoxantua Aurivillius
(156) 1. Mesoxantha ethosea (Drury)
Papilio ethosea Drury, 1782, Ill. Exot. Ent., IIT, p. 51, Pl. xxxvn, figs. 3, 4.
Mesoxantha ethosea Aurtvitiius, 1913, Seitz, Gross-Schmett., XIII, p. 208, Pl.
XLrxb,
A single male captured at Medje, August, 1910. This is a quite
common insect on the Ogové River, and the writer has a long
series taken by the late Dr. A. C. Good at Kangvé.
Bysuia Hiibner
(157) 1. Byblia crameri Aurivillius
Byblia crameri Aunivititius, 1894, Ent. Tidskr., XV, p. 279; 1918, Seitz, Gross-
Schmett., XIII, p. 208, Pl. xirxe, figured as “ B. vulgaris U.”
This species or local race of B. ilithya is the one which prevails on
the western coast from Sierra Leone to northern Angola, penetrating
eastward as far as Uganda. It is remarkably constant throughout its
range, as extensive series of specimens from many places show. |
Eleven males and three females. One female was caught at Kwa-
mouth, July 14, 1909, the other two at Niangara in November, 1910,
Three males were taken at Medje in August, all the rest at Faradje and
Niangara in November.
1920] Holland, Lepidoptera of the Congo 159
Eunicine
AsTEeropeE Hiibner
Asterope Hispner 1816, type amulia Cramer, 1779, Pap. Exot., II, Pl. cuxxx, figs.
C7 .D. ‘
_Crenis BotspuvAL 1833, type madagascariensis. Cf. Rothschild and Jordan, 1903,
Nov. Zool., X, p. 528.
(158) 1. Asterope occidentalium (Mabille)
Crenis occidentalium MaBI.uE, 1876, Bull. Soc. Zool. France, I, p. 275. AURIVILLIUS,
1913, Gross-Schmett., XIII, p. 205, Pl. xuviuig.
Asterope occidentalium ROTHSCHILD AND JORDAN, 1903, Nov. Zool., X, p. 529.
Fourteen specimens: one male taken at Gamangui in February;
the rest captured at Medje (July-September).
(159) 2. Asterope boisduvali (Wallengren)
Crenis boisduvali WALLENGREN, 1857, Rhop. Caffr., p. 30. AuRtviLuius, 1913, Seitz,
Gross-Schmett., XIII, p. 205, Pl. xivinig.
Asterope boisduval: ROTHSCHILD AND JORDAN, 1903, Nov. Zool., X, p. 530.
Nineteen specimens, of which one was taken at Gamangui in June,
two at Niangara in November, and the rest at Medje (May—September).
(160) 3. Asterope natalensis (Boisduval)
Crenis natalensis BoispuvaL, 1847, Delegorgue, Voy. Afr. Australe, IT, p. 592. Aurt-
vituius, 1913, Seitz, Gross-Schmett., XIII, p. 206.
Asterope natalensis ROTHSCHILD AND JORDAN, 1903, Nov. Zooi., X, p. 531.
One male captured at Niangara in November.
Marpesiinze
Cyrrestis Westwood
(161) 1. Cyrestis camillus (Fabricius)
Papilio camillus Fasricrus, 1781, Spec. Ins., IT, p. 11.
Cyrestis camillus AURIVILLIUS, 1913, Seitz, Gross-Schmett., XIII, p. 203, Pl. xurxb.
Twenty-two specimens, mostly taken at Medje from May to
September, though there are a couple captured at Niangara in November
and one from Bafwabaka caught in January.
Neptidinz
Neptis Fabricius
The genus Neptis is well represented in the woodlands of Africa.
The collection upon which I am reporting contains a number of
species which are already more or.less well known and some rarities.
160 Bulletin American Museum of Natural History [Vol. XLIII
(162) 1. Neptis metella Doubleday and Hewitson
Neptis metella DoUBLEDAY AND HEewitTson, 1850, Gen. Diurn. Lep., I, p. 272, Pl. xxxv,
fig. 2. HoLuanp, 1892, Ent. News, ITI, PI. rx, fig. 3.
Nine specimens, all taken at Medje in July and August, ‘except one
caught at Gamangui in June.
(163) 2. Neptis marpessa Hopffer
Neptis marpessa Hoprrer, 1855, Monatsber. Akad. Wiss. Berlin, p. 640; 1862, Peters,
Reise n. Mossambique, Ins., p. 383, Pl. xxrv, figs. 9, 10.
Neptis pasteuri SNELLEN, 1882, Tijd. v. Ent., XXV, p. 221.
Neptis marpessa AuURIVILLIUvs, 1912, Seitz, Grow-eluslits XII, p. 199.
One specimen was captured at Niangara, November 20, 1910. This
is the continental form of the insect which isfoundin Madagascar and
was named JN. saclava by Boisduval. It is very abundant on the east-
ern coast and I have long series from Natal and Mombasa.
(164) 3. Neptis nemetes Hewitson
Neptis nemetes Hewirson, 1868, Exot. Butt., IV, Neptis, Pl. 1, figs. 1,2. Ho.uanp, |
1892, Ent. News, III, Pl. rx, fig. 4.
Seven specimens, one taken at Kwamouth, July 15, one from
Risimu, taken September 8, 1909, the rest from Medje, June to August.
(165) 4. Neptis agatha (Stoll)
Plate VI: Figure 8, Forest Form, o; Figure 7, Open Country Form,
Papilio agatha Srou1, 1780, in Cramer’s Pap. Exot., IV, p. 76, Pl. cccxxvn, figs. A, B.
Neptis agatha AuriviLx10s, 1912, Seitz, Gross-Schmett., XIII, p. 200, Pl. xuviiid.
Of this species the collection contains eighty-seven specimens, in-
cluding some of the varieties to which discriminating students have
given names founded in some cases upon the measurement of the size
of the spots and bands. The form lativittata Strand, the author of which
informs us that the mesial band of the secondaries is eight millimeters
wide, is represented. I discover that all of the specimens falling into
this category are large and well-developed females, as is also the case
with a lot of specimens I have from other parts of Africa. I call into
question the specific validity of this so-called “species,’’ which is not
even a variety.
I note, however, that there is a distinction, which I do not believe
has been previously pointed out, between the specimens which come
from the hot woodlands and those from the more open country of the
eastern and southern grass-lands. As it happens, most of the speci-
mens representing this species were taken at Niangara and Faradje.
s
=
&-
=
j
‘
1920] Holland, Lepidoptera of the Congo 161
They agree perfectly with spoeliaionias taken on the hills back of Mombasa,
of which I have a long suite, and with series of specimens coming from
Natal and other more southern localities. By selecting from the mass
all those which were caught at Medje about the middle of the year, I
discover that the latter as a group are not only generally larger but have
the transverse mesial band of the primaries a little straighter and
narrower than the other lot of specimens. Comparing them with long
suites of specimens coming from Cameroon, Sierra Leone, and the
valley of the Ogové, I discover that the specimens from Medje are iden-
tical in appearance with these, and that there is a slight but constant
difference between the whole assemblage of specimens from the wooded .
tropics and those which come from the steppes of the east and the
south. By pinning out fifty of each of the forms selected according
to locality, as I have done, the different facies of the two lots
becomes very apparent and striking to the eye, even more so than
when only two individuals are compared with each other.
(166). 5. Neptis seeldrayersi Aurivillius
Neptis seeldrayersi AuRivILLIus, 1895, Ent. Nachr., p. 379; 1898, Rhop. Aithiop.,
p. 167, Pl. 1, fig. 7; 1912, Seitz, Gross-Schmett., XIII, p. 200, Pl. xnvimid.
Eight specimens, all taken at Medje in July and August. There is
some individual variability. In one specimen the fourth white spot
reckoning from the costa is obsolete in the mesial band and gives the
wing a different facies. The species is not uncommon farther west and
we have it in some numbers from the region of the Ogové River.
(167) 6. Neptis nysiades Hewitson
Neptis nysiades Hew1tson, 1868, Exot. Butt., IV, Neptis, Pl. 1, figs.’3, 4.
Six males and four females, all caught at Medje, the dates of cap-
ture including April, July, August, and September.
(168) 7. Neptis metanira Holland
Neptis metanira HOLLAND, 1892, Ent. News, III, Pl. rx, fig. 6.
Aurivillius, in the ‘Rhopalocera Acthiopica,’ expresses the opinion
that this form is simply an aberration of the preceding species. He may
be right but, as the insect has a very different facies from N. nysiades
on the upper side, though I agree that it closely resembles nysiades
on the under side, and, as it is constantly turning up and we have quite a
good series from Cameroon and other parts, I am inclined to let the
matter stand as I originally put it until the test of breeding shall show
that Dr. Aurivillius is correct in his surmise.
One male, taken at Medje, August 13, 1910.
162 Bulletin American Museum of Natural History [Vol. XLIII
(169) 8. Neptis continuata Holland
Neptis biafra var. continuata HoLuanp, 1892, Ent. News, III, Pl. rx, fig. 9.
Aurivillius in the ‘Rhopalocera Aithiopica’ says that this is, like
the preceding, an aberration of N. nysiades Hewitson. I admit that I
was in error in making it a varietal form of N. biafra Ward, which is a
different insect, though there are resemblances on the under side of the
wings which led me to take the step. I have a good series of this thing
from various localities, many more than I had when I first named the
insect. As it is every now and then being received from different parts
of the hot lands of Africa and is quite constant and recognizable, I am
inclined to apply the remarks which I made under the preceding
species and let the matter stand until we shall receive more light. It is
not conspecific with NV. bafra Ward, as I have already said. The figure
of the latter insect, which I gave on the same plate in the Entomological
News, is that of a typical N. biafra, o, and it does not take much effort
to see the great difference between the two insects. Twenty-six years
of study since I wrote that article on the genus Neptis has broadened my
knowledge of the subject.
There are four examples of this form, three of which were taken at
Medje in July and August, the fourth at Munie Katoto, September
1909.
(170) 9. Neptis nicomedes Hewitson
Neptis nicomedes Hewrrson, 1874, Ent. Mo. Mag., X, p. 205. Kirsy, 1894, Handbook
Lep., I, p. 147, Pl. xx, fig. 3.
I refer to this species four specimens which agree both with the
description and the figure cited above, except that all of them have the
mesial band of the primaries slightly inangulated on its inner margin
at vein 4, thus resembling the variety quintilla Mabille!| The only
difference which I am able to affirm definitely between typical nicomedes
and quintilla is the fact that in the latter the longitudinal streak in the
cell of the fore wing is reduced and obsolete basad and persists merely
as a short bar or curved line near the end of the cell. I have good speci-
mens of N. quintilla which were collected in Cameroon, before me as I
write, and quite agree with Dr. Aurivillius in regarding the two forms as
merely varietal.
The specimens upon which I am reporting were taken at Medje,
one in June, one in July, and two in August. There are two males and
two females.
a ~~ -~—-
‘Kirby’ A Seiten’ in mr ‘ieiubal and may not be true to type.
See aS i ee es
ee ee
1920] Holland, Lepidoptera of the Congo 163
(171) 10. Neptis strigata Aurivillius
Neptis strigata AURIVILLIUS, 1894, Ent. Tidskr., XV, p. 284, fig. 10; 1913, Seitz,
Gross-Schmett., XIII, p. 201, Pl. xivute.
There are five specimens of this species, three males and two females,
all taken at Medje, except one of the females which was caught at Pawa
in October. The dates of capture at Medje cover the months of April,
July, August, and September. One of the specimens agrees so closely
with the figure given by Aurivillius in the ‘Gross-Schmetterlinge’
(loc. cit.) that it might have served the artist as his model. The rest do
not have the longitudinal streak in the cell of the fore wings divided
into two parts, but at the point where this division is shown in the figure
given by’ Aurivillius there is a small black spot which, by becoming
only a little larger, would cause the band to be interrupted as shown by
the author of the species. We have this insect also from Cameroon and
the Ogové Valley.
(172) 11. Neptis biafra Ward
Plate VIII, Figure 3, 2
Neptis biafra Warp, 1871, Ent. Mo. Mag., VIII, p. 121; 1874, Afr. Lep., p. 12, Pl
Ix, figs. 1,2. Ho Luann, 1892, Ent. News, III, Pl. rx, fig. 10, #.
The collection contains one female caught at Medje, June 30,
1910. In this specimen the two small spots at the basal end of the cell
of the fore wing coalesce, while the larger spot at the end of the cell re-
mains free. We have some fine specimens of the male in our collections
gathered for us in Cameroon and on the Ogové River. It is one of the
most beautiful of the species in the genus and is related to N. strigata
but is larger and more strikingly marked. I strongly suspect that my
valued friend, Dr. Aurivillius, was in some doubt as to the identification
of N. biafra, both when he queried the identity of my figure of the male
with his N. strigata, as he does in the ‘Rhopalocera A®ithiopica,’ and
when in the ‘Gross-Schmetterlinge der Erde’ he places N. biafra in
another group, separate from strigata, to which it is allied, and gives a
description, which is not at all conformed to the description given by
the author of the species. I wish to say that the figure of N. biafra
given by me in the Entomological News is that of a specimen which con-
forms absolutely to the description of Ward, and that the insect, while
not far removed from N. strigata, is totally distinct. With a good series
of both species before me for comparison, I am able to affirm this positively.
(173) 12. Neptis nicoteles Hewitson
Neptis nicoteles Hewitson, 1874, Ent. Mo. Mag., X, p. 206. Hotianp, 1892, Ent.
News, ITI, Pl. rx, fig. 8.
One male, lacking head and antenne, taken at Medje, July 13, 1910.
164 Bulletin American Museum of Natural History [Vol. XLIII
(174) 13. Neptis nicobule Holland
Neptis nicobule HoLuanD, 1892, Ent. News, III, p. 249, Pl. rx, fig. 7. AurtviLirus,
1898, Rhop. thiop., p. 168; 1913, Seitz, Gross-Schmett., XIII, p. 202.
One male, taken at Gamangui, June 17,1910. It agrees perfectly
with the type with which it has been compared.
(175) 14. Neptis lermanni Aurivillius
Nept,s lermanni AuRIVILLIUS, 1896, Ofvers. Sv. Vet. Akad. Férh., LIII, p. 431; 1913,
Seitz, Gross-Schmett., XIII, p. 202, Pl. xivui/.
Seven males and three females: two males caught at Gamangui in
June; all the rest at Medje, from June to August.
(176) 15. Neptis melicerta (Drury)
Papilio melicerta Drury, 1773, Ill. Exot. Ent., II, p. 34, Pl. xxx, figs. 3, 4.
Neptis melicerta Hotuanp, 1892, Ent. News, III, Pl. rx, fig. 5. Aurtviiiius, 1913,
Seitz, Gross-Schmett., XIII, p. 202, Pl. xtvute.
Of this common insect there are thirty-five examples, all taken at
Medje from July to September, except two which were captured at
Niangara in November.
NEPTIDOMIMA, new genus
(177) 1. Neptidomima exaleuca (Karsch)
Neptis exaleuca Karscu, 1894, Berl. Ent. Zeit., XX XTX, p. 9, fig. 5. AuRrvrirus,
1913, Seitz, Gross-Schmett., XIII, p. 202.
There is a single specimen of this rare and curious insect, taken at
Medje in July. Although it agrees in the neuration of the wings with the
genus Neptis, I nevertheless think that it should be separated generically
from that genus because of the structure of the palpi, which are more
robust, porrect, and hirsute thanin any species of Neptis known to
me. Furthermore, the insect is throughout more robust in its structure
than is the case in the genus Neptis. I propose the generic name Nep-
tidomima for the creature, of which genus it is the type.
Nymphalinez
Pseupacraa Westwood
(178) 1. Pseudacrwa hostilia warburgi Aurivillius
Pseudacraa hostilia var. warburgi Aurivitii0s, 1892, Ent. Tidskr., XII, p. 200;
1912, Seitz, Gross-Schmett., XIII, p. 194, Pl. xivie.
One male, captured at Medje, August 1910.
aa a Pe oe
1920] Holland, Lepidoptera of the Congo 165
(179) 2. Pseudacrea semire (Cramer)
Papilio semire Cramer, 1779, Pap. Exot., III, p. 3, Pl. oxcry, figs. B, C.
Pseudacrea semire AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 194, Pl. xurva.
Four males: one taken at Gamangui in June, and re caught at
Medje in July and August.
(180) 3. Pseudacrea lucretia (Cramer)
Papilio lucretia CraMER, 1775, Pap. Exot., I, p. 71, Pl. xtv, figs. C, D.
Pseudacrea lucretia AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 197, Pl. xivia.
Represented by fifty-four specimens, nineteen of which are not
expanded. The specimens were mainly taken at Medje, though about
half a dozen bear other scattering locality labels. The dates of capture
range from May into September.
The typical form is well represented, but the majority of the speci-
mens belong to the slight variety named Pseudacrea protracta by Butler
in which the light discal area of the hind wing is yellowish instead of
pure white as in the typical form. But there are intergrades and it is
hard to say in the series where one form begins and the other ends.
(181) 4. Pseudacrma clarki Butler
Pseudacrea clarki BuTLER, 1892, Trans. Ent. Soc. London, p. 201, Pl. x, figs. 1, la.
AURIVILLIvS, 1912, Seitz, Gross-Schmett., XIII, p. 197, Pl. xiv1b.
This lovely butterfly is represented by six males taken at Medje
(July to September).
(182) 4a. Pseudacrea clarki egina Aurivillius
Pseudacrea clarki var. egina AuRIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 197.”
Four males taken at Medje (May to June) and one at Niangara in
November.
(183) 5. Pseudacrea gottbergi Dewitz
Pseudacrea gottbergi Dewit1z, 1884, Berl. Ent. Zeit., XXVIII, p. 187, Pl. 1, fig. 1.
AvRIVILLIvs, 1912, Seitz, Gross-Schmett., XIII, p. 197, Pl. xiv1b.
Five males: four taken at Medje (June to September) and one at
Munie Katoto, September 10, 1909.
(184) 6. Pseudacrea hobleyi Neave
Pseudacrea hobleyi Neave, 1904, Nov. Zool., XI, p. 331. Aurtvitii1us, 1912, Seitz,
Gross-Schmett., XIII, p. 196.
One female taken at Medje, June 28, 1910. The subapical band of
the fore wing is a trifle wider than usual.
166 Bulletin American Museum of Natural History [Vol. XLIII
PsEUDONEPTIS Snellen
(185) 1. Pseudoneptis conobita (Fabricius)
Papilio cenobita Fasrictus, 1793, Ent. Syst., III, part 1, p. 247.
Pseudoneptis cenobita AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 193, Pl.
xivib.
Thirty specimens, mostly males, a few of which were taken at
Gamangui in June, the rest at Medje in the middle months of the year.
Catuna Kirby
(186) 1. Catuna crithea (Drury)
Papilio crithea Drury, 1773, Ill. Exot. Ent., II, p. 29, Pl. xvt, figs. 5, 6.
Catuna crithea AuRIviLuivs, 1912, Seitz, Gross-Schmett., XIII, p. 192.
Thirty-one specimens, mostly males, taken at Medje from June to
September.
(187) 2. Catuna angustata (Felder)
Euomma angustatum Fever, 1867, Reise Novara, Lep., III, p. 425.
Catuna angustata AuRIvILLIvs, 1912, Seitz, Gross-Schmett., XIII, p. 192.
Fifty-seven specimens taken at the same place and times as the
foregoing species.
(188) 3. Catuna oberthiiri Karsch
Catuna oberthiiri Karscu, 1894, Berl. Ent. Zeit., XX XIX, p.4. Aurrvitiius, 1912,
Seitz, Gross-Schmett., XIII, p. 192, Pl. xvid, e.
Twenty-two examples to which the remarks made under the two
preceding species apply.
In addition to the specimens enumerated above there are in the
collection thirty-nine unexpanded specimens representing all of the three
species, but which cannot be separated easily without being spread
so that the upper side of the wings can be examined.
CyNANDRA Schatz
(189) 1. Cynandra opis (Drury)
Papilio opis Drury, 1773, Ill. Exot. Ent., I, p. 33, Pl. xvii, figs. 5, 6.
Cynandra opis Auntvitiius, 1912, Seitz, Gross-Schmett., XIII, p. 191, Pl. xuvie,
9.
Of this insect there are five males and eight females, all of which
were taken at Medje, except one badly damaged male and one female
which were taken at Niangara, in November 1910. The dates of
capture at Medje include the months of April, June, July, August, and
September.
1920] Holland, Lepidoptera of the Congo 167
Aterica Boisduval
(190) 1. Aterica galene (Brown)
Papilio galene Brown, 1776, New Ills. of Zool., p. 94, Pl. xxxvut.
Aterica theophane Horrrer, 1855, Monatsber. Akad. Wiss. Berlin, p. 641.
Aterica galene AuRIvILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 191, Pl. xivia, b.
There are thirty males and twenty-three females of this species
and its varieties. The form theophane is an extreme in which the light
area of the hind wing is broadly suffused with red, but there are inter-
grading specimens connecting up these with such specimens as have the
light area quite white. One male is labelled as having been captured
at Noki in Angola, a few came from Gamangui and were taken in June,
but the great majority were captured at Medje in the middle months of
the year.
Hamanvumipa Hiibner
(191) 1. Hamanumida dedalus (Fabricius)
Papilio dedalus Fasricivs, 1775, Syst. Ent., p. 482.
Hamanumida dedalus Aurtvituius, 1912, Seitz, Gross-Schmett,. XIII, p. 191, Pl.
xivif.
Of this form, which differs from the following by the duller coloring
of the lower side of the wings and the partial suppression of the white
spots on that side, there are sixteen males and five females taken at
Niangara in November. -
(192) la. Hamanumida dedalus meleagris (Cramer)
Papilio meleagris CraMER, 1775, Pap. Exot., I, p. 102, Pl. xuv1, figs. A, B.
Hamanumida meleagris Aurtviturus, 1912, Seitz, Gross-Schmett., XIII, p. 191, Pl.
xuivif.
This variety is represented by a long series of males and females,
all of which were taken at Niangara and Faradje in November, except
one taken at Noki in Angola, June 24, 1909, and another caught at
Avakubi in November 1909.
The two forms, H. dedalus and H. meleagris, intergrade and, as both
were taken at Niangara in large numbers at the same time of the year,
the opinion of Dr. Aurivillius that they represent seasonal forms is
open to question. They are mere color varieties, H. dedalus having the
under surface dull and the white spots for the most part suppressed,
and H. meleagris having the under surface of the wings brighter and the
white spots present. C
The writer has a large series of both forms collected for him on the
Ogové River and at Gaboon by the late Dr. A. C. Good and these, too,
168 Bulletin American Museum of Natural History [Vol. XLIII
by the dates of capture seem to confirm the opinion that the two forms
are probably the issue of one and the same brood and that they are
not “‘seasonal varieties’? in the sense in which that term must be ac-
cepted.
EvupHzpra Hiibner
The genus Eupheedra is one of the most puzzling genera among the
diurnal Lepidoptera of Africa. A vast number of so-called species, or
forms, have been distinguished by writers, in many instances based
apparently upon solitary individuals coming into their’ possession
without sufficient material at hand to institute broad comparisons.
Until in coming years some careful student on the ground shall succeed
in breeding these insects from ova laid by individual females there
necessarily will remain more or less confusion in certain groups. This is
especially true of the forms allied to EH. preussi Staudinger, EH. ceres
(Fabricius), and E. xypete (Hewitson).
Of these I have before me as I write a vast assemblage of specimens,
many of which were brought home by the American Museum Congo
Expedition. For example, there were collected by this expedition one
hundred and eight specimens representing EH. preusst and varieties,
males and females. We have in the collections contained in the Carnegie
Museum as many more from various localities. In all this assemblage
of specimens it is almost impossible to find two individuals in either
sex which are absolutely alike. There are slight differences in the shade
of color on both the upper and under sides, in the number of spots in
the cells on both sides of the wings, and in the discal and submarginal
markings, which may be strongly, feebly, or not at all indicated. In
‘addition to this, a close and accurate study of the literature shows that
the descriptions and the figures which have been given by authors
represent individuals rather than species; so that it is exceedingly
difficult to state in exact terms where one so-called species begins and
another ends in this group. The same remarks apply to other so-called
“snecies,”’ in other groups in which almost infinite variety is plainly
visible, but appearing to me to have as little real significance as the
shape and location of the spots on common cattle. On the other hand,
there are certain forms recognized by authors which appear to be quite
constant over the entire range in which they occur and which are not
difficult to discriminate. In some cases, however, it may well be, as has
been suggested, that some of these which are not distantly related to
each other may, when the test of breeding is applied, turn out to be
1920] Holland, Lepidoptera of the Congo 169
mere color varieties or seasonal forms which have not as yet been recog-
nized as such. The genus as a whole presents as much difficulty to the
systematist as the genus Argynnis and, in fact, I am disposed to regard
it as more difficult than the latter genus. The arrangement and classi-
fication of the specimens contained in the collection upon which I am
now reporting must, therefore, in certain of the groups be regarded as
. in a measure provisional.
I. Prrseis Group
(193) l. Euphedra imitans Holland
Plate VII, Figure 2, type, 2
Euphedra imitans Houianp, 1893, Canad. Ent., XXV, p. 3. AurrviLirus, 1898,
Rhop. Athiop., p. 185; 1912, Seitz, Gross-Schmett., XIII, p. 190.
Of this well-marked species, which has never before been figured,
there are in the collection one male and three females, all taken at
Medje, the male on September 27, one of the females in July, and two
of the females in August 1910.
II. Rusprna Group
I may say in passing that I assign EF. crowleyi Kirby to this
group. It is a small species, which Aurivillius in error classifies under
Euptera, but I have both males and females which show that the insect
is a true Euphedra.
(194) 2. Euphedra ruspina (Hewitson)
Romaleosoma ruspina Hewitson, 1865, Exot. Butt., III, Romaleosoma, Pl. u,
figs. 6, 7.
Buphedra Piette AURIVILLIUS, 1898, Rhop. Aithiop., p. 185; 1912, Seitz, Gross-
Schmett., XIII, p. 190, Pl. xumb. (In error designated on the plate as E. eleus.)
Of this well-known species there are eleven males and three females.
All of the males and two of the females are labelled as captured at Medje;
one female is marked as taken at Gamangui in July.
III. Exuevs Group
(195) 3. Euphedra eleus (Drury)
Papilio eleus Drury, 1782, Ill. Exot. Ent., III, p. 14, Pl. x, figs. 1, 2.
Euphedra eleus AURIVILLIUS, 1898, Rhop. Aithiop., p. 185; 1912, Seitz, Gross-Schmett.,
XIII, p. 190, Pl. xumb. (Middle figure, 2, erroneously labelled ruspina on
plate.)
The figure of E. eleus given by Drury does not appear to be for-
tunate. The insect on his plate (loc. cit.) is represented as having the
hind wing strongly produced at the anal angle. Such a specimen, so
170 Bulletin American Museum of Natural History [Vol. XLIII
far as my knowledge goes, does not exist, and I think the outline of the
wing in his drawing is due to the fancy of the draftsman, who has ex-
aggerated in this respect. The color and markings of the upper side of
the wings in all other respects agree with the multitude of specimens
before me, which may be accepted as typical E. eleus. The color and
markings. of the lower side of the wings in Drury’s figure represent an
extreme variety.
Of what I long have determined to be H. eleus (Drury) there are in
the collection nine males and four females, all taken at Medje at dates
ranging from June to September, except one female which was taken
at Niangara in November 1910. These specimens all have the trans-
verse subapical band white, relatively narrow, consisting of four spots of
which the third, reckoning from the costa, is the largest, and the ground-
color of both wings is dark brownish red as shown in Drury’s figure.
(196) 3a. Eupheedra eleus hybrida Aurivillius
Euphedra eleus ab. hybridus (Staudinger, in litt.), AURtVILLIUS, 1898, Rhop. Aithiop.,
p. 186; ab. hybrida Aurtvitiivs, 1912, Seitz, Gross-Schmett., XIII, p. 190.
This varietal form, or subspecies, is represented in the collection by
twenty-five males and thirteen females. The dark apical area and, to
some extent, the dark marginal band of the secondaries in these speci-
mens, shows in certain lights a greenish luster. In the males the sub-
apical transverse band is, in almost all cases, composed of only three
spots, yellowish white in color, the lowermost of. which is the largest.
In ten of the specimens in the collection there is, however, a fourth spot
indicated as a mere pointlet in the interspace between the second and
third submedian nervules. The red ground-color of the wings is much
paler than in typical eleus, being orange-red and not dark brown.
The specimens were almost all taken at Medje, at dates ranging
from June to September, though there are several specimens, both male
and female, recorded as captured at Gamangui in June, a male taken
at Niangara in November, and a female at Avakubi in October.
This varietal form with the paler ground-color of the wings, the
greenish luster of the dark margins, and the yellow subapical bands,
when pinned out in a series, contrasts rather strikingly with the darker
clored specimens which I refer to EZ. eleus Drury, but when it comes to a
comparison of the under side of the wings the difference is so trifling as
to make it appear certain that the two forms are merely varietal.
1920] Holland, Lepidoptera of the Congo 171
(197) 3b. Euphedra eleus coprates (Druce)
Romaleosoma coprates Druce, 1875, Proc. Zool. Soc. London, p. 411.
Euphedra eleus var. coprates AuRIvILLIUS, 1898, Rhop. Athiop., p. 186; 1912, Seitz,
Gross-Schmett., XIII, p. 190, Pl. xia, o (in error hybridus), Pl. xumb, 2.
Of this varietal form, distinguished by the presence of spots in the
cell of the fore wings, both on the upper and under side, there are three
males taken at Medje, one in June and the other two in August.
(198) 4. Euphedra edwardsi (Hoeven)
Aterica edwardsi Horven, 1845, Tijd. Nat. Gesch., XII, p. 251, Pl. rv, figs. la, 1b.
Romaleosoma pratinas DouBLEDAY AND Hewitson, 1850, Gen. Diurn. Lep., II, p.
284, Pl. xxxvin, fig. 3, o.
Euphedra edwardsi Aurivituius, 1898, Rhop. Athiop., p. 186; 1912, Seitz, Gross-
Schmett., XIII, p. 189, Pl. xu11c ( 9, non o, as figured on plate).
The collection contains one female taken at Medje on July 29, 1910.
IV. Crres Group
As already indicated, the collection contains a great many speci-
mens referable to this group showing innumerable slight variations in
color and markings, so that it becomes very difficult to determine by
fixed lines specific and varietal distinctions.
(199) 5. Euphedra preussi Staudinger (?)
Euphedra preussi StauvinGER, 1891, Iris, IV, p. 119, Pl. 1, fig. 1.
To this species I refer, with some doubt, eighteen males and four
females, all taken at Medje, at dates ranging from July to September,
except one male captured at Niangara in November and one female
taken at Bafwasende on January 7, 1910.
Not one of these specimens agrees absolutely on the lower side with
the figure given by Staudinger, though the males agree almost per-
fectly with the figure which he gives of the upper side of the wings.
In this connection it is to be observed that the insect figured as H. preussi
by Aurivillius in Seitz, ‘Gross-Schmetterlinge,’ XIII, Plate xurid, is not
the insect figured by Staudinger in Iris (loc. cit.) but represents the
variety named njamnjami by Staudinger. Staudinger’s figure represents
an insect which is bright ochraceous on the under side. The insects
upon which I am reporting are prevalently greenish on the under side,
though some of them are brownish, but almost all are inclined to be
more or less ochraceous on the inner margin of the lower side. The
number of the spots in the cells of both the fore and hind wings on the
lower side is somewhat variable. All-have a pale discal spot beyond the
172 Bulletin American Museum of Natural History [Vol. XLIIT
cell, while in some specimens this discal spot forms the lower end of a
pale band extending costad to the pale costal marginal border. No two
specimens are absolutely alike. The submarginal spots on the upper
and under side are also variable, though generally indicated on the
upper side, and sometimes quite strongly; in other specimens, particu-
larly on the lower side, these submarginal spots are more or less obsolete.
The transverse apical band in the males is, as in Staudinger’s figure,
narrow and greenish, and the apical region on the upper side in the males
is more or less suffused with green. In the females the transverse sub-
apical band is white, showing strongly against the black apical area of
the fore wings on the upper side, and is composed of four spots.
None of the specimens are absolutely typical #. preussi Staudinger, but
knowing, as I do, how greatly these insects tend to vary, I am not
inclined to set up a new species merely basing it upon the slightly
different shade of the lower side of the wings.
(200) 5a. Euphedra preussi njami Staudinger
Euphedra preussi var. njami Staudinger, 1891, Iris, IV, p. 125.
I refer to this varietal form erected by Staudinger twenty-one males
and six females, all taken at Medje at dates ranging from April to
September, except two males taken at Niangara in November and one
female captured at Gamangui in June.
Staudinger differentiates this form from his #. preussi by stating
in his description that the fore wings of the male on the upper side are
darker, not laved with green, and that on the under side they are ferru-
ginous and, further, that the submarginal spots on the lower side are
distinctly visible. The long series of specimens before me which I refer
to this form correspond closely with Staudinger’s description, but
there is considerable variability in the shading of the upper and under
sides of the wings. On the upper side, some are distinctly greenish; in
others the ground-color passes into bluish green. On the. under side,
some are indeed ferruginous, as he states; others, however, are somewhat
greenish, and no two are absolutely alike in every minute particular.
Almost all have a small light spot on the under side of the secondaries
beyond the end of the cell, and in some specimens this light spot is
extended costad, forming a light bar which loses itself in the pale costal
border. There is extreme variability in this respect. The females are
like the females of typical preussi, but almost all of them have the pale
diseal transverse band on the secondaries running from the end of the
cell upward toward the costa more distinctly marked than in the males.
1920] Holland, Lepidoptera of the Congo 173
(201) 5b. Euphedra preussi njamnjami Staudinger
Euphedra preussi var. njamnjami StaupINGER, 1891, Iris, IV, p. 125.
Euphedra preussi AuriviLii0s, 1912, Seitz, Gross-Schmett., XIII, p. 189, Pl. xumd.
To this varietal form, characterized by Staudinger as having the
upper side of the wings black and not laved with green at the apex as in
typical preussz, the under side ferruginous, without submarginal mark-
ings, I refer twelve males and twenty-one females from Medje and
Gamangui. The males and females are all distinctly somewhat larger in
size than either of the two foregoing forms. The outer two-thirds of the
wings of the males are black, the subapical transverse band narrow,
composed of four spots separated by the nervules, and the lower side
without submarginal markings save as these may be indicated by small
white dots in a few of the specimens. The prevalent color of the under
side of the wings is brownish ochraceous, or ferruginous more or less
laved with green in the case of the males, but not in the case of the
females. As in the two preceding forms, the hind wings beyond the end
of the cell may have a discal light spot, or this light spot may be ex-
tended eostad in the form of a pale transverse discal band, losing itself
in the light costal margin.
While I am averse to adding to the apparent interminable confu-
sion which already exists in this species, I am, nevertheless, constrained
by facts which are too apparent to be overlooked to set off from among
the specimens before me several varietal forms which are quite as
worthy of being discriminated as those which have already been men-
tioned.
(202) 5c. Euphedra preussi notata, new variety
Plate XI, Figure 1, #7
I apply this name to two males and three females before me. The males and
females are relatively smaller than any specimens of the varieties njami Staudinger
and njamnjami Staudinger, which they resemble on the upper side of the wings,
save that at the end of the cell on the upper side of the secondaries they all have a
distinct black spot. On the under side the ground-color of the wings is chocolate-
brown, with a more or less bluish suffusion, and the spots in the cell of both the fore
and hind wings are large and distinct, and the submarginal spots are also well indi-
cated, having an inner dark nucleus accentuated externally by light spots. The
transverse subapical band in the males is pale whitish green; in the females pure white.
Expanse, o’, 65-67 mm.; 9, 70-80 mm.
The type @ and allotype 2 are in the The American Museum
of Natural History; two paratypes in the Holland Collection in the
Carnegie Museum. All are from Medje.
174 Bulletin American Museum of Natural History (Vol. XLII
(203) 5d. Euphedra preussi subviridis, new variety
Plate XI, Figure 4, 7
Closely resembling E. preussi, but with the upper outer two-thirds of the fore
wings in the case of the male velvety black, the postapical transverse band diffuse,
greenish, and toward the apex fading into a lustrous green subapical area which, in
certain lights, shows as a brilliant green tract covering the apical area from near the
apex to the outer end of the cell; the hind wings and the posterior margin of the fore
wing for some distance iridescent greenish blue, this area on the fore wings reaching
the lower margin of the cell near the base, but not reaching the outer angle. The
posterior wings broadly bordered with dark green, accentuated with a submarginal
series of black velvety spots. On the under side in the male the wings are more or
less grass-green, tinged with chocolate-brown, the transverse subapical band of the
primaries being whitish. The spots in the cells of the primaries.and the secondaries
are variable in number and size, as is true of all the varieties, and this is also true of the
submarginal series of dark spots. The female is marked on the upper side like the
male except that the transverse subapical band is pure snow-white. The outer
margins, as in all the forms of E. preussi, have the fringes dark, interrupted on the
interspaces with white, and the fore wings are at the apex tipped with white.
There are two males and one female from Medje which I refer to
this form. The male and female types are in The American Museum of.
Natural History; the second male, a paratype, is in the Holland Collec-
tion in the Carnegie Museum.
This form, which is near typical preuss?, may be at once distin-
guished from it by the velvety black band which crosses the fore wings
from the base to the outer margin, leaving the diffuse paler subapical
transverse band and the elongated brilliantly green apical area in strik-
ing contrast with the rest of the wing, and by the brilliantly deep green
color of the under side of the wing.
(204) 5e. Euphedra preussi fulvofasciata, new variety
Plate XI, Figure 6, @
This variety differs from the preceding in having the transverse subapical band
on the upper sides of the primaries bright orange in both sexes; the band on the
under side is paler, inclining to white, though in a few specimens in the series before
me the orange color of the upper side reappears. There is a considerable variety in.
the shape of this band, especially in the female sex. There are seven females before
me, and in none of them is the band absolutely the same in outline, and in some of
them it is much broader than in others. The markings on the under side are more or
less variable, as in 2. njamnjami and the other varietal forms which have been men-
tioned.
I assign to this varietal form three males taken at Medje, one in
April, one in July, and one in August, and seven females, all taken at
the same place, two in April and the others from July to September.
In size and in all other respects, except the color and shape of the sub-
1920] Holland, Lepidoptera of the Congo 175
apical band, the form agrees rather closely with E. njamnjami Staud-
inger, from which, however, it may easily be discriminated.
(205) 5f. Euphedra preussi latefasciata, new variety
Plate XI, Figure 8, @
This form is very much like the preceding but differs in having the transverse
subapical band of the fore wing broader, and by the great reduction, amounting to
almost a complete suppression on the under side of the black markings in the cells,
especially in the case of the female. The submarginal markings on the under side of
the secondaries on the other hand are very distinct, inwardly black, margined ex-
ternally by light blue.
This variety is represented by a male captured at Medje in April
and a female taken at the same place in July.
(206) 5g. Euphedra preussi angustior, new variety
Plate XI, Figure 7, @
I apply this name to four males and two females which are characterized by
having the transverse postapical band orange on the upper side of the fore wings
and greatly reduced in width, being much narrower than in any of the other forms in
this group. On the under side the spots in the cell of the fore wing are much enlarged
and very conspicuous, much more so than in any other of the varieties hereinbefore
mentioned. The transverse postapical band in what I take to be the female is not
yellow in this form, as it is in the case of the male, but white.
There are four males, one taken at Medje in April and three captured
at the same place in August. The two females which I associate with
the males because of the markings on the under side of the wings were
also captured at Medje, one in July and the other in August.
Whether the forms which I have mentioned in the foregoing para-
graphs are all really referable to Huphedra preussi and are to be regarded
as mere varieties of that insect can only be decided by the test of breed-
ing. They all agree in having on the anterior margin of the hind wing a
broad white band extending from the base in cell 7 almost to its outer
extremity. This pale band is generally tinged with bluish, but sometimes
inclines to greenish or greenish yellow. There is a general resemblance
among these insects and, if not varietal forms of the same insect, they
represent species which are wonderfully closely related to each other.
(207) 6. Euphedra inanoides, new species
Plate XI: Figure 2, <7, type; Figure 3, 9, allotype (under side)
On the under side of the wings like Huphedra inanum Butler, but with the white
transverse bands not neariy as distinctly defined, especially in the male, and differing
on the upper side from EZ. inanum by having the postapical transverse band of the
fore wings blue, as in E. cwrulescens Grose-Smith, and not pale yellowish white as in
176 Bulletin American Museum of Natural History [Vol. XLIII
E. inanum, specimens of which I have from Sierra Leone, the type locality. In the
female sex the postapical band inclines to whitish or paler blue than in the male sex.
On the upper surface this form so closely resembles EZ. cerulescens Sharpe that at
first glance it might be mistaken for it, but the bluish median band of the fore wing
does not extend as far costad as in E. cerulescens, being restricted to a narrow longi-
tudinal stripe bordering the hind margin of this wing, and of course the markings of the
lower side of the wings are totally different.
There are eight males, which were taken at Medje (type locality)
from June to September, and two females, one captured at Medje in
September and another at Bafwabolion September 10, 1909. The types
co and 2 are in the American Museum of Natural] History; paratypes
in the Holland Collection in the Carnegie Museum.
(208) 7. Eupheedra afzelii (Felder)
Romaleosoma afzelii FELDER, 1867, Reise Novara, Lep., p. 430.
Euphedra afzelii AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 187, Pl. xb, 9 .
I refer to this species a single male captured at Medje, April 6,
1910. The specimen agrees most nearly with Felder’s species, but the
reference is doubtful. In view of the great variability of the forms
under discussion, I do not care to erect a new species upon a single
individual, but it differs almost enough from what I have determined
to be typical EZ. afzelit to justify its separation as a valid variety or
subspecies.
(209) 8. Euphedra phaéthusa (Butler)
Romaleosoma phaéthusa ButTuER, 1865, Proc. Zool. Soc. London, p. 670, fig. 4.
Euphedra phaéthusa Aurtvi.urus, 1913, Seitz, Gross-Schmett., XIII, p. 188.
There are two specimens of this form, both taken at Medje, one in
April, the other in August. It is characterized by the obliteration of the
dark markings on the under side of the wings.
(210) 9. Euphedra ravola (Hewitson)
Romaleosoma ravola Hewrrson, 1864, Exot. Butt., II1, Romaleosoma, Pl. trv, figs.
19-20.
Euphedra ravola Aurtvitiius, 1913, Seitz, Gross-Schmett., XIII, p. 188, Pl. xi,
There are two males and two females which I refer with some
uncertainty to this species. They do not agree perfectly with speci-
mens from the western coastal regions of which I have a long suite.
They are much brighter green upon the lower side of the wings, and
the dark spots and markings are greatly reduced in size imparting to the
under side of the wings a different facies from that of typical 7. ravola.
However, I do not think it expedient to bestow a new varietal name
upon them. They are from Medje.
Otte WY
1920] ° Holland, Lepidoptera of the Congo 177
(211) 10. Euphedra rezioides, new species
Plate XI, Figure 5, @
I am constrained to describe as new two females, which it is impossible to refer
to any of the hitherto described forms belonging to this group. On the upper side
they resemble Z. rezia, 9, and have the postapical band narrow and straight as in
that species. This band is pale orange in the specimens before me, as is also the case
in about half of the females of typical EZ. rezia, the other half, however, in E. rezia
having this band white. On the under side there is a marked difference. The sub-
marginal spots of the fore wing, which are invariably present in E. rezia, are obsolete,
except the one near the inner angle. The cell of the hind wing has one or two black
spots in it and is closed at its extremity by a black bar, which is not found in £Z. rezia,
and beyond the cell in interspaces 3, 4, 5, and 6 there are inwardly pointing sagittate
dark spots arranged in a straight series, after which is a transverse series of pale
markings; of these the one in space 7 is the longest and immediately follows the longi-
tudinal black streak which partly fills the interspace basad, but does not reach the
basal end of this interspace. The other pale spots of the series, which I am describing,
diminish rapidly in size in the direction of the lower margin of the wing. The species
at first glance recalls the pale spots on the under side of EZ. eberti (cf. Aurivillius, Seitz,
XIII, Pl. xurvd), but differs by the existence of the strongly defined dark spots .in
the cell of the fore wing and the dark markings of the hind wings which have been
described. Expanse, 80-85 mm.
The two specimens were both taken at Medje in August. The type
is in The American Museum of Natural History. The paratype is in the
Holland Collection in the Carnegie Museum.
THEMIS GROUP '
(212) . 11. Euphedra adonina (Hewitson)
Romaleosoma adonina Hewirson, 1865, Exot. Butt., III, Romaleosoma, Pl. m,
figs. 11, 12.
Euphedra adonina AurivILuivs, 1912, Seitz, Gross-Schmett., XIII, p. 187, Pl. xu1vb.
Six males and three females, one male caught at Gamangui in J une,
the rest at Medje from July to September.
(213) 12. Euphedra vetusta (Butler)
Romaleosoma vetusta BuTLER, 1871, Lep. Exot., p. 82, Pl. xxx, fig. 5, 9.
Euphedra vetusta AURIVILLIUS, 1912, Seitz, Grossflehimett. III, p. 187, Pl. xxiva, re
With a measure of hesitation I refer to this species three males
taken at Medje in July and August. Butler’s figure is that of a female,
and so also is that given by Aurivillius. The specimens correspond
more nearly with what is known of Butler’s species than with any
other, and I leave them here subject to query.
(214) 13. Euphedra gausape (Butler)
Romaleosoma gausape BUTLER, 1865, Proc. Zool. Soc. London, p. 671, fig. 5.
Euphedra gausape AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 186.
178 Bulletin American Museum of Natural History [Vol. XLIII
There are five males and eleven females before me, not any two of
which exactly agree with each other in all respects, but they are so
closely related to each other that it seems impossible to separate them.
They come nearer to the insect named E. (Romalaosoma) gausape by
Butler than to any other form known to me, and I place them here provi-
sionally. The males and females have the postapical band of the fore
wing orange in color; in the case of the females it is wider than in the
case of the males, and varies somewhat in form and breadth. One
female has this band pure white, but otherwise. I am enable, to distin-
guish it by its markings from the other females.
The specimens were all taken at Medje, a male and a female in
April, the rest in July and August.
XYPETE GROUP
(215) | 14. Euphedra xypete maxima, new variety
There are three males and three females, which are referable to Romaleosoma
xypele Hewitson (cf. Exot. Butt., 1865, III, Romaleosoma, Pl. u, figs. 8-10), but which
differ from specimens in my collection received from Sierra Leone, Cameroon, and
Gaboon, in being much larger in size, by the prevalently bluish (not greenish) cast
of the lighter portions of their wings on the upper side, and by having the spots which
define the inner margin of the postapical band on the under side of the primaries
much narrower and less strongly developed than is the case in specimens from the
localities named. The form may be designated as var. maxima, the specimens aver-
aging fully fifteen per cent more in expanse of wing than specimens taken on the
west coast, of which I have many scores before me as I write.
Male type, Medje; female allotype, Ngayu; paratypes, o’ 2 , Ngayu.
(216) 15. Euphedra cyanea, new species
Plate IX: Figure 1, o; Figure 2, 9
Related to E. cerulescens Grose-Smith, but differing markedly on both the
under and the upper side of both wings. In Z. cerulescens the basal portion of the
fore wing on the upper side is black throughout, except for a few greenish blue scales —
along the edge of the inner margin. In Z. cyanea the basal area is bright blue from
the base to the middle of the cell and thence outwardly to nearly the inner angle of
the wing. Beyond this bright blue area, the outer line of which is quite straight, the
remainder of the wing is rich velvety black, interrupted, however, by a sharply
defined postapical band of bright blue, which is much more sharply defined than is
the case in FE. cwrulescens, and does not extent as far downward toward the inner
margin as is the case in the latter species. In 2. cwrulescens this band, which is green-
ish blue, passing into yellowish toward the costa, reaches vein 2 near the outer border,
but in 2. cyanea the band never extends beyond vein 3, The white spots of the
cilia, which are present in both species, differ, being wider in 2. carulescens, forming
little lunules at the end of the interspaces, but in 2. cyanea they are mere pointlets,
or dots at the middle of the interspaces on the outer margin. The middle of the
ES as
Peery
1920] Holland, Lepidoptera of the Congo 179
upper side of the hind wings in E. cyanea is bright blue, while in Z. cerulescens this
area is dull greenish blue. On the under side in EZ. cerulescens the cell of the
fore wing is solidly blue at the base, in H. cyanea this region is green or greenish
ochraceous. The dark spots in EZ. cyanea which appear in the cell of the fore wing are
much smaller than the corresponding spots in the wing of EZ. cerulescens, and this holds
good also of the spots of the hind wings on the under side. Below the red costal band of
E. cerulescens (Vide Aurivillius, 1912, Seitz, Gross-Schmett., XIII, Pl. xutva,
where the under side of FL. cwrulescens is shown, in error named EF. gausape) large dark
spots intervene between the red costal band and the rest of the wing. This is not the
case in EZ. cyanea. There are no such spots, or they are at most feebly indicated
by a faint dark shade near the basal end of interspace 7. The spots in the cell of the
secondaries are also very greatly reduced or disappear and the discal spots beyond
the cell in HZ. cyanea are smaller and the red of the costal border in some speci-
mens spreads downward into the discal area of the wing, as a faint reddish shade.
The females in E. cyanea are marked exactly as are the males, but have much
greater expanse of wing, and the postapical band of the primaries on the upper side
is paler blue. The postapical band of the primaries is less prominent on the under
side in both sexes of #. cyanea than it is ia E. cerulescens, and in some specimens
scarcely appears. Expanse, o’, 60-65 mm.; 9, 70-85 mm.
Types in The American Museum of Natural History; paratypes in Holland Col-
lection in the Carnegie Museum.
The Expedition brought back fourteen males and five females of
this species, which I have carefully compared with a series of male
specimens of E. cerulescens which I have in my collection from the lower
banks of the Ogové River. The specimens were all captured at Medje,
two males in April, the rest from July to September. Two of the males
are aberrant in that they show a tendency in the postapical band of the
primaries on the upper side to become whitish towards the costa.
(217) 16. Euphedra karschi Bartel
Euphedra karschi Barter, 1905, Nov. Zool., XII, p. 141. Aurrvruxrus, 1912, Seitz,
Gross-Schmett., XIII, p. 186.
The collection contains four males of this well-defined species which
were taken at Medje, two in June, one in July, andone in August. The
specimens agree perfectly with a series in the possession of the writer,
which were collected for him many years ago by Mrs. Reutlinger at
Benito, Spanish Guinea.
Mepon Group
(218) 17. Euphedra medon innotata, new variety
Plate IX, Figure 6, type, @
Papilio medon Linn=vs, 1763, Cent. Ins., p. 19; 1767, Syst. Nat., 12th Ed., p., 753.
Euphedra medon Avrivituius, 1912, Seitz, Gross-Schmett., XIII, p. 184, Pl. xiva.
180 Bulletin American Museum of Natural History [Vol. XLII
Euphedra medon is represented in the collection by a series of seven
males. All belong to a varietal form, long known to me but not
hitherto described, to which I venture to apply the varietal or subspecific
name 7znnotata.
The form is characterized by being paler on the upper side than typical EL. medon,
the coloration being glaucous and less inclined to greenish than in the typical form,
and the dark markings less pronounced, harmonizing more thoroughly with the
ground-color of the wings. The differences on the under side are pronounced and may
be summed up in the statement that the submarginal spots on both the fore and hind
wings are either entirely obliterated or so greatly reduced as to be almost invisible,
and the transverse median light band of the hind wing, which is characteristic of the
typical form, is either wanting entirely or survives merely as a small light-colored
quadrate spot on the costal border.
I have a small series of this form collected upon the Ogové River
by the late Dr. A. C. Good, and they have long been standing in my
collection awaiting a name. On the under side, because of the suppres-
sion of the markings noted above, they are strikingly different from
the typical form. The specimens returned by the American Museum
Congo Expedition were all taken at Medje in June, July, and August.
No other representatives of EZ. medon are in the collection. The type.
a male, and several paratypes are in The American Museum of
Natural History, paratypes are also in the Holland Collection in the
Carnegie Museum. é
(219) 18. Euphedra spatiosa (Mabille)
Romalaosoma spatiosa MasruE, 1877, Bull. Soc. Zool. France, I, p. 278.
Euphedra spatiosa Aurtvitutius, 1912, Seitz, Gross-Schmett., XIII, p. 183, Pl. xive.
There are thirty-six males and thirteen females of this species, all
taken at Medje, except one male captured at Niangara in November.
A few were taken in April, but the majority were caught in June, July,
and August, several bearing the later date of September.
(220) 19. Euphedra losinga (Hewitson)
Romaleosoma losinga Hewr1son, 1864, Exot. Butt., III, Romaleosoma, PI. 1, fig. 5.
Euphedra losinga Aurtvitui0s, 1912, Seitz, Gross-Schmett., XIII, p. 183, Pl. xuvb, ec.
There are two males and two females taken at Medje, one male in
April, the rest in July and August 1910.
EuryPHEeNne Westwood
This is a large genus. There is great dissimilarity between the
sexes in many of the species. A clue to the relationship between the
males and females is, however, generally found by a close examination
1920] Holland, Lepidoptera of the Congo 181
of the markings on the under side of the wings. The females in many of
the groups are wonderfully alike on the upper surface, and it is only
by a careful study of the bandings and markings on the under side of the
wings that it is possible to make a correct reference of the females to
the corresponding males. Many of the females have not yet been figured
in any work, although verbal descriptions of a number of them have been
recently given by Dr. Aurivillius in the latest revision of the genus,
which is contained in Seitz’s ‘Gross-Schmetterlinge der Erde,’ Vol. XIII.
(221) 1. Euryphene carshena Hewitson
Euryphene carshena Hewrtson, 1871, Exot. Butt., IV, Euryphene, Pl. vu, figs. 31,
32, %. AuRIVILLIUs, 1912, Seitz, Gross-Schmett., XIII, p. 171, Pl. xuid, f.
This species is characterized in both sexes by the presence on the
under side of the wings, above the outer upper angle of the cell of the
hind wing, of a broad, very conspicuous brown spot, which shows itself
even more conspicuously in the case of the females than in the case of
the males.
The collection contains four males; two taken in April and two in
September, and four females taken in July and August. All! the speci-
mens were captured at Medje.
(222) 2. Euryphene subtentyris Strand
Euryphene subtentyris StRAND, 1911, Fauna Exot., I, p.37; 1912, Archiv f. Naturg.,
LXXVII, Suppl., 4, p. 118. Avrivitirus, 1912, Seitz, Gross-Schmett.,
XIII, p. 172.
This insect, which may be only a seasonal variety of EH. tentyris
Hewitson, differs from the latter in the entire absence of the brilliant
blue-green coloration of the costal area of the fore wing in the male sex.
In indirect light there is a faint violet sheen discernible upon the disc
of the fore and the hind wings. On the under side of the wings this form
is absolutely like typical tentyris. We have large numbers of this form
collected for us in southern Cameroon.
The species is represented in the Congo Collection by a single male
‘specimen captured at Niangara, November 26, 1910.
(223) 3. Euryphene abesa Hewitson
Euryphene abesa Hewritson, 1869, Trans. Ent. Soc. London, p. 84; 1871, Exot.
Butt., IV, Zuryphene, Pl. vu, figs. 29, 30, o. Auriviuuius, 1912, Seitz, Gross-
Schmett., XIII, p. 172, Pl. xd, e. .
This species is represented by a pair caught at Medje on April
6, 1910.
182 Bulletin American Museum of Natural History [Vol. XLIII
(224) 4. Euryphene absolon (Fabricius)
Papilio absolon Fasrictvs, 1793, Ent. Syst., III, part 1, p. 56.
Euryphene absolon AURIVILLIUS, 1898, Rhop. 4thiop., p. 201, Pl. m1, fig. 5, #7.
There are six males and one female of this species in the collection.
- The female was taken at Medje about the middle of August 1910, and
two males were likewise captured at the same place, one in May, the
other in July. One male was taken at Avakubi, on October 20, 1909,
and two males were caught at Niangara in November 1910.
(225) 5. Euryphene entebize Lathy
Euryphene entebie Latuy, 1906, Trans. Ent. Soc. London, p. 5, Pl. m1, fig. 1, o.
AURIVILLIvs, 1912, Seitz, Gross-Schmett., XIII, p. 172.
Dr. Aurivillius regards this as a varietal form of E. absolon. I differ
from him, however, for the reason that the female which I associate
with the form (the male of which has been very accurately depicted by
Lathy) is more like the female of zonara than the female of absolon;
in fact, in my preliminary arrangement of the species I had referred the
specimens of the suite before me to zonara, marking them as “dark var.”
Typical zonara from the west coast of Africa is a somewhat smaller
insect, the prevalent color of the upper side of the wings being bright
reddish fulvous. The specimens of E. entebie before me are most of
them larger than typical zonara, much darker in color, warm brown,
banded with very dark brown, almost black, and they are all abso-
lutely alike on the under side in having on the hind wings on either side
of vein 6, just beyond the upper outer angle of the cell, a dark shade,
accentuated, as is well shown in Lathy’s excellent figure, by two small
light points, arranged vertically, one on either side of vein 6. The soli-
tary female before me has a dark shade located at exactly the same spot,
with indications of the lighter points. If designated as a variety of any
of the well-known species already described, it seems to me it should be
of zonara rather than of absolon. For the present I recognize it as a
distinct species.
The specimens vary a little in size.. Held in a very oblique position,
they reveal a very slightly purplish blue iridescence, in this respect being
somewhat like E. absolon var. micans Aurivillius, though in the latter
the blue iridescence is much more evident.
The collection contains twelve males and one female, which were
all taken at Medije, the dates of capture including the months of April,
July, August, and September.
EE ES Se
1920] Holland, Lepidoptera of the Congo 183
(226) 6. Euryphene zonara (Butler)
Aterica zonara BUTLER, 1871, Proc. Zool. Soc. London, p. 81; 1871, Lep. Exot., p. 72,
Pl. xxviu, figs. 1, 2.
There are three males, all taken at Medje, one in April, the other
two in September, and one female caught in the month of July. The
figure of the under side of the wings of a male given by Aurivillius in his
‘Rhopalocera A&thiopica,’ Pl. m1, is that of a specimen in which the
characteristic markings are not as strongly accentuated as in the speci-
mens upon which I am reporting.
(227) 7. Euryphene lucasi, new species
Plate IX: Figure 4, @, type; Figure 5, 9, allotype
o&. Upper side of both wings reddish fulvous marked with bands and spots of
dark brown. Fore Wina: a dark spot filling the base of the cell, succeeded by a
narrow angulated line, which in turn is followed by a large spot having the outline
of the figure 8, succeeded by a second narrow angulated line, and this in turn followed
- at the end of the cell by a large spot, which has its inner margin straight, but is
sinuate on its outer margin; below the cell the basal area as far as the first mesial
outer band of light spots is dark brown; the dark spot at the end of the cell and
the dark area below the cell are succeeded by a mesial band of light spots, which,
beginning on the costa beyond the end of the cell, runs first transversely toward the
outer margin as far as vein 6, then turns abruptly and sweeps backward and
downward to the inner margin of the wing which it reaches less than half its
distance from the base. Beyond this series of light spots is a series of dark subquad-
rate spots gradually increasing in width from the costa as far as the first submedian
nervule, and then rapidly diminishing toward the inner margin. This band is suc-
ceeded by a series of light lunate spots convex basad as far as vein two, and continued
to the inner margin in a narrow curved line parallel to the line of the lower part of the
first median series of light spots. This last line of light spots is followed by a series
of subcircular dark spots on the interspaces extending from vein 6 to vein 1, below
which the last spot of the series is quadrate. Beyond these spots there is another
series of light sublunate spots convex marginad. The effect of the arrangement of
the last-mentioned two series of light lines with their reversed curvatures is to produce
the appearance of eyelike markings, with a dark pupil partly surrounded by red.
The marginal border of the wing is dark, but not as dark as the inner series of dark
bands and spots. At the apical extremity of the wing there is a general dark suffu-
sion in which the spots and bands lose themselves. The cilia are dark brown, check-
ered at the middle of each interspace with white. Hinp Wtna: fuscous on the inner
margin, with the area below the cell for a short distance, and the base of the gell dark
brown; the middle of the cell isreddish, marked with a figure 8 and a thin dark line at
its end, both darker; beyond the cell is a dark shade or band, interrupted by a
mesial band of bright reddish brown, which, beginning as a point in interspace 7,
rapidly widens to vein1, where it ends abruptly. The two bands of light lines en-
closing cireular dark spots, found on the primaries, are continued upon the
secondaries and the marginal borders and cilia of the latter are much as on the former.
The head, thorax, and abdomen are blackish above, as are the palpi; on the lower
184 Bulletin American Museum of Natural History {Vol. XLIII
side they are grayish. The under side of both wings is pale gray, and all the mark-
ings of the upper side are reproduced with modifications, most of them being greatly
reduced in size, and those of the cells and the discal areas being pale but sharply de-
fined externally by fine dark lines.
Q. The female is larger than the male, as is always the case in this genus, and
has greater expanse of wing. The arrangement of the spots and lines is substantially
the same as in the male sex, with this difference that the light red spots of the upper
side are prevalently light gray, except the inner mesial and the two succeeding series
of light spots, which in this sex are light lemon-yellow, and the former of which on the
secondaries is greatly expanded, forming a large diffuse yellow band occupying the
middle of the wing. On the under side the markings near the end of the cell are
compacted to form a very irregular dark band sharply defined externally, and having
as one of its characteristic marks a narrow projection at the origin of the second
and third submedian nervules. This is also to be detected in the male sex, and is
a clue to the relationship of the two sexes. Expanse: o’, 50-55mm.; 9, 60-65 mm.
Types in The American Museum of Natural History; paratypes in the Holland
Collection in the Carnegie Museum.
The species may easily be distinguished from all hitherto described
by the narrow band of light yellow spots which crosses the primaries oi
the female, as stated in the description, and which corresponds in its
location to the inner band of fulvous spots in the male.
I take pleasure in naming this species in honor of my friend of many
years, Dr. F. A. Lucas, the Director of The American Museum of Natural
History.
The collection before me contains three males and three females,
all captured at Medje, one female in April, the rest from June to Sep-
tember.
(228) 8. Euryphene mandinga Felder
Euryphene mandinga Fevver, 1860, Wien. Ent. Monatschr., [V, p. 108. AuRIVILLIUs,
1898, Rhop. Aithiop., p. 201, Pl. m1, fig. 7, o.
Of this-well known species there are four males and three females, all
taken at Medje in August, except one female taken on September 1,
1910.
(229) 9. Euryphene oxione Hewitson
Euryphene orione Hewrrson, 1866, Exot. Butt., III, Zuryphene, Pl. v, text; 1871, loc,
cit., 1V, Euryphene, P|, vint, figs. 36, 37, o. Dewrrz, 1866, Berl. Ent. Zeit., XXX,
Pl. vu, figs. 1,2, 9. Avrivittrus, 1912, Seitz, Gross-Schmett., XIII,
p. 172, Pl. xute.
The collection contains two males and five females of this well-
marked species, all taken at Medje, the dates of capture including the
months of April, June, August, and September.
1920] Holland, Lepidoptera of the Congo 185
(230) 10. Euryphene partita Aurivillius
Euryphene partita Aurivituivs, 1895, Ent. Nachr., X XI, p. 380.
Euryphene aurivillii SraupinGeEr, 1896, Iris, VIII, p. 371, Pl. vit, fig. 3, &.
Euryphene partita AuRIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 172, Pl. xuic, @.
Of this easily recognized species the collection contains four males
and eleven females, all taken at Medje, a couple of females captured in
April, and all the remainder taken in the months of July and August.
It may be worthy of note that the two females captured in April
differ from the females taken later in the year in that they are without the
bluish iridescence which is found upon the fore wings of the latter, indi-
cating a seasonal variation in this respect.
(231) 11. Euryphene iturina Karsch
Euryphene iturina Karscu, 1894, Ent. Nachr., XX, p. 215. Aurtvi.urus, 1898, Rhop.
AXthiop., p. 200, Pl. m1, fig. 1, 9; 1912, Seitz, Gross-Schmett., XIII, p. 173,
Pl..xuib, co’, 9.
The collection contains twenty-two males and seventeen females.
The specimens were all taken at Medje, except two males which are
ticketted as captured at Gamangui in June. The labels attached to
the specimens taken at Medje show that a few were captured in April,
but by far the greater number were taken in August and September.
(232) 12. Euryphene fulgurata Aurivillius
‘ Plate X, Figure 11, 9
Euryphene fulgurata Aurtivituus, 1904, Ent. Tidskr., XXV, p. 95, fig. 35; 1912,
Seitz, Gross-Schmett., XIII, p. 173.
Two females taken at Medje, one in July, the other in August.
(233) 13. Euryphene congolensis Capronnier
Euryphene congolensis CAPRONNIER, 1889, Ann. Soc. Ent. Belgique, XX XIII, Bull.,
p. CXxii. AuRIVILLIUs, 1898, Rhop. ASthiop., p. 200, Pl. m1, figs. 8,9, 7, @.
This species is represented in the collection by thirty-five males and
thirty-three females, all of which appear to have been taken at Medje,
except one female caught at Stanleyville on August 18, 1909, and two
females taken at Gamangui on June 17, 1910. The dates attached to
the specimens captured at Medje show that a few were caught in April,
but the great majority were taken in the months of July, August, and
September.
(234) 14. Euryphene phranza Hewitson
Euryphene phranza Hewitson, 1865, Exot. Butt., III, Euryphene, Pl. u, figs. 7, 8.
AURIVILLIUs, 1912, Seitz, Gross-Schmett., XIII, p. 173, Pl. xb.
186 Bulletin American Museum of Natural History [Vol. XLIII
This species is represented by a male and female, the former cap-
tured at Medje on May 25, the latter taken at the same place on June
21.
(235) _ 15. Euryphene severini Aurivillius
Euryphene severini Aurtvitiius, 1897, Ofvers. Sv. Vet.-Akad. Férh., LIV, part 1,:
p. 280, fig. 2, 2; 1898, Rhop. Aithiop., p. 200, Pl. m1, fig. 10, 7.
Of this species there are thirteen males, all labelled as captured at
Medje in June, July, and August. With these males I associate five
females, taken at Medje in the same months, which agree almost
absolutely with the figure of this sex given by Aurivillius (loc. cit.),
save that in the specimens before me the apex of the fore wing is some-
what more pointed and not of precisely the shape given in the wood cut
supplied by Aurivillius. I can see no difference in other respects; the
markings are identical and, in spite of the fact that the fore wing of
these females is somewhat more like the female of EL. sophus in showing a
tendency to become falcate at the tip, I am reasonably certain that my
reference is correct. It is to be observed that, while the fore wings are,
as I have said, faleate, they are rounded at the extreme apex and not
sharply acuminate as in sophus.
(236) 16. Euryphene letitia Ploetz
Euryphene letitia Putz, 1880, Stett. Ent. Zeit., XLI, p. 192. Aurtviturus, 1898,
Rhop. A&thiop., p. 200, Pl. 1, fig. 2; 1912, Seitz, Gross-Schmett., XIII, p. 174,
Pl. x1Ic, o, 9.
I refer to this species two males taken at Medje on May 6, 1910.
While agreeing pretty thoroughly on the upper side with the figure of
the male given by Aurivillius, there are some minor discrepancies on the
under side of the wing but, without more material before me, I do not
feel justified in differentiating the insect from £. letitia, with which it
agrees more closely than with any other species hitherto described.
In passing, I must call attention to the fact that Dr. Aurivillius is
quite in error in sinking my £. castanea (cf. Canadian Entomologist,
1893, XXV, p. 1) as asynonym of EZ. letitia. It is totally distinct and,
had Dr. Aurivillius known it other than by the verbal description I
published, he would not have included it under £, letitia.
(237) 17. Euryphene sophus (Fabricius)
Papilio sophus Fasnicivus, 1793, Ent. Syst., III, part 1, p. 46.
Euryphene sophus Avnivit.ii0s, 1912, Seitz, Gross-Schmett., XIII, p. 174, Pl. xure.
Two males and two females captured at Medje are referable to this
species. One of the males was caught on August 1, the other on Sep-
1920] Holland, Lepidoptera of the Congo 187
tember 27. One of the females was taken on August 13, the other on
July 29. The latter female, which has the pre-apical band white, is
referable to the varietal form named phreone Feisthamel. Except for
the difference in the color of the transverse postapical band there is no »
difference whatever between this form and true E. sophus (Fabricius).
(238) 18. Euryphene phantasia Hewitson
Euryphene phantasia Hewirson, 1865, Exot. Butt., III, Zuryphene, Pl. u, figs. 9-11.
AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 176, Pl. xu1a.
Three males and three females. One female which has the post-
apical band of the primaries yellowish instead of white, as is the case
with the other two, was taken at Ngayu in December, 1909. All the
other specimens were captured at Medje, one male in April, the rest in
July and August.
(239) 19. Euryphene flaminia Staudinger
Euryphene flaminia Svaupincer, 1891, Iris, IV, p. 110, Pl. 1, fig. 4. AURIvILLIUs,
1912, Seitz, Gross-Schmett., XIII, p. 176, Pl. xte.
I refer to this species two males and a female taken at Medje, the
female in April, the males in July and August.
(240) 20. Euryphene maximiniana Staudinger
Plate VII, Figure 10, 9
Euryphene maximiniana SiauvinGER, 1891, Iris, IV, p. 112. Avurtviturus, 1912,
Seitz, Gross-Schmett., XIII, p. 176.
Four males and three females are referable to this species, which
hitherto has only been reported from Cameroon. Six specimens were
taken at Medje in June, July, and August, and one at Gamangui.
(241) 21. Euryphene mardania (Fabricius)
Papilio mardania Fasricivs, 1793, Ent. Syst., III, part 1, p. 249.
Euryphene mardania Aurtvituius, 1912, Seitz, Gross-Schmett., XIII, p. -177, Pl.
XLe, d.
Of this common species there are six males and four females. Two
of the males and one of the females were taken at Medje in July and
August, the rest were captured at Niangara in November 1910.
(242) 22. Euryphene plistonax Hewitson
Euryphene plistionax Hewson, 1874, Exot. Butt., V, Zuryphene, Pl. 1x, figs. 38, 39.
AURIVILLIUs, 1912, Seitz, Gross-Schmett., XIII, p. 177, Pl. xe.
Of this large and striking species there are two males and five fe-
males taken at Medije, the dates of capture ranging from June to August.
188 Bulletin American Museum of Natural History [Vol. XLIII
(243) 23. Euryphene barce (Doubleday)
Aterica barce DouBLEDAY, 1847, Proc. Zool. Soc. London, p. 59.
Euryphene lesbonax Hewrison, 1864, Exot. Butt., IIl, Huryphene, Pl. 1, figs. &, 6, #7.
Euryphene barce var. achillena Bartel, AurtviLuius, 1912, Seitz, Gross-Schmett.,
XIII, p. 179, Pl. xub, 9.
This species is represented in the collection by a single female, be-
longing to the varietal form named achillena by Bartel. It was caught
at Medje, May 6, 1910.
(244) 24. Euryphene chloéropis Bethune-Baker
Euryphene chloéropis BerHUNE-BAKER, 1908, Ann. Mag. Nat. Hist., (8) II, p. 474.
AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 178.
I refer a single male captured at Medje in the early part of April to
this species, with the description of which it appears to agree very closely,
except in the expanse of wing. Baker records 72 mm. for the spread of
the specimen he described, while the individual before me has a spread
of only 58 mm.
(245) 25. Euryphene luteola Bethune-Baker
Euryphene luteola BetHunE-BakeEr, 1908, Ann. Mag. Nat. Hist., (8) II, p. 474.
AURIVILLIvs, 1912, Seitz, Gross-Schmett., XIII, p. 179.
There are two males which agree very closely with the description
given by Bethune-Baker and which I accordingly refer to this species.
There are also two females which appear to me to be undoubtedly that
sex of this species. As the female of Z. luteola awaits description, I may
briefly point out that, as is always the case in this genus, they exceed
the males in expanse of wing. The prevalent colors of the wings are
paler on the upper side, the light areas inclining to blue and not to
green as in the male. On the under side both wings are pale reddish
brown, the base of the costal margin blue as in the male, and the other
spots and markings as in the male but more diffuse and expanded. As
in the species listed just before this, there is a discrepancy in the matter
of the expanse of wing between the individuals before me and the male
described by the author of the species. Bethune-Baker gives 77 mm.
as the expanse, but the two males before me measure not more than 57
mm., while the females have a spread of from 68 to 70 mm.
The specimens are from Medje.
(246) 26. Euryphene rubrocostata Aurivillius
Euryphene rubrocostata Auntvit.ivs, 1897, Ofvers. Sv. Vet.-Akad. Férh., LIV, 5, p.
279, fig. 1, 9; 1898, Rhop. Acthiop., p. 197, Pl. 1, fig. 5, o.
1920] Holland, Lepidoptera of the Congo 189
The collection contains fifty males and four females, all taken at
Medje, a couple in April, and all the rest including all the females, from
July to September.
Diestoeyna Karsch
(247) 1. Diestogyna camarensis (Ward)
Euryphene camarensis Warp, 1871, Ent. Mo. Mag., VIII, p. 35.
Diestogyna camarensis AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 159, PI.
xxxixd.
A single male taken at Medje, in September 1910,
(248) 2. Diestogyna goniogramma Karsch
Diestogyna goniogramma Karscu, 1894, Berl. Ent. Zeit., XXXIX, p. 5, fig. 1.
AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 160, Pl. xxxrxd.
Of this species there are thirteen males and four females, all taken
at Medje, a pair in April, all the rest from July to September.
(249) 3. Diestogyna mawamba Bethune-Baker
Diestogyna mawamba BeTHUNE-BakER, 1908, Ann. Mag. Nat. Hist., (8) II, p. 476.
AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 163, Pl. xxxvime.
To this comparatively recently described species I refer a male
taken at Medje on August 11, 1910 and a female taken at the same
place on July 17, 1910. While the specimens slightly differ from the
description given by Bethune-Baker, they nevertheless agree so thor-
oughly with the figures given by Aurivillius that I think I am right in
my determination.
(250) 4. Diestogyna tadema (Hewitson)
Aterica tadema Hewitson, 1866, Exot. Butt., III, Pl. Aterica and Harma, figs. 10-
12, o, 2.
I refer to this species a male taken at Medje, on August 1, anda
female taken on July 21, 1910.
(251) 5. Diestogyna saphirina Karsch
Diestogyna saphirina Karscu, 1894, Ent. Nachr., XX, p. 220. Aurrviiirus, 1912,
Seitz, Gross-Schmett., XIII, p. 166, Pl. xxxvuf.
I refer to this species a series of eighteen males and fourteen females,
taken at Medje, a few in April, the rest from July to September.
(252) 6. Diestogyna melanops Aurivillius
Diestogyna melanops Avrivituius, 1897, Ofvers. Sv. Vet.-Akad. Forh., LIV, 5,
p. 282; 1912, Seitz, Gross-Schmett., XIII, p. 168, Pl. xxxvue.
190 Bulletin American Museum of Natural History [Vol. XLII
This species is represented in the collection by three males, all
taken at Medje, one in April, one in August, and one in September.
(253) 7. Diestogyna doriclea infusca (Capronnier)
Euriphene (sic) infusca CaPRONNIER, 1889, Ann. Soc. Ent. Belgique, XXXII,
Bull., p. exlv.
Diestogyna doriclea infusca AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 168.
This variety of D. doriclea (Drury) (ef. Ill. Exot. Ent., 1782, III,
p. 50, Pl. xxxv1, figs. 5, 6), which occurs sporadically in collections which
we have received from the tropical western coast, appears to be, as
Aurivillius has already pointed out, the dominant form in the interior
of the Congo Basin. It is represented in the collection by eight males
and two females, all taken at Medje, a isdn in April, the rest in August
and September 1910.
(254) 8. Diestogyna amaranta Karsch
Diestogyna amaranta Karscu, 1894, Berl. Ent. Zeit., XTX, p. 6, fig. 2. AURIVILLIUs,
1912, Seitz, Gross-Schmett., XIII, p. 169, Pl. xxxvutrf.
There are two males of this species which agree perfectly with the
figure and description given by Karsch. They were both captured at
Medje in September 1910.
(255) 9. Diestogyna gambie (Feisthamel)
Euryphene gambie FristHamMe., 1850, Ann. Soc. Ent. France, (2) VIII, p. 251, Pl.
Ix, fig. 2.
Diestogyna gambiae AuRIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 169, Pl. xxxviia.
Of this well-known and widely distributed species there are eight
males and nine females, all taken at Medje, a pair in April, the rest in
August 1910.
(256) 10. Diestogyna atossa (Hewitson)
Euryphene atossa Hewrtson, 1865, Exot. Butt., III, Zuryphene, Pl. 11, figs. 1, 2, 2.
Aterica amaxia Hewi1rson, 1865, Exot. Butt., III, Aterica and Euryphene, Pl. v1, figs.
8, 9,9.
Diestogyna atossa AuRIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 169, PI.
xxxvuila, b,
Of this familiar species there are five males and three females, all
taken at Medje in August and September.
In addition to the ten foregoing species which are referable to forms
already known and described, I find in the collection two specimens
which have given me a good deal of trouble and which I can not refer to
any species the descriptions and figures of which are known to me,
1920] Holland, Lepidoptera of the Congo 191
They belong apparently to what Dr. Aurivillius designates as his
“Second Group” in this great genus. While I dislike very much to
found a specific description upon single individuals, I nevertheless am
constrained to do so in this case, as the insects are so evidently distinct
from all other forms which have been described.
(257) 11. Diestogyna kahli, new species
Plate XII, Figure 13, 7
&. Closely allied to D. amaranta Karsch, but having the hind wing somewhat
narrower than in the species described by Karsch, and differing totally in the distribu-
tion and shade of the blue color on the upper side of the wings. In D. amaranta
the blue color is evenly distributed over the entire surface of the fore wing and shows
in certain lights a tendency to play into green. In D. kahli the blue of the wings on
the upper side is deep pavonine blue, or intense cobalt, confined on the anterior
-wing to a broad patch above the lower margin near the lower angle, and not invading
the cell; the cell and the remainder of the wing being dark velvety brown. There is also
in the type no indication of the presence of the small subapical pale dots which are
shown in the figure given by Karsch as characterizing his species. On the under side
of the wings there is a resemblance in the markings to those of amaranta Karsch, and
mawamba Bethune-Baker, but in D. kahli the outer area of the fore wing is much
lighter and is sharply contrasted with the darker basal area which sends forth
tooth-like projections in a regular series on each nervule from the costa to vein 2.
The hind wing is marked very much as in amaranta, but is richer and redder in tone,
the basal and inner areas being laved with dark maroon. The fringes of both the fore
and the hind wings are narrowly pure white on the upper sides, not so conspicuous
on the lower side. In D. amaranta Karsch the white border of the hind wing .
seems to be restricted to small white dots at the middle of each interspace. Expanse,
45 mm.
The type, which is unique, was taken at Medje and is in The American Museum
of Natural History.
I take pleasure in naming this beautiful species after my friend and
associate, Mr. Hugo Kahl, who has assisted me in arranging the collec-
tion upon which I am reporting.
(258) 12. Diestogyna rotundata, new species
Plate XII, Figure 14, #7
&. This species in the matter of form is distinguished by the relatively great
expanse of the hind wings, which, in their rounded outline and breadth, suggest the
outlines of the females of this genus, rather than the males. The species in the outline
of the wings comes nearer D. tadema in this respect, but the wings are relatively much
broader and more fully rounded in outline than is the case even in that species. The
wings on the upper side are brown, shot with pale lilacine blue over most of the surface
of the primaries and upon the inner half of the secondaries, which have the costal
area very broadly pale brown, shading on the costal margin into pale fuscous. The
inner margin, as far as vein 1, is also fuscous. In the fore wings there are the usual
192 Bulletin American Museum of Natural History [Vol. XLIII
dark transverse markings in the cell. An obscure dark band runs from the costa to the
end of the cell outwardly and downwardly to near the lower angle of the cell, and then
passes perpendicularly downward toward the inner margin, which it does not reach,
being lost in the general ground-color of the wing. This dark band is followed out-
wardly by a paler bluish band succeeded by a dark shade extending from the costa
toward the inner margin. Beyond this there are five or six light points, one on each
interspace, these points are succeeded externally by a series of dark quadrate spots
on the interspaces. There is a deep, dark brown submarginal band running evenly
from just before the apex to the lower angle of the wing. The fringes are concolorous.
The hind wing on the upper side is traversed by three bands of darker color, which are
most strongly accentuated on the inner half of the wing and which fade out toward the
costal area. The dark submarginal band which is found on the primaries reappears
upon the secondaries, being most intense opposite the end of the cell. The wings on
the lower side are pale reddish brown, variegated with darker brown spots and bands.
The fore wings are paler in color than the hind wings; the basal area is pale chestnut-
brown with an oval brown spot near the base of the cell and two transverse, quite
broad, dark spots, one near the middle of the cell and the other at its end. ° All of these
spots in the cell are defined outwardly by very narrow pale lines. A very pale light
transverse band marks the division between the darker basal area and the
lighter outer area of the wing. The series of small light-colored points near the apex
of the wing on the upper side reappear on the lower side, where they are much more
distinct than on the upper side, and are accentuated externally by deep brown shades.
The submarginal band of the upper side appears quite faintly on the lower side, and
the outer margin from vein 2 to vein 7 is laved with dark chestnut-brown.
The hind wings on the lower side have the basal area darker than is the case in
the fore wing, and are distinctly defined externally by a band of pale lilac. There is a
minute dark spot in the middle of the cell, pupiled with lighter color and surrounded
by yellowish scales. The outer half of the hind wing is more or less deep chestnut-
brown, with a regular evenly curved row of six sharply defined white spots located,
at the middle of the interspaces from just behind the costa, terminating before vein 2.
The submarginal dark band of the secondaries reappears on the lower side in a series
of dark lunulate markings parallel to the outer border but not quite reaching the
inner border.
The palpi on the upper side are dark, on the lower side pale brown. The thorax
and the abdomen on the lower side are pale brownish white; on the upper side they
are dark brown.
The lower side of the wings recalls in some respects the markings of the under
side of the species named wmbrina by Aurivillius, but in the outline of the wings, the
color and shape of the markings, it is very different, and assuming that the figure of
the species given by Aurivillius in Seitz’s work is adequate, I cannot reconcile myself
at all to refer the insect before me to that species. Expanse, 53 mm.
The type, which is unique, was taken at Medje. It is deposited in The Amer-
ican Museum of Natural History.
EuryPHuRA Staudinger
This genus is but poorly represented in the collection, as is usually
the case, for the insects do not appear to be common. It is a difficult
genus, both because of the dissimilarity of the sexes and the variabilitv
of the females, which are polymorphic in some species.
1920] Holland, Lepidoptera of the Congo 193
(259) 1. Euryphura achlys (Hopffer)
Harma achlys Horrrer, 1862, Peters, Reise n. Mossambique, Ins., p. 390, Pl. xxu,
figs. 5, 6.
There is one female, taken at Niangara in November, which agrees
well with the figure given by Hopffer, and with specimens in my collec-
tion from Zanzibar.
(260) 2. Euryphura chalcis (Felder)
Harma chalcis Feivrr, 1860, Wien. Ent. Monatschr., IV, p. 234.
Euryphura chalcis AuRivILui0s, 1912, Seitz, Gross-Schmett., XIII, p. 157, Pl. xxxvid.
To this species I refer four males and an aberrant female. One
male and the female were taken at Medje in August, the other three
males were captured at Niangara in November.
(261) 3. Euryphura plautilla albofasciata Staudinger
Euryphura albofasciata StaupiINGER, 1896, Iris, LX, p. 213.
I refer to this form a single female taken at Bafwabaka, January 7,
1910. It agrees quite thoroughly with the description given by Staud-
inger.
CymorHoE Hiibner
The genus Cymothoé is characteristic of the hot, wooded lands of the
Ethiopian subregion, to which it is confined. It offers a number of difti-
culties to the student because of the great dissimilarity of the sexes in
most of the species and the further fact that many species in the female
sex are known to be polymorphic. Until the test of breeding the various
species shall have been made, there will necessarily remain a measure of
uncertainty as to the true relationship of various forms, some of which
have been already named and described in one or the other sex and some
of which prudent students have been holding in the hope of obtaining
more light. '
The genus is rich in species, some of which are among the most
beautiful insects found in the region they inhabit. The American
Museum Congo Expedition returned good series of a number of the com-
moner forms and, in addition, three species which I believe hitherto
undescribed, the males of C. angulifascia Aurivillius and of C. aramis
(Hewitson), of which hitherto only the females have been known, and
other material which helps to confirm conclusions, already reached by
the writer with the help of collections made for him in other parts of
_ the general region, which in some respects are at variance with the pub-
194 Bulletin American Museum of Natural History [Vol. XLIII
lished opinions of some of his friends, among them Dr. Aurivillius, who
has given us in Seitz, ‘Gross-Schmetterlinge’, Vol. XIII, the latest revi-
sion of the genus. These things will be touched upon in what I shall
have to say in regard to the several species in the following pages.
(262) 1. Cymothoé theobene (Doubleday and Hewitson)
Harma theobene DouBLEDAY AND HEwitson, 1850, Gen. Diurn. Lep., II, Pl. xt, fig. 3.
Cymothoé theobene AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 144, Pl. xxxrvd.
There are one hundred males and twenty-four females of this species
in the collection. Most of them were taken at Medje, a few in April,
the rest from June to October. A number are from Niangara, captured
in November, and there are other individuals labelled as coming from
Ngayu, Gamangui, and Munie Katoto.
This is one of the commonest species of the genus, and almost
every collection from tropical Africa contains at least a few examples.
There is considerable variation in the size and marking of individuals,
especially in the case of the females. I find, however, no female as large
and as darkly colored as some I have from the Ogové River, the outer
third of the wings in these specimens from the Congo being paler and
the dark spots smaller in size than is the case with the material from
nearer the western coast. There are several male specimens from
Niangara and one from Medje which come near to the form named C.
blassi by Weymer in which the transverse Ifght band on the primaries
of the males is not sharply defined outwardly near the costa but fades
insensibly into the general ground-color. The form is scarcely worthy of
a varietal name, though it can easily be discriminated and, in fact, is the
prevalent form on the eastern coast, whence I received a series some
years ago, which were collected for me by the late William Doherty on
the hills back of Mombasa.
(263) 2. Cymothoé reinholdi (Ploetz)
Harma reinholdi Putz, 1880, Stett. Ent. Zeit., XLI, p. 194 (7).
Cymothoé reinholdit Aurivitiius, 1894, Ent. Tidskr., XV, p. 305 ( 9); 1898, Rhop.
AXthiop., p. 211, Pl. rv, figs. 6, 7 (cand 9).
There are three males and one female, all taken at Medje, a pair
in July and two males in August.
(264) 3. Cymothoé theodosia Staudinger
Cymothoé theodosia Siacvinorer, 1889, Stett. Ent. Zeit., L, p, 416. AuRtvi.itvs,
1912, Seitz, Gross-Schmett., XIII, p. 146, Pl. xxxvia, 9.
Of this beautiful insect, stated by Staudinger to be a local race of
C. beckeri Herrich-Scheffer, there are thirty-two males and eleven fe-
OE a eS le
1920] Holland, Lepidoptera of the Congo 195
males. They were all taken at Medje, except a pair caught at Munie
Katoto and a male taken at Risimu in September 1909. One rather
dwarfed specimen is recorded as taken in April, the rest from July to
September, but by far the larger number in the month of August. The
insect is not a race of C. beckeri Herrich-Scheffer, but undoubtedly a
valid species.
(265) 4. Cymothoé confusa Aurivillius
Cymothoé confusa Auriviiivs, 1887, Ofvers. Sv. Vet.-Akad. Férh., XLIV, p. 310;
1912, Seitz, Gross-Schmett., p. 146, Pls. xxx1va, b, and xxxvia,
One male taken at Niangara, November 20, 1910.
(266) 5. Cymothoé colmanti Aurivillius
Cymothoé colmanti Avurivitiius, 1898, Ent. Tidskr., XIX, p. 180, fig. 6; 1912,
Seitz, Gross-Schmett., XIII, p. 147.
A single male captured at Gamangui, February 4, 1910, is referred
to this species. It corresponds very closely to the description and figure
given by Aurivillius, and the very slight differences may be due to the
fact that the specimen is a trifle worn and the marginal borders of the
wings are not quite as distinctly marked as in the published figure.
(267) 6. Cymothoé cyclades (Ward)
Plate VII, Figure 6, 7
Harma cyclades Warp, 1871, Ent. Mo. Mag., VIII, p. 119; 1874, Afr. Lep., p. 14,
Pl. x1, figs. 4, 5.
Cymothoé cyclades AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 147, Pl. xxxrvb,
o.
One male taken at Medje, July 8, 1910. I refer this specimen to
Ward’s species in spite of some minor, apparently individual, differences.
It agrees very closely with specimens which I have from the tropical
western coast and which I do not hesitate to identify with C. cyclades.
(268) 7. Cymothoé diphyia Karsch
Plate VI, Figure 6, 9
Cymothoé diphyia Karscu, 1894, Ent. Nachr., XX, p. 211. Avrrvinurus, 1912,
Seitz, Gross-Schmett., XIII, p. 149, Pl. xxxva, .
Ten males caught at Medje, one in April, the rest in July and
August, and one female, which I believe I am right in associating with
the males, though it does not quite agree with the brief characteriza-
tion of the female given by Aurivillius (loc. cit.). As no figure of the
female of C. diphyia has thus far been published and all we know about
196 Bulletin American Museum of Natural History [Vol. XLIII
that sex is contained in the couple of lines printed by Aurivillius, it
seems to me fitting that I should give a figure of the insect which I have
determined to be the female of this species. The female assigned to C.
diphyia by Karsch is unmistakably the female of C. theodosia, as has
already been pointed out by Aurivillius. The species is closely related
to C. fumana (Westwood). It differs in the male sex in having the outer
third of the fore wing on the upper side tinged with ochreous (in C.
fumana the apical third is creamy white) and in having the outline of the
dark inner basal area of the fore wing straight or concave costad, while
in C. fumana this area is strongly convex on the margin toward the
costa.
(269) 8. Cymothoé herminia Grose-Smith
Plate VIII: Figure 1, &%; Figure 4, 9
Cymothoé herminia Grose-Smitu, 1887, Ann. Mag. Nat. Hist., (5) XIX, p. 63.
AvRIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 149, Pl. xxxva, o.
There are eighteen males of this species, all taken at Medje in July
and August. With these I associate a female, which I believe to be that
sex of the species, and which I cause to be figured, as no description or
figure of the female has heretofore been published.
(270) 8a. Cymothoé herminia poénsis, new variety
Plate VIII, Figure 2, 7
All the specimens of C. herminia taken by the American Museum
Congo Expedition conform to the figures and descriptions of this species,
which have been published by Grose-Smith and by Aurivillius, and are
typical, having the dark band which outwardly defines the pale middle
band of the primaries incomplete at its upper extremity, thus leaving the
costal margin broadly of the same color as the middle of the wing, except
at the tip, on which the outer marginal border is continued. There is,
however, in my possession a male specimen of this species which was
taken on Fernando Po for me by the late Dr. A. C. Good, in which the
black band above mentioned extends all the way to the costa, and in
which the dark submarginal markings are heavier than in the specimens
from the Congo upon which I am reporting. To this varietal (insular)
form I propose to give the name poénsis, and take the opportunity to
figure it. Type in Holland Collection, Carnegie Museum.
EE —————— eS
1920]. Holland, Lepidoptera of the Congo 197
(271) 9. Cymothoé langi, new species
Plate VIII: Figure 7, &%; Figure 8, 9
¢@. This species, which is closely allied to C. weymeri, C. staudingeri, and C.
johnstoni, is easily distinguished from all of them by the fact that the light sagittate
markings which cap internally the black dart-shaped submarginal spots on the inter-
spaces are entirely suppressed on the hind wings, and also on the fore wings except in
space 2, where the lower limb of one of these markings survives as a narrow line point-
ing downwardly and outwardly, and on space 5, where there is an oblong light spot,
slightly bifid at its outer extremity, and on space 6, where there is a faint linear streak
of lighter scales on the dark ground of the broad outer margin. On the under side
the mesial white band, which is much whiter than on the upper side, is narrower.
and tapers from the costa of the fore wing to nearly the anal angle of the hind wing,
being sharply defined outwardly by a thin almost straight dark line. Expanse,
60-63 mm.
Q. With the males I associate, I believe correctly, a female, which rather
closely resembles that sex of C. hewitsoni Staudinger. It differs from C. hewitsoni 9
in not having the white bar at the outer end of the cell of the fore wing and in not
having the small black bar which occurs in C. hewitsoni at the end of the.cell of the
hind wing. On the under side a difference presents itself in the fact that the inner
dark area is defined outwardly by a straight line in C. langi, while in C. hewitsoni it
is inwardly curved or concave on the secondaries, and at the point where this dark
area terminates abruptly on the lighter ground of the fore wings in C. hewitsoni there
is in-C. langi a thin dark line, which extends forward until it reaches the costal margin,
about one-third of the distance from the apex of the fore wing. Expanse: <7, 60
mm.; ?, 70 mm.
The type o’, and allotype 2, are in The American Museum of Natural History,
together with a number of male paratypes. There are also several male paratypes
in the Holland Collection in the Carnegie Museum. Type locality, Medje.
The American Museum Congo Expedition took eleven males and
the single female above described, one male having been captured at
Gamangui in June, the female, which is the allotype, at Medje in
June, and all the other specimens at the latter place in July and
August. We also have two males taken by Mr. A. I. Good at Lolodorf,
Cameroon, in November 1914,
(272) 10. Cymothoé staudingeri Aurivillius
Cymothoé staudingerit AURIVILLIUS, 1898, Rhop. A2thiop., p. 212, Pl. rv, fig. 5, o.
Two somewhat defective males which I refer without doubt to
this species, in spite of the fact that the light mesial band tends to
coalesce with the light sagittate submarginal markings about the middle
of the fore wing, the dark lunate lines which define these inwardly in the
present specimens being fainter than in the figure given by the author
of the species. In all other respects the specimens agree perfectly, both
on the upper and lower sides, with the descriptions and figures of Auri-
villius. The specimens were taken at Medje, one in July, and the other
in September.
198 Bulletin American Museum of Natural History (Vol. XLIIT
(273) 11. Cymothoé jodutta (Westwood)
Harma jodutta Westwoop, 1850, Gen. Diurn. Lep., p. 289.
Harma cyriades Warp, 1871, Ent. Mo. Mag., VIII, p. 120.
Cymothoé aralus MaBrL_F, 1890, Ann. Soc. Ent. France, (6) X, p. 22, Pl. 1, fig. 8.
Cymothoé jodutta AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 152, Pl. xxxve, 7,
Four males and one female taken at Medje in August and Sep-
tember, and one female caught at Niangara in November.
(274) 12. Cymothoé ehmckei Dewitz
Cymothoé ehmckei DewitTz, 1886, Berl. Ent. Zeit., XXX, p. 302, Pl. vu, figs. 3, 4.
AvRIVILLIs, 1912, Seitz, Gross-Schmett., XIII, p. 152, Pl. xxxvd, 0’, 9.
Of this form, which is no doubt merely a local race of C. jodutta,
there are twenty-two males and six females. All were captured at Medje
from May to September, except one male which was taken at Niangara
in November.
(275) 13.. Cymothoé capellides, new species
Plate VIII: Figure 6, C. capella Ward, &; Figure 5, C. capellides Holland, #7
Allied to C. capella Ward, but smaller in size, and easily discriminated from the
latter species by the paler gray of the basal areas of both wings on the upper side and
the outward extension of this darker area on both wings as well as by the presence
of the characteristic dark markings of the gerius in the cell and beyond it on the
upper side of the fore wings, these dark markings being suppressed in C. capella,
the end of the cell and the apical third of the fore wing in Ward’s species being im-
maculate, except for the marginal series of spots. Expanse, o", 50-55 mm.
There are four males in the collection, all taken at Medje, one in May, the others
in August. They show no variation among themselves. The type is in The American
Museum of Natural History. Paratypes are in the Holland Collection in the Carnegie
Museum.
To make the distinction between the two species plain to the student
I give a figure of a typical male specimen of C. capella Ward and of the
type of the new species.
(276) 14, Cymothoé cenis (Drury)
Plate IX: Figure 3, C. conformis Aurivillius, 9; Figure 7, C. rubida Holland, 9
Papilio cenis Drury, 1773, Ill. Exot. Ent., I, p. 33, Pl. xrx, figs. 1, 2.
Cymothoé canis Aurivitiius, 1912, Seitz, Gross-Schmett., XIII, p. 151, Pl. xxxve,
re
There are fifty-six males and three females in the collection. Two
of the females belong to the form named conformis by Aurivillius and
one to the reddish form which I described many years ago in Psyche,
VI, p. 215, without giving it a name. C. canis was bred for me in
large numbers at Kangvé in the valley of the Ogové River and later at
1920] Holland, Lepidoptera of the Congo 199
Efulen in Cameroon by Dr. A. C. Good. The female is polymorphic, and
at least four well-defined varieties in this sex are known. The commonest
is that named althea by Cramer. The prevalent color of this variety is
black, with the wings crossed by a white mesial band. Closely allied
to it is the form called euthalioides by Kirby, in which the white mesial
band is broader and more irregular than in althea. Our plate gives
accurate representations of the other two varieties to which reference
has been made and a verbal description is not called for. I take pleasure
in designating the single specimen obtained by the American Museum
Congo Expedition at Medje as the type of C. cenis form rubida, new
form, 2, but have selected a more perfect specimen from my own col-
lection taken at Kangvé for representation on the plate. I have many
ex larva.
The specimens of C. cenis brought back by the Expedition were
almost all taken at Medje from June to September 1910, but there are a
couple ticketted as taken at Gamangui in June, one is labelled as from
Munie Katoto, September 1909, a few from Avakubi caught in October
of that same year, and several from Niangara captured in November
1910.
(277) 15. Cymothoé adelina (Hewitson)
Plate VIII: Figure 9, o; Figure 10, ?
Harma adelina Hewitson, 1869, Exot. Butt., IV, Harma, Pl. 1m, figs. 9, 11, 9.
(Non Cymothoé adelina, #) AuRIVILLIUS, 1912, Seitz, Gross-Schmett., XIIT, p. 152,
Pl. xxxvd. ;
Aurivillius in his ‘Lepidoptera Asthiopica,’ p. 215, makes C’. (Harma)
altisidora (Hewitson) a synonym of C. adelina, regarding the former as
being the male of the latter, which has priority. In Seitz, Gross-Schmet-
terlinge, XIII, Pl. xxxvd, he depicts a male of the following species
as that sex of adelina, and also gives a figure of a female which certainly
does not conform to Hewitson’s type of C. adelina. However, this species
is wonderfully variable in the female sex, as I have pointed out. I am
convinced that Dr. Aurivillius isin error. I give on Plate VIII, fig. 9,
a representation of the true male of C. adelina (Hewitson). Itisan insect
closely allied to C. cenis, from which it consistently differs in always
having the ground-color of the wings on the upper side deep Naples
yellow and not white or creamy white, as is the case with C. cenis.
With some hundreds of C. ceenis before me as I write, and a good series
of C. adelina both males and females, the difference is plainly visible
and strikingly constant. The females of C. adelina are exceedingly
variable in the ground-color or tint of the wings, although the black spots
°
200 Bulletin American Museum of Natural History [Vol. XLIII
and bands are quite uniform in all specimens I have seen; and I have
examined many scores of this sex. I do not possess a female as light in
color as the one figured by Aurivillius, but some which come very near
to it. Most specimens are some shade of orange-red, from that passing
into dark sienna and even deep umber. Two bred specimens received
some years ago from Dr. A. C. Good are almost black, the maculation
being almost lost on the darkly colored ground of the wings. With
these dark female specimens there emerged some females which are
much lighter.
The specimens belonging to the collection upon which I am report-
ing were taken at Medje from June to September, except one pair, which
was taken at Niangara in November.
(278) 16. Cymothoé angulifascia Aurivillius
: Plate X, Figure 9, @#
Cymothoé angulifascia Aurtviiiies, 1897, Ofvers. Sv. Vet.-Akad. Forh., LIV, 5, p.
285, fig. 3, 2; 1912, Seitz, Gross-Schmett., XIII, p. 153, P]. xxxvib, Q.
This species, of which hitherto only the female has been recognized,
is represented in the collection before me by eight males and twofemales.
I also have a fine pair collected for me more than twenty years ago by
Dr. A. C. Good at Kangvé on the Ogové River. The females agree per-
fectly with the description and figures given by Dr. Aurivillius. The
males, by the markings on the under side of the wings, disclose their
specific identity with the females. Aurivillius has provisionally placed
C. angulifascia in the same group with C. sangaris (Godart) and itsallies.
Now that we know the male sex of the species, it seems to me better
to put it into what Aurivillius terms the ‘Canis Group.”’
I do not think it necessary to give a detailed verbal description of
the male, as the excellent figure on the plate will enable any one to
recognize the insect. It is, in fact, so far as the wings show, almost
exactly like the male C. adelina in Seitz, XIII, Pl. xxxvd, but the
markings on the under side agree with those of C. angultfascia
Aurivillius and are reddish throughout.
The specimens were all captured at Medje, three of the males in
April, the rest from July to September. One of the females was caught in
August, the other in September.
(279) 17. Cymothoé sangaris (Godart)
Nymphalis sangaris Govar', 1823, Enc. Méth., LX, p. 384, co”.
Harma uselda Hewrrson, 1869, Exot. Butt., [1V, Harma, Pl. mm, figs. 18, 14, 9.
Two male specimens, one taken at Munie Katoto, September 10,
1909, the other at Medje, June 10, 1910.
1920] Holland, Lepidoptera of the Congo 201 -
(280) 18. Cymothoé aramis (Hewitson)
Plate X: Figure 1, #; Figure 2, 9
Euryphene aramis Hewrtson, 1865, Exot. Butt., III, Zuryphene, Pl. 1, figs. 16,17, 2.
Adolias fuluomacula CapronniER, 1889, Ann. Soc. Ent. Belgique, XX XIII, Bull., p.
exliv, 9.
Cymothoé aramis AuRIVILLIUs, 1912, Seitz, Gross-Schmett., XIII, p. 153, Pl. xxxvie,
ie
This species has hitherto only been known by the female sex, except
by myself. For many years a series of males has been standing in my
cabinets awaiting the coming of a favorable opportunity to describe
them, which now fortunately arrives. Instead, however, of giving a long
verbal description, I shall rely more upon the figure on the plate to aid
the student in the identification of the species. The male is in outline
and in the shape and location of the markings very much like C. anatorgis
(Hewitson) (see Plate X, fig. 7, @; fig. 8, 2; specimens in Holland
Coll.) but may be discriminated from that species at a glance by
the quite different color of the upper side of the wings. In C. anatorgis
the ground-color of the wings is deep blood-red, as in C. sangaris;
in aramis the ground-color is dark ochraceous, the tint being very much
the same as the ground-color of the wings in C. lurida or C. cyclades.
The American Museum Congo Expedition brought back two males,
which were taken at Medje, September 27, 1910. I havea series of males
and females taken at Kangvé on the Ogové River more than twenty years
ago.
(281) 19. Cymothoé coccinata (Hewitson)
Harma coccinata Hewitson, 1874, Exot. Butt., V, Harma, Pl. v1, figs. 24-26, #.
Cymothoé coccinata AuRIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, Pl. xxxvic, 3;
xxxvid, 9.
Of this species the Expedition returned seventy-four males, all
captured at Medje, except two which are labelled as taken at Gamangui
in June. A few were taken in April, the remainder were caught from
June to September.
(282) 20. Cymothoé Regine-Elizabethe, new species
Plate X: Figure 5, o, type; Figure 6, 9, ailotype
o. The males are without the light spot about the middle of the costa of the
secondaries on the upper side, which is a marked feature in C. aramis, anatorgis, and
coccinata. The edge of the costa towards the base is simply narrowly laved with pale
yellowish, which cannot be seen except by parting the fore and hind wings. The
prevalent color of the upper side of the wings is warm vermilion, somewhat paler than
in the case of C. coccinata, and much lighter in tone than in C. sangaris and C. ogova.
202 Bulletin American Museum of Natural History [Vol. XLIII
The wings are crenulate, slightly concave about the middle of the margin of the
primaries and truncated at their lower angle thus resembling in outline the wings of
C. sangaris, which is a much larger species. The hind wings are somewhat produced
at the anal angle, but not more so than is the case in C. aramis and C. anatorgis.
Both wings are defined outwardly by very fine black lines, except on their inner
margins; both have a submarginal series of quite small black dots on the inter-
spaces, those of the foie wing tending in a number of specimens before me to become
obsolete, those on the hind wings, which are larger and more conspicuous than those
of the fore wings, in some specimens tending to become arcuate or sagittate toward
the anal angle.
Q. The female in general appearance is not unlike the female of C. ogova
(Ploetz), but the white transverse band on the upper side of the secondaries lies much
nearer the base than in that species, and the dark basal area, which succeeds it in-
wardly is correspondingly reduced in extent. The differences between the two sexes
as shown by the upper side of the wings is clearly revealed in the figures given upon
Plate X.
Expanse, o°,52mm.; 9,65 mm.
There are thirteen males and two females of this beautiful species in
the collection. They were all taken at Medje (type locality) in July and
August, except one male which is labelled as caught at Niangara in
November. ;
On the occasion of the visit to the Carnegie Institute by their
Majesties, the King and the Queen of the Belgians, and of His Royal
Highness, the Crown Prince Leopold, Duke of Brabant, on October 23,
1919, I had the honor of showing to Her Majesty, the Queen, a proof of
Plate X of this paper, which was lying upon my desk. I requested Her
Majesty to accord to me the privilege of naming this lovely butterfly
in her honor, and she most graciously acceded to my request, expressing
pleasure at the thought. It therefore bears the name of the Queen of the
Belgians.
The types are in The American Museum of Natural History;
paratypes in the Holland Collection.
(283) 21. Cymothoé ogova (Plcetz)
Plate X: Figure 3, o"; Figure 4, 9
Harma ogova Pi@1z, 1880, Stett. Ent. Zeit., XLI, p. 1938, 9.
Cymothoé ogova Aurtvi.i10s, 1912, Seitz, Gross-Schmett., XIII, p. 154, Pl. xxxvib, 9.
The male of this species has never hitherto been described or figured.
I possess a series of eighteen males and twenty-two females bred for me
more than twenty years ago by Dr. A. C. Good at Kangvé on the Ogové
River. On Plate X, Fig. 3, I give a figure of the upper side of the wings of
the male and in Fig. 4 a representation of the wings of the female. The
male has a small shining white spot, girdled with black, at the very base
1920] - Holland, Lepidoptera of the Congo 203
of the hind wing, where it joins the body. In the general color of the
wings it closely approaches C. sangaris, from which, however, it may at
once be discriminated by the different form of the outline of the primaries,
which are not as much excavated on the margin and truncated at the
lower angle as is the case in C. sangaris. In the fine suite of females in
my possession I detect some variation, and one female has the apical
third of the wing not white, but reddish. For this slight variety the
name rubescens may be proposed, if such things deserve to be discrimi-
nated nomenclatorially. Expanse, 3; 52-60 mm.; @2, 60-75 mm.
Messrs. Lang and Chapin did not take any specimens of this
species, but I atone for the lack by donating to The American Museum
of Natural History a pair taken from my cabinet. I have no doubt that
it occurs on the Upper Congo, as well as in the valley of the Ogové.
EvuPTEeRA Staudinger
(284) 1. Euptera pluto (Ward)
Euryphene pluto Warp, 1873, Ent. Mo. Mag., X, p. 59.
One male taken at Medje the first week in August.
PsEUDATHYMA Staudinger
* (285) 1. Pseudathyma sibyllina (Staudinger)
Pseudacrea sibyllina StaupINGER, 1890, Iris, III, p. 338, Pl. 1m, fig. 8.
Pseudathyma sibyllina StaupinGER, 1891, Iris, IV, p. 90. Aurivillius,, 1912, Seitz,
Grosz:-Schmett., XIII, p. 156, Pl. xirxe.
One male taken at Medje the first week in August 1910.
The specimen differs slightly from the descriptions and figures of
this species which have been given by both Staudinger and Aurivillius.
It would, however, be a mistake to attempt to describe it as a different
species without more material at hand. There is before me a good speci-
men of Pseudathyma neptidina from the Ogové Valley, which the speci-
men somewhat closely resembles on the under side, but the absence
on both sides of the long white bar in the cell of the fore wing, which is
one of the diacritical marks of P. neptidina, forces me to assign it to P.
sibyllina, with which it agrees better than with any other species which
has been described. The specimens representing this genus in the col-
lections of the world are thus far very few, and, beside the individual I
am reporting upon and a few in my own collection, there are no others in
America, and only half a dozen in the museums of Europe. No doubt
the insect is common enough, but its mimetic resemblance to other forms
which are so common as to make them almost unworthy of notice by
collectors, has led to its having been overlooked.
204 Bulletin American Museum of Natural History [Vol. XLIII
EvuxanTHE Hiibner
(286) 1. Euxanthe trajanus (Ward)
Godartia trajanus Warp, 1871, Ent. Mo. Mag., VIII, p. 36.
Euzxanthe trajanus AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 123, Pl. xxrxf.
One male, Medje, September 1910.
(287) 2. Euxanthe ansellica (Butler)
Godartia ansellica ButiEerR, 1870, Trans. Ent. Soc. London, p. 525.
Euxanthe ansellica AuRIVILLUS, 1911, Seitz, Gross-Schmett., XIII, p. 124, Pl. xxrxf,
by error on plate there given as eurinome.
Four males, one taken at Bafwabaka, December 3: the others
captured at Medje, one in each of the months, May, peer: and Sep-
tember.
(288) 3. Euxanthe crossleyi (Ward)
Godartia crossleyi Warp, 1871, Ent. Mo. Mag., VIII, p. 36.
Euzxanthe crossleyi Aurtvitutus, 1911, Seitz, Gross-Schmett., XIII, p. 124.
Three males caught at Medje, July 1910.
CuHaraxes Ochsenheimer
(289) 1. Charaxes epijasius Reiche
Charazes epijasius Reicun, 1849, in Ferret and Galinier, Voyage en Abyssinie,
Entomologie, p. 469, Pl. xxxn, figs. 1, 2. Aurivintirus, 1911, Seitz, Gross-
Schmett., XIII, p. 126, Pl. xxxa.
The species is represented by two males, one taken at Faradje, and
simply ticketted ‘‘1911-1912,”’ the other labelled ‘‘Niangara, Nov-
ember 20-25, 1910.” The latter is the more perfect specimen.
This is the first record of this species from the Belgian Congo and
extends the known range. It has been recorded from Senegal, Nigeria,
and the Togo country and eastward to Abyssinia and Unyoro.
(290) 2. Charaxes brutus angustus Rothschild and Jordan
Papilio brutus Cramer, 1779, Pap. Exot., III, p. 82, Pl. coxut, figs. £, F.
Charazxes brutus angustus ROTHSCHILD AND JORDAN, 1900, Nov. Zool., VII, p. 432.
Charaxes angustus AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 126, Pl. xxxa.
Without exception the specimens before me belong to the form to
which Rothschild and Jordan have given the above name. This is also
true of all the specimens which I have received from Cameroon and the
valley of the Ogové River. It is the prevailing form in the hot, wooded
lands of the central Ethiopian region, and is easily distinguished from
typical C. brutus from Sierra Leone and adjacent parts.
1920] Holland, Lepidoptera of the Congo 205
The collection contains ninety-six males, of which twenty-one are
not as yet expanded. With the exception of one example taken at Baf-
waboli, September 11, and another taken at Bafwasende, September
27, 1909, all of the specimens were captured at Medje, a few in the first
week in April, the rest from early in July to about the middle of Sep-
tember.
(291) 3. Charaxes castor (Cramer)
Papilio castor CraMER, 1775, Pap. Exot., I, p. 61, Pl. xxxvu, figs. C, D.
Charaxes castor AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 127, Pl. xxxa.
There are ten males of this well-known insect belonging to the
characteristic West African form, which has the basal spots of the under
side black instead of chestnut. Professor Aurivillius has proposed the
subspecific name C. godarti for this form. One specimen was captured
at Niangara on November 8, 1910. |The other examples were all taken
at Medje, two in the month of May, the rest in July and August.
(292) 4. Charaxes pollux (Cramer)
Papilio pollux Cramer, 1775, Pap. Exot., I, p. 61, Pl. xxxvm, figs. Z, F.
Charazxes pollux AuRIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 127, Pl. xxxb.
One male taken at Bafwasende, October 23, 1909.
(293) 5. Charaxes eudoxus (Drury)
Papilio eudoxus Drury, 1782, Il. Exot. Ent., III, p. 44, Pl. xxx, figs. 1, 4.
Charazes eudoxus AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 128.
There are three males which I refer to this species. They were taken
at Medje, one in May, one in July, and another in August. They are
absolutely alike and may be distinguished at once from the following
species by the great reduction in width of the silvery bands on the lower
side of the hind wings, as well as by the prolongation costad of the median
fulvous band of the fore wings on the upper side.
The species is rare in collections.
(294) S “Charaxes sischow! Rothaehild gad Joian
Charaxes eudoxus mechowi ROTHSCHILD AND JORDAN, 1900, Nov. Zool., VII, p. 419;
1899, loc. cit., VI, Pl. vim, fig. 3.
Charaxes mechowi AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 128, Pl. xxxe,
name eudorus on plate an error.
There are seven males of this species, agreeing throughout with
others which we have from Cameroon. The figure given by Aurivillius
(ef. Seitz, ‘Die Gross-Schmetterlinge des Afrikanischen Faunengebietes,’
Pl. xxx) does not agree with the specimens before me in not having the
206 Bulletin American Museum of Natural History [Vol. XLIII
dark markings of the outer angle of the fore wings accentuated, as they
are in these specimens, by a short dark submarginal bar extending from
vein 1 and vein 3, and defining the median band outwardly.
Like the preceding form this is a rare species, the female of which
has never, I believe, been found or described.
(295) 7. Charaxes etesipe (Godart)
Nymphalis etesipe Gopart, 1823,"Enc. Méth., LX, p. 355.
Charazes etesipe AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 128, Pl. xxxe.
This seems to be a very common species, judging from the number
of specimens taken. Curiously, all of them are males. There are one
hundred and thirty-three examples, all taken at Medje, a.number in the
early part of April, the rest from the beginning of July to the early part
of September.
(296) 8. Charaxes protoclea Feisthamel
Charazes protoclea Fe1sTHAMEL, 1850, Ann. Soc. Ent. France, (2) VIII, p. 260.
Charazes eson HerricH-ScuFFeErR, 1850, Aussereur. Schmett., figs. 9,10, 9.
Charazes protoclea Aurtvitulus, 1911, Seitz, Gross-Schmett., XIII, p. 129, Pl.
XXXIlIla.
The collection includes one hundred and twelve males and four
females. One male is labelled “‘Niangara, November 8-9, 1910,”
another is from Faradje, captured in 1911. All the rest were taken at
Medje, a number early in April, the remainder from the beginning of
July to the middle of September.
(297) 8a. Charaxes protoclea marginepunctata, new variety
A number of specimens of this species have a well-developed series
of submarginal black spots located on the cadmium-orange border of
the hind wings. The vast majority of specimens of C. protoclea have only
one such spot on the interspace between veins 7 and8. For this form with
the submarginal spots I propose the above varietal name. The type
is in the Holland Collection from Gaboon, paratypes are in the
present collection, belonging to The American Museum of Natural
History, and in the Holland Collection from various localities in tropical
Africa.
(298) 9, Charaxes cynthia Butler
Charaxes cynthia Burier, 1865, Proc. Zool. Soc. London, p. 626, Pl. xxxvu, fig. 3.
Aunivitirivs, 1911, Seitz, Gross-Schmett., XIII, p. 130, Pl. xxxt1a,
Of this species there are in the collection seven unexpanded and
thirty-four expanded males. They were collected at Medje, a few in
April, the rest from the end of June to the beginning of September.
ee ae i...
1920] Holland, Lepidoptera of the Congo 207
(299) 10. Charaxes lucretius (Cramer)
Papilio lucretius CRAMER, 1777, Pap. Exot., I, p. 129, Pl. txxxu, figs. EZ, F.
Charaxes lucretius AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 130, Pl. xxxd.
Of this well-known and common species there are before me in the
collection forty-two males. One is labelled ‘‘Ngayu, December 14,
1909,” another “ Niangara, November 20-25, 1910,” and a third “ Baf-
wasende, September 23, 1909.”’ All the others were taken at Medje, a
couple in April, the rest from early in June to the middle of September.
(300) 11. Charaxes smaragdalis Butler
Charaxes smaragdalis ButTLeR, 1865, Proc. Zool. Soc. London, p. 630, Pl. xxxv1, fig. 5.
AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 131.
There are twenty-two males in the collection, one labelled “‘Munie
Katoto, September 10, 1909,” another “ Niangara, November 20-25,
1910.” The rest were taken at Medje, two in April, the remainder from
July to the middle of September, most of them, however, in August.
(301) 12. Charaxes numenes (Hewitson)
Nymphalis numenes Hewirson, 1859, Exot. Butt., Il, Nymphalis, Pl. 11, figs. 9, 10, 11.
Charaxes numenes AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 132, Pl. xxx1c.
The collection includes ninety-eight males and one female, all of
which were taken at Medje, a few in April, the rest from July to Sep-
tember 1910.
(302) 13. Charaxes tiridates (Cramer)
Papilio tiridates CRAMER, 1777, Pap. Exot., I, p. 100, Pl. cux1, figs. A, B.
Charazes tiridates AuRIviILLIvs, 1911, Seitz, Gross-Schmett., XIII, p. 132, Pl. xxx1b.
There are one hundred and seventy-six males and one female of this
species in the collection. They were all taken at Medje, quite a large
number in April, the rest in July and August, except a few captured in
the early part of September.
(303) _ 14. Charaxes bipunctatus Rothschild
Charaxes bipunctatus RoruscuiLp, 1894, Nov. Zool., I, p. 536. Auriviuxtus, 1911,
Seitz, Gross-Schmeit., XIII, p. 132.
Seven males, one not expanded. All were taken at Medje, six in
August, and one in September.
(304) 15. Charaxes imperialis Butler
Charaxes imperialis Butter, 1874, Trans. Ent. Soc. London, p. 531, Pl. x1, fig. 3.
AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 132.
Ten males, all taken at Medie, one in April, one in July, and four in
each of the months of August and September.
208 Bulletin American Museum of Natural History [Vol. XLII
(305) 16. Charaxes amelie Doumet
Charaxes amelie DouMET, 1861, Rev. Zool., (2) XIII, p. 171, Pl. v, fig. 1. Aurtvriuivs,
1911, Seitz, Gross-Schmett., XIII, p. 133, Pl. xxx1b.
There are one hundred and ninety-two specimens of this species in
the collection, forty-one of which have not been expanded. They are
all males. They were taken at Medje, a number early in April, the
rest from July to the middle of September.
(306) 17. Charaxes hadrianus Ward
Charaxes hadrianus Warp, 1870, Ent. Mo.Mag., VIII, p. 120. AURIVILLIUS, 1911,
Seitz, Gross-Schmett., XIII, p. 133, P]. xxxtd.
Represented by twenty-seven males, six not expanded, all taken at
Medje. One was caught in April, the rest from July to September, but
the greater number were taken in August.
(307) 18. Charaxes nobilis Druce
Charazes nobilis Druce, 1873, Ent. Mo. Mag., X, p. 13. Aurtviniius, 1911, Seitz,
Gross-Schmett., XIII, p. 133, Pl. xxxutc.
A single male specimen of this rare species was taken at Medje on
May 29, 1910. It lacks one antenna and is slightly rubbed.
(308) 19. Charaxes anticlea (Drury)
Papilio anticlea Drury, 1782, Ill. Exot. Ent., II, p. 36, Pl. xxvu, figs. 5, 6.
Charazes anticlea AURIVILLIUS, 1911, Seitz, Gross-Schmett., XIII, p. 134, Pl. xxxmid. _
Six males captured at Medje, the dates varying from July to Sep-
tember.
(309) 20. Charaxes hildebrandti (Dewitz)
Nymphalis hildebrandtii Dewrrz, 1879, Nov. Act. Acad. Nat Cur., XLI, p. 28, Pl. 0,
fig. 16.
Charaxes hildebrandti Auriviturus, 1911, Seitz, Gross-Schmett.., XIII p. 134, Pl.
XXXIIICc,
Nine males taken at Medje from July to September.
(310) 21. Charaxes etheocles (Cramer)
Papilio etheocles Cramer, 1777, Pap. Exot., II, p. 34, Pl. exrx, figs. D, 2.
Charazes etheocles AvniviLiius, 1911, Seitz, Gross-Schmett., XIII, p. 135, Pl. xxxmb,
Of this polymorphic and decidedly puzzling species there is a large
series of males and a solitary female. The female, which was taken at
Medje, June 28, 1910, does not agree absolutely with any form of the
female figured by authors, but comes nearest to that given by Staud-
as
1920] Holland, Lepidoptera of the Congo 209
inger. (Iris, 1896, TX, Pl. 11, fig. 4), which Rothschild and Jordan in their
revision of the genus accept as the typical female of C. etheocles (cf.
Nov. Zool., 1900, VII, p. 486).
The males, of which there are one hundred and twenty-one speci-
mens, belong principally to the form described by Butler as Charazxes
holland: (ef. Ann. Mag. Nat. Hist., 1893, (6) XII, p. 266), and the form
described by Staudinger under the name Charazes catochrous (cf. Iris,
IX, 1896, p. 216), in which the basal two-thirds of the wings are
whitish on the under side. There is one male taken at Niangara in
September which fits the description given by Rothschild and Jordan
of the form to which they have applied the name picta. There are some
intergrading forms which are not exactly referable to any of those just
mentioned. As is well known to students, the species with which we
are dealing is excessively variable and it is not wise to attempt without
the test of breeding to decide what are the exact relationships of the
variant forms which every new collection from Africa reveals.
The specimens, with the exception of the example of C. etheocles
picta taken at Niangara, as mentioned above, and a male of C. etheocles
hollandi taken at Faradje, were all collected at Medje or near by.
The dates of capture are early in April in the case of a few specimens, but
the majority are labelled as having been taken from July to September,
but principally in August.
(311) 22. Charaxes candiope (Godart)
Nymphalis candiope Gopart, 1813, Enc. Méth., UX, p. 353.
Charazes candiope AurRIvILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 138, Pl
XXxIla,
Of this species there are in the collection ninety-two males. With
the exception of two taken at Niangara in November and one captured
at Gamangui in June, they were all collected at Medje, a few early in
April, the rest from July to September, principally from the middle of
July to the end of August.
(312) 23. Charaxes kahldeni Homeyer and Dewitz
Charaxes kahldeni HomEYER AND Dewr1z, 1882, Berl. Ent. Zeit., XX VI, p. 381,
Pl. vu, figs. 1, 2. Aurtvitirus, 1912, Seitz, Gross-Schmett., XIII, p. 139.
Of this pretty little species there are thirty-eight males, all taken at
Medje from June to early September 1910.
(3138) 24. Charaxes eupale (Drury)
Papilio eupale Drury, 1782, Ill. Exot. Ent., III, p. 7, Pl. v1, fig. 3.
Charazes eupale AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 140, Pl. xxxtte
210 Bulletin American Museum of Natural Hictory (Vol. XLII
One hundred and sixteen males, two taken at Niangara in November,
one at Gamangui in June, and the rest at Medje, a few early in April,
the rest from June to September, but principally in July and August.
Of the “washed out”’ form named C. dilutus by Rothschild and Jordan
there are several examples. It is hardly worthy of a subspecific name,
being connected with the more strongly marked form by intergrades.
(314) 25. Charaxes nichetes Grose-Smith
Charazes nichetes GRosr-Smitu, 1883, Ent. Mo. Mag., XX, p. 58.
Charazes ogovensis HoLLAND, 1886, Trans. Amer. Ent. Soc., XIII, p. 330, Pl. vm,
fig. 2. Sr:
Charaxes nichetes AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 140, Pl. xxxi1b.
One male taken at Medje in September 1910.
(315) 26. Charaxes porthos Grose-Smith
Charazes porthos GrosE-SmiT#, 1883, Ent. Mo. Mag., XX, p. 57. Smr1n anp Kirsy,
1887, Rhop. Exot., I, Charazes, p. 2, Pl. 1, figs. 4, 5.
One male taken at Medje in the beginning of August.
(316) 27. Charaxes zelica Butler
Charazes zelica BUTLER, 1869, Ent. Mo. Mag., VI, p. 28. Aurivituius, 1912, Seitz,
Gross-Schmett., XIII, p. 141, P!. xxx1b.
One male captured at Medje early in August.
(317) 28. Charaxes laodice (Drury)
Papilio laodice Drury, 1782, Ill. Exot. Ent., III, p. 34, Pl. xxvu, figs. 1, 2.
Charazes laodice AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 141, Pl. xxxtic.
An expanded male taken at Medje, April 6, 1910, and another not
expanded.
(318) 29. Charaxes doubledayi Aurivillius
Charazes doubledayi Aurivitutus, 1899, Kongl. Sv. Vet.-Akad. Handl., XX XI, p. 244;
Seitz, Gross-Schmett., XIII, p. 141, Pl. xxx11e.
A series of twenty-one males taken at Medje, a few in April, the
rest from July to September.
(319) 30. Charaxes mycerina (Godart)
Nymphalis mycerina Govart, 1835, Ene, Méth., TX, p. 369.
Charaxes mycerina AURIVILLIUS, 1912, Seitz, Gross-Schmett,, XIII, p. 141,
One male taken at Medje in the first week of September 1910.
The separation of the form C. doubledayi Aurivillius from C,
mycerina (Godart) made by Prof. Aurivillius on the basis of the absence
of the marginal blue spots on the fore wing of the latter form appears
VO ee a
1920] Holland, Lepidoptera of the Congo 211
to the writer justifiable, but with long series of specimens before him
from all pafts of tropical west and central Africa, he finds that a regular
intergradation in this character occurs, and a regular series running
from spotless C’. mycerina to much-spotted C. doubledayi can be arranged.
Patta Hiibner
(320) 1. Palla vologeses Mabille
Palla vologeses MABILLE, 1876, Bull. Soc. Zool. France, I, p. 280.
Charazes vologeses AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 139, PI.
xxxuid,
A solitary male specimen taken at Niangara about the middle of
November.
(321) 2. Palla fulvescens Aurivillius
Palla fulvescens Auriviuuivs, 1891, Ent. Tidskr., XII, p. 216.
Charazes fulvescens AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 139, Pl. xxxud.
There are seventy-four specimens of this species before me, all of
them being males. With the exception of one specimen which is ticketed
“Niangara, XI, 20-23, 1910,” all were taken at Medje. A few were
captured in the month of April, and bear no later date than the 6th of
that month. One is stated to have been taken on June 11. All the rest
were taken at dates ranging from July 3 to September 6, 1910.
From the evidence of the labels it would appear that the species
is at least double-brooded, and there may be three broods during the
year.
(322) 3. Palla decius (Cramer)
Papilio decius CRAMER, 1777, Pap. Exot., II, p. 26, Pl. cxrv, figs. A, B.
Palla decius Auriviiiius, 1912, Seitz, Gross-Schmett., XIII, p. 141, Pl. xxxuid.
There are three males before me, two of which were taken at Medje
about the middle of July, and one at the same place near the end of
August. |
(323) 4. Palla ussheri Butler
Palla ussheri Butier, 1870, Trans. Ent. Soc. London, p. 124; 1871, Lep. Exot.,
p. 52, Pl. xx1, fig. 3. AuRivrLiius, 1912, Seitz, Gross-Schmett., XIII, p. 141,
Pl. xxxiid.
This species seems to be vastly more common at the places where
collections were made than the preceding. There are fifty-seven males.
With the exception of one specimen, labelled ‘Risimu, September 8,
1909,”’ all were taken at Medje, a few early in April, the rest from the
first week of July to the middle of September. |
212 Bulletin American Museum of Natural History [Vol. XLIII
PuHitocnoma Westwood
(324) 1. Philognoma lichas bebra (Rothschild)
Charazes lichas bebra Rotuscuixp, 1900, Nov. Zool., VII, p. 507.
Charazes bebra AuRIviLLIvs, 1912, Seitz, Gross-Schmett., XIII, p. 139.
The collection contains thirty-four males and one female referable
to this form. The males differ markedly from the typical form which
occurs on the Gold Coast, the dark area of the apical portion of the wing
being invaded internally by the lighter color of the body of the wing,
as pointed out by Rothschild. The female, however, does not differ at
all from females of the typical form in the collection of the writer from
Sierra Leone and Cameroon.
They were all collected at Medje, except one labelled as taken at
Gamangui on February 6, 1910. The dates of capture run through the
months of July and August, with the foregoing exception.
(325) 2. Pbilognoma paphianus (Ward)
Charazes paphianus Warp, 1871, Ent. Mo. Mag., VIII, p. 120.
Philognoma falcata Butter, 1872, Lep. Exot., p. 101, Pl. xxxvum1, fig. 1.
Charazes paphianus AURIVILLIUS, 1912, Seitz, Gross-Schmett., XIII, p. 139, Pl. xxxua.
There are in the collection sixty-five males, of which nine are not
expanded. With the exception of a single specimen labelled as taken at
Gamangui, June 6, 1907, all were captured at Medje, the dates ranging
from early in June to about the middle of September. The specimens
show very little variation.
Monvra Mabille
(326) 1. Monura zingha (Cramer)
Papilio zingha Cramer, 1780, Pap. Exot., IV, p. 53, Pl. ccexv, figs. B, C.
Charazes zingha Aurtvitutus, 1911, Seitz, Gross-Schmett., XIII, p. 128, Pl. xxxnd.
This species is represented by one hundred and thirty-one males
and three females. With the exception of one male which is labelled
as having been taken at Niangara on September 26, all of the specimens
were captured at Medje. A few were taken in the first week of April.
The greater number were captured in July and August, the earliest date
being June 30, and.the latest September 9, 1910.
Libytheide
LisytH#ea Fabricius
(327) 1. Libythea labdaca Westwood
Libythea labdaca Westwoon, 1851, Gen. Diurn. Lep., I, p. 413, note, Pl. yxvin, fig. 6.
Dichora labdaca Scuvpen, ef, infra.
1920] Holland, Lepidoptera of the Congo 213
Libythea labdaca Avriviuutus, 1913, Seitz, Gross-Schmett., XIII, Pl. xia.
Of this interesting insect, for which the late Dr. 8S. H. Scudder
proposed the generic name Dichora (cf. Report U.S. Geol. Survey, VIII,
part 1, p. 470) there are nineteen specimens: six taken at Medje, four
in June, one in August, and one in September 1910; three collected at
Basoko in July, 1909; three taken at Isangi and one at Stanleyville in
August 1909; four collected at Avakubi in September 1909; and two
badly worn examples taken at Lubila in the same month and year.
This butterfly in some years is quite scarce and only occasional
examples are taken; and again it appears in enormous numbers, fairly
swarming, as I have been informed by collectors resident in Cameroon
and on the Gold Coast.
Lemoniidez
Azpisara Felder
(328) 1. Abisara rogersi Druce
Abisara rogersi Druce, 1878, Ent. Mo. Mag., XV, p. 101.
Abisara geryon Dewrrz, 1889, Ent. Nachr., XV, p. 105, Pl. 1, figs. 3, 4.
A solitary male, captured at Niangara in November 1910.
(329) 2. Abisara intermedia Aurivillius
Abisara intermedia Aurtvitutus, 1895, Ent. Nachr., X XT, p. 381.
There are two males of this form, both taken at Medje, one in April,
the other in September. With these I associate three females, one taken
in each of the months April, July, and September at the same place.
They agree with the males on the under side of the wings.
(330) 3. Abisara rutherfordi Hewitson
Abisara rutherfordii Hnwrrson, 1874, Ent. Mo. Mag., XI, p. 56.
Abisara rutherfordi AuRIvILLIus, 1913, Seitz, Gross-Schmett., XIII, Pl. ux1b.
There are three males and two females of this species captured at
Medje, one female in June, all the others in August.
Lycenide
The genera and species of this family known to occur in the region
of the Congo are not only very numerous, but of the highest interest.
The Ethiopian Subregion is the metropolis of some of the most aber-
rant forms of this great family, which display to a wonderful degree the
phenomena of “‘mimicry.’”’ Many species are gorgeously beautiful and
bizarre in their coloring, and in their habits are no less wonderful, quite
a large number of them being in their larval state carnivorous, or ento-
mophagous, and many being myrmecophilous.
214 Bulletin American Museum of Natural History (Vol. XLIIT
The collection assembled by the American Museum Congo Expe-
dition is very limited both in the number of specimens and species, and
is by no means representative of this portion of the fauna. Nevertheless,
it possesses the highest interest, and it is to be regretted that the gentle-
men in charge had not the opportunity to devote more time to careful
collecting in the parts which they visited. Small as is the collection, it
contains a number of novelties, thus showing how much might have been
ascertained had the native collectors been induced to turn their attention
to this particular group. In proportion to the number of specimens brought
back, the number of species new to science is considerable. ‘There are
also some species which, until the present time, have been very inade-
quately represented in the collections of the world and which it has been
a pleasure to the writer to study and examine.
Lipteninse
Teurpna Aurivillius
(331) 1. Telipna rothioides, new species
Plate XII, Figure 7, @
9. Near 7. rothi Grose-Smith, but may be distinguished from that species by
the facts that on the upper side of the primaries the transverse reddish yellow band
does not reach the costa, as it does in 7’. rothi, that the black marginal border on the
upper side of the secondaries is broader than in 7’. rothi, and by the further fact that
on the under side of the secondaries the marginal row of white spots surrounded by
black terminates abruptly at vein 5, and does not completely encircle the wing as is
the case in 7. rothi. There are other minor differences, but those stated will enable the
student to discriminate this form from 7. rothi, to which, until I had made a critical
examination, I was inclined to refer it, and which it otherwise superficially resembles.
The collection contains two females, both captured at Medie, one
in April, the other in July, The latter is designated as the type and is
in The American Museum of Natural History in New York; the former,
which is the paratype, isin the Holland Collection in the Carnegie Museum
in Pittsburgh.
(332) 2. Telipna medjensis, new species
Plate XII, Figure 8, 9?
9. Near 7. nyanza Neave (cf. Novitates Zoologice, 1904, XI, p. 335, Pl. 1,
fig. 19). It differs, however, from the species described and figured by Neave in im-
portant particulars. The subapical white spot of the primaries is much larger than
in 7. nyanza, extending from vein 4 to vein 8, and reappears conspicuously on the
under side of the wing, which is not the case in Neave’s species. On the under side
this spot is bordered inwardly by a heavy black bar, running from the costa to vein 4,
and on the under side this spot is not defined outwardly by dark markings, and the
entire apical region beyond it, save immediately on the costa, is immaculate yellowish
‘
1920] Holland, Lepidoptera of the Congo 215
red, like the ground-color of the rest of the wing. The under side of the secondaries
also differs from 7. nyanza in that there are five, instead of three black bars on
the costa, as in the latter species, the third from the base running downward across
the end of the cell. There are also two minute black dots, one above the other, in the
cell beyond its middle, and in interval 5 there is a conspicuous squarish black spot.
Furthermore the black outer border of the secondaries on the lower side terminates
abruptly at the extremity of vein 5, as in the preceding species, the pale yellowish red
ground-color between veins 5 and 7 extending outwardly quite to the border of the
wing, separated from the cilia only by a very narrow black marginal line. The cilia
are broadly checkered with white between the extremities of the nervules where they
are black. Succeeding the fine black marginal line there is a row of sublunate white
spots sharply defined upon the deep black border, extending from the anal angle as
far as the interval between veins 3 and 4. At a remove of about two millimeters
from this row of spots basad there is another row of similar larger white spots,
each spot totally surrounded by the deep black of the border, which terminates in-
wardly just above them. Expanse, 50 mm.
The type, which is from Medje and is unique, is in the American
Museum of Natural History.
(333) 3. Telipna bimacula (Ploetz)
Pentila bimacula Pia@rz, 1880, Stett. Ent. Zeit., XLI, p. 199.
The species is represented by but one female specimen, which is in
The American Museum of Natural History. It was taken at Niangara
about the middle of November 1910, and does not differ in the least
from long suites of the species which we possess from the valley of the
Ogové and from Cameroon.
PENTILA Westwood
(334) 1. Pentila clarensis Neave
Plate XII, Figure 11 upper side”, from Faradje; Figure 12 upper side , from Basoko —
Pentila clarensis NEAveE, 1903, Ent. Mo. Mag., XX XIX, pp. 136-137.
There are two specimens taken at Faradje, “1911-1912,” which
agree so closely with the description of this species that I have no doubt
that they represent it. There are four other specimens, one taken at
Basoko in July 1909, two captured at Gamangui in June, and one taken
at, Medje in July 1910, which in the main agree with those taken at
Faradje and which cannot be separated from them specifically. These
four appear to me to be at most representatives of a seasonal or local
variety, the only difference being the increased size and therefore more
prominent appearance of the spots on the upper and lower sides of the
wings. In the specimens from Faradje the discal spots are much re-
duced in size, and some are almost obsolete, as Neave points out to have
been the case with some of the specimens before him when he wrote his
216 Bulletin American Museum of Natural History [Vol. XLIIT
description. The specimens from Gamangui, Basoko, and Medje
agree very closely with each other in every particular and have a facies
which is somewhat different from the specimens taken at Faradje.
When an analysis of the facts is made, however, it clearly appears that
the only real difference is due to the uniform enlargement of the various
spots upon the wings. As Neave points out, this insect is a close mimic
of Pardopsis punctatissima. This is particularly true of the specimens
with the enlarged spots. From P. pauli Staudinger the insect may be
discriminated by the fact that the elongated marginal spots, one in each
interval of the upper and lower wings, do not entirely reach to the margin,
as they do in P. pauli (ef. Rhop. Athiopica, p. 261. where Aurivillius
has given us a figure of that species), but constitute a distinct and regular
submarginal series, separated from the thin marginal line by an appreci-
able interval. Many of the spots are, as Neave points out, ‘‘monili-
form,’’—I should say having the shape of a dumb-bell, especially in the
apical region of the fore wing. The species is in.my judgment valid and
not to be confounded with P. pauli, which in size and general appearance
it somewhat resembles. I discover that we have a series of finely
preserved specimens of the species from the interior of Cameroon, collect-
ed some years ago by Mr. A. I. Good and awaiting a convenient season
for its study, which the arrival of the Lang-Chapin material has created.
(335) 2. Pentila cletensi Aurivillius (?)
Plate XII, Figure 9, @
Pentila cletensi AuRtvILLIvs, 1897, Ent. Tidskr., X VIII, p. 214, fig.
In the markings of the fore wing agreeing entirely with the descrip-
tion and figure of P. gletensi given by Aurivillius, but quite different in
the form and arrangement of the spots on the outer border of the hind
wing. In the specimens before me the marginal row of spots on the
secondaries are located at the tips of the veins and are quite large,
sagittate, pointing inwardly and running for some distance along the
nervules basad, much as in P. auga Karsch, or in some heavily marked
specimens of P. abraxas Doubleday and Hewitson. I refer the specimens
provisionally to P. cletensi Aurivillius, with which they agree more
closely than with any other species, realizing that there is in this genus
more or less variability in the number and size of some of the markings,
as is shown by long suites of many of the species in my possession. Itisa
doubtful procedure to erect species in the genus Pentila, basing them
upon the presence or absence of some of the spots, or their size.
The collection contains two males taken at Medje, one in August,
the other in September.
1920] Holland, Lepidoptera of the Congo 217
PsEUDERESIA Butler
(336) 1. Pseuderesia libentina (Hewitson)
Liptena libentina Hewirson, 1866, Exot. Butt., III, Pl. Liptena and Pentila, figs. 8, 9.
Pseuderesia libentina Aurivi.utus, 1898, Rhop. Aithiop., p. 267.
A somewhat poorly preserved female taken at Medje about the
middle of August is referable to this species. The type of P. (Liptena)
libentina (Hewitson) is lost, and some years ago, when I consulted the
collection of Hewitson in the British Museum, it had been replaced by
two specimens of other species. The specimen from Medje, however,
agrees so well with Hewitson’s figure and other specimens referred to this
species in the collections at Pittsburgh that there seems to be no doubt
of the correctness of the determination.
Epitoline
Epr1irota Westwood ’
(337) 1. Epitcla langi, new species
Plate XII, Figure 10, @#
&. The fore wing acuminate at apex, slightly concave below the apex and
a little rounded at the lower angle. Hing wing rounded externally, a trifle elongated.
Thorax and abdomen black above, pale brown below. On the upper side both wings
are dark brown, almost black, feebly revealing in certain lights a deep violet-blue
sheen, especially on the disk of the primaries and the inner half of the secondaries.
This color only reveals itself in a strong lateral light; in direct vision the wings
appear to be almost uniformly dark brown or black. The cilia are concolorous. On
the under side both wings are pale smoky brown, except on the inner margin of the
primaries near the angle, where they are pale bluish gray, a shade or two lighter
than the ground-color of the wings. About the end of the cell of the fore wing there
is a vertical row of four small pale spots extending from the costa to the lower angle
of the cell, and in the apical area of this wing there is a row of four or five similar
spots just before the outer margin extending from the apex to the interval between
veins 3 and 4, at the furthest. The hind wings on the lower side have a row of pale
basal spots, three in number, quite small and faint. These are followed by a series
of similar subbasal spots, which are also not conspicuous, but clearly defined. In
the middle of the cell is a quadrate darker spot defined before and behind by thin
pale lines, which I reckon as belonging to the subbasal series. Crossing the wing
from the costa to the inner margin is a curved and twice dislocated line of small pale
spots, two of which just at the end of the cell are the most conspicuous. This median
row of small light spots is succeeded by a postmedian row, quite regularly curved and
extending from the costa to the inner margin, a spot in each intraneural space. This
row is in turn succeeded just before the margin by a regular row of submarginal spots
somewhat crescentic in form. The cilia on the underside are dark as on the upper
side. Expanse, 29 mm.
The type, which lacks antenne and is in this respect defective, is unique. It
was taken at Medje, August 19, 1910 and belongs to The American Museum of
Natural History.
218 Bulletin American Museum of Natural History [Vol. X LIII
I dislike to found a new species upon a solitary defective specimen,
but, after examining every picture of an Epitola which has been published
and reading carefully every description which has been printed, I am
convinced that the little butterfly before me has never been described
or figured, at least not in such manner as to make either figure or de-
scription recognizable. The insect most nearly approaches EL. mangoénsis
Bethune-Baker (cf. Proc. Zool. Soc. London, 1908, Pl. vu, fig. 6). The
outline of the wings is the same, but the markings below differ. It is
odd that the only specimen of the genus Epitola brought back from the
Congo by the expedition should turn out to be hitherto nondescript.
There are now nearly fifty species of the genus known from the region of
which the Belgian-Congo forms a part.
Lycenine
MEGALOPALPUS Rober
The separation of the African species under the generic name Megalo-
palpus Rober from the Asiatic forms, which have been described under
the generic names Gerydus Boisduval and Paragerydus Distant is in the
opinion of the writer a rather unnecessary refinement, based upon struc-
tural differences which are so microscopic as hardly to be worthy of
regard. We are reaching a time when the discriminating instincts of
authors, who carry on their labors with the help of compound micro-
scopes, will demand the erection of a genus for every species, not only in
entomology, but all the other zoological sciences.
The so-called ‘species’ of Megalopalpus found in tropical Africa
seem to the writer to be in a state of confusion at the present time.
Aurivillius in his ‘Rhopalocera Awthiopica,’ p. 300, recognizes three species.
The first is M. zymna (Westwood), originally figured in Doubleday and
Hewitson’s ‘Genera of Diurnal Lepidoptera,’ who in their plate represent
a rather small insect, apparently belonging to the female sex, in which the
posterior border of the secondaries is broadly margined with black. The
second species recognized by Aurivillius is M. simplex Réber, in which
the hind wings are more narrowly bordered with black and in which the
markings of the under side of the wings are as described by Aurivillius
in his analytical key. This species Aurivillius regards as having been
redescribed by Capronnier under the name bicoloraria, and also re-
described by Kirby and refigured by Smith and Kirby under the specific
name similis. The third species admitted by Aurivillius is 7. metaleucus
Karsch, which, according to Professor Aurivillius, is the insect figured as
M. zymna by Smith and Kirby.
1920] Holland, Lepidoptera of the Congo 219
It happens that the present writer has before him a very large
series of specimens of the genus from various parts of Africa where it is
found. Over this mass of specimens he has long pored in the attempt to
bring order out of what seems chaos. Every one of the forms admitted
by Aurivillius to have specific rank is to be found in this series, but
there are so many intergrading forms that it is impossible to decide
where one species begins and the other ends. Of course, by selecting
those which tally with the forms already named and described and de-
stroying the rest, a semblance of specific security for these forms might
apparently be produced. But such a procedure would not be scientifi-
cally honest. In the first place, there is great variability in size among
specimens marked alike. Some are only 22 mm. in expanse, and they
range through various measurements up to 45 mm. in expanse. Some
have the outer margins of the secondaries heavily bordered with black;
in other specimens the hind margins are altogether free from black on
the margins; and there are intergrades between the two extremes.
Some are absolutely free from markings of any kind whatever on the
lower side of both primaries and secondaries; others are heavily marked,
as described and figured by authors; and there is every grade of
difference from those which are plain white on the under side to those
which are figured as is the insect named M. zymna by Smith and Kirby
and figured by them as such. Those with the light hind margins have
the same maculation on the under side as those which have the heavy
hind margins. Nothing is absolutely fixed, and there is no key to the
puzzle, if regard be had to the maculation and markings of the wings.
The fact is, I am convinced, that we are dealing here with a species
characterized by great variability, both in size and markings. We have
over one hundred specimens collected at one locality in the Cameroons
during one summer. All of the so-called species are represented in the
bunch, and the writer, if he were disposed to do so, might describe several
others from the same catch, if he were willing to select some of the inter-
grading forms and dwell upon the presence or absence of this or that
spot, or the slight variations in shade which occur. Differences are
plainly and clearly distinguishable, but to the mind of the writer they
do not show specific diversity. They represent merely individual
variation, and he is inclined to the belief that the whole congeries of
variant specimens are after all only referable to M. zymna (Westwood), a
protean species. There are vastly more females than males in the collec-
tion and, strangely enough, a multitude of the females are greatly
dwarfed, most of the specimens only measuring 22 mm. in expanse of
220 Bulletin American Museum of Natural History [Vol. XLII
wings, though some females, identically marked, are greatly hypertro-
phied. The insects are myrmecophilous, and perhaps some peculiarity
in their mode of nutrition in the larval state, the relative ease or diffi-
culty with which they secure their sustenance in this stage of their
existence, may account for the very abnormal difference in the size of
individuals. Until some observer carefully works out their life-history
by breeding we shall not have a solution of the problem with which the
present writer feels himself confronted.
(338) * 1. Megalopalpus zymna (Westwood)
Miletus zymna WeEstwoop, 1852, in Doubleday and Hewitson, Gen. Diurn. Lep., II,
p. 503, Pl. uxxv1, fig. 7.
(? M. simplex RéseEr, M. bicoloraria CAPRONNIER, M. similis Kirpy, M. metaleucus
Karscu, M. zymna Smita and Krrsy.)
The collection contains three specimens, all females, taken at Medje,
no two of which are exactly alike, though each of which can be matched
in any large collection such as that in the possession of the writer. They
are as follows.
a. Q. Dwarfed, expanse 24 mm., markings of the under side as
in M. simplex, but almost obsolete and so pale as only to be detected
by close scrutiny. Like a score of specimens before the writer from Lolo-
dorf, Cameroon. Taken at Medje, June 27, 1910.
b. Q. Expanse 43 mm. Combining on under side characteristics
of M. simplex, as defined by Aurivillius and shown in Rdéber’s photo-
graph, and characteristics of M. zymna, as depicted by Smith and Kirby
=M. metaleucus Karsch, fide Aurivillius. On the upper side of the
secondaries dark border reduced to a mere trace. Taken at Medje,
September 27, 1910. Can be matched by numerous specimens in collec-
tion of writer, some with black hind borders on upper side of hind wings,
some without such borders, some light on the under side, some dark,
some fulvous, some slaty gray, as in Smith and Kirby’s figure (‘Afr.
Lycenide,’ Plate x11, figs. 1, 2).
c. 9. Smaller than the preceding specimen. Expanse, 33 mm.,
with the hind margins of the secondaries much broader and darker than in
that specimen, but the markings of the under side much paler, though
closely resembling those of specimen 6. Taken at Medje, August 1910,
Lacunocnema Trimen
This is another myrmecophilous genus which stands in much need
of intelligent revision. The writer has a mass of material at his com-
mand, collected in tropical East Africa and in tropical West Africa, as
1920] Holland, Lepidoptera of the Congo 221
well as material received in the past from the late Dr. Roland Trimen,
collected in the Transvaal. The study of this shows that there are
either a number of closely related forms which have not been hitherto
discriminated or that the species are remarkably variable. This is not,
however, the place to take up the discussion of this subject.
(339) 1. Lachnocnema reutlingeri Holland
Lachnocnema reutlingeri Houuanp, 1893, Ann. Mag. Nat. Hist., (6) X, p. 286.
AURIVILLIUS, 1898, Rhop. Athiop., p. 302, °.
Arrugia umbra SmiTrH AND Kirpy, 1901, Rhop. Exot., III, Afr. Lycenide, p. 128,
Pl. xxvu, figs. 5, 6, @.
_ The American Museum Congo Expedition brought back two males,
neither very well preserved, which represent this species. We have in the
Carnegie Museum fifty-five males and ten females, including the type -
of the species. The two specimens taken at Medje in July and August
have been compared with these and they agree absolutely.
The writer has a suspicion that L. luna H. H. Druce (Proc. Zool.
Soc. London, 1910, p. 368, Pl. xxxrv, fig. 5) is only a slight varietal
form of this species, and that L. magna Aurivillius may only be an albinic
female of the same insect. In fact, he has a specimen of a female in
which the yellow color of the upper side is so pale that it might almost be
termed white and which seems rather closely to agree with the descrip-
ion given by Dr. Aurivillius.
Devporix Hewitson
(340) 1. Deudorix elealodes Bethune-Baker
Deudorix elealodes BETHUNE-BAKER, 1908, Proc. Zool. Soc. London, p. 112, Pl. rx,
fig. 6. :
There are two somewhat imperfect male specimens of this pretty
species taken at Medje, one in June, the other in August. As the author
points out, the species is near to D. eleala Hewitson, from which, how-
ever, upon comparison it may be distinguished at a glance by the
heavenly ultramarine blue of the upper surface, which strongly contrasts
with the greenish blue ground-color of its ally, by the almost complete
obsolescence of the lobe-spot on the under side of the hind wing, and by
the much reduced width of the transverse lines on the under side of
both wings.
(341) 2. Deudorix batikelides, new species
&. Closely resembling D. batikeli Boisduval on the upper side, but with dark
markings near the anal angle of the secondaries less prominent. On the under side
lacking altogether the three basal spots which are conspicuous upon the secondaries
222 Bulletin American Museum of Natural History [Vol. XLII
of Boisduval’s species, and with the wings throughout evenly pale warm gray, with
the tranverse lines and markings only a little lighter than the ground-color. These
markings recall in their disposition those of certain oriental Lycenide belonging to
the genus Lampides.
Q. Like the male, but with the blue of the discal areas of both wings reduced in
extent, due to the widening of the dark costal and outer marginal areas of black on
both wings. :
Type @ taken at Niangara, November 1910; allotype Q taken at Mombasa,
E. Africa by William Doherty, and in Holland Collection, Carnegie Museum.
I am under the impression that this is the form which has in the
past been referred by some authors to the species D. batzkeli, and re-
ported as such from various points on the East African mainland. D.
batikeli is found on the island of Madagascar.
I have for a number of years had a small but very fine series of
this insect collected for me on the hills about twelve miles up from
the coast back of Mombasa by the late William Doherty. They have
puzzled me, because, while resembling D. batikeli,
they could not be found to agree with either
figures or descriptions or actual specimens from
Madagascar and were labelled “‘? batikeli Bsd.”
The occurrence in the collection upon which
I am reporting of a good male specimen,
picked up at Niangara in November 1910,
brings matters to a focus, and I venture to’ Fig. 1. Deudorix batikelides
: ee : Holland. 9 (Nat. size). Drawn
give a name to this insect, which I am sure 4,01, specimen takenat Mombasa.
is not the same as the insect named by Bois-
duval and which apparently has been overlooked by other students.
(342) 3. Deudorix antalus (Hopffer)
Dipsas antalus Horrrer, 1855, Monatsber. Akad. Wiss. Berlin, p. 641.
Sithon antalus Horrrer, 1862, Peters, Reise n. Mossambique, Ins., p. 400, Pl. xxxv,
figs. 7-9.
Deudorix antalus Aurivitiivs, 1898, Rhop, Athiop., p., 309.
One female of this widely distributed species taken at Faradje,
January 11, 1912.
Oxyuipes Hiibner
(343) 1. Oxylides homeyeri (Dewitz)
Plate XII, Figure 5, 9°
Hypolycana homeyeri Dwwrrz, 1879, Nov, Act, Acad, Nat. Cur., XLI, part 2, p.
206, Pl. xxvi, fig. 13.
Oxrylides homeyeri Aunivitiius, 1898, Rhop, Athiop., p, 313.
1920] Holland, Lepidoptera of the Congo 223
There is one female specimen in moderately good condition, taken
at Medje, July 20, 1910. As this sex of the insect has never heretofore
been described, unless, as Aurivillius suggests, O. melanomitra Karsch
be it, I venture to briefly gives its characteristics.
Q. On the under side apparently agreeing completely with the description and
figure given by Dewitz (loc. cit.). On the upper side the ground-color is gray laved
with lilac, especially on the cells and discal areas of both wings. The costa and outer
margin of the front wings are darker, and on the ‘hind wings there is just below the
upper angle a dark spot or band of limited size, which is darker than the rest of the
wing. The lower end of the hind wing is broadly white on this side, with the three
marginal spots standing out conspicuously upon the light ground. On the under
side the anal angle and the adjacent parts of the wing are somewhat broadly laved
with yellow, which is also the case in the male.
Hypotyc#na Felder
(344) 1. Hypolycena hatita Hewitson
Hypolycena hatita Hewrrson, 1865, Ill. Diurn. Lep., Lyczenide, p. 51, Pl. xxx,
figs. 21-24. Aurivituius, 1898, Rhop. Athiop., p. 315.
There are eighteen males and one female of this insect, all in more or
less damaged condition, as is usually the case with them when received
from collectors. It appears to be almost impossible to preserve the long
narrow tails with which the hind wings of the species of this genus are
adorned. I have hundreds of them, and none seem to be absolutely
perfect. All of the specimens returned by the expedition were taken at.
Medje on dates ranging from March to September, except two, one of
which was caught at Risimu in September 1909. and the other at
Gamangui on June 14, 1910.
(345) 2. Hypolycena antifaunus (Doubleday and Hewitson)
Tolus antifaunus DouBLEDAY AND Hewitson, 1852, Gen. Diurn. Lep., II, p. 481,
Pl. txxv, fig. 1.
Hypolycena antifaunus AURIVILLIUS, 1898, Rhop. Aithiop., p. 315.
There are four damaged specimens of this species all captured at
Medje, one in April, the other three in July.
(346) 3. Hypolycena dubia Aurivillius
Hypolycena dubia AuRIviLuivs, 1895, Ent. Tidskr., XIV, p. 211.
This is indeed a very dubious species, as its name implies. It is
probably only a seasonal variety, or local race of H. lebona Hewitson,
characterized by being a somewhat brighter blue on the upper side of the
fore wings, and having a wider inward extension toward the cell of the
light color of the inner margin. It is a very common insect in Cameroon
and on the Ogové River, and I have hundreds of specimens, both males
224 Bulletin American Museum of Natural History (Vol. XLIII
and females, of H. lebona and H. dubia, and while the diacritical points
emphasized by Aurivillius hold well enough in many cases, so that it is
possible to make up a good series of both forms, there are many inter-
grading forms which suggest that we may well be in doubt as to the
validity of the species as such.
The collection contains a-male and female in fairly good preserva-
tion taken at Medje..
(347) 4. Hypolycena liara H. H. Druce
Hypolycena liara H. H. Druce, 1890, Ann. Mag. Nat. Hist., (6) V, p.27. AuriviL-
Lius, 1898, Rhop. A®thiop., p. 316.
A male and a female were captured, the former at Niangara in
November, the latter at Medje in June.
Apanzus Hiibner
(348) 1. Aphnezus orcas (Drury)
Thecla orcas Drury, 1782, Ill. Exot. Ent., ITI, p. 46, Pl. xxxrv, figs. 2, 3.
Aphneus orcas HOLLAND, 1893, Ent. News, IV, Pl. 1, fig. 13. Aurtvitiius, 1898,
Rhop. 2thiop., p. 327.
Six males, one taken at Lubila, September 1909, four at Medje from
April to September, and one at Niangara in November 1910.
Spinpasis Wallengren
(349) 1. Spindasis natalensis (Hewitson)
Aphneus natalensis Hewrtson, 1865, Ill. Diurn. Lep., Lycenide, p. 62, Pl. xxv,
figs. 1, 2.
Aphneus caffer Trimen, 1868, Trans. Ent. Soc. London, p. 88.
The specimens have been compared with others named Aphneus
caffer and obtained from Mr. Trimen himself in exchange. They agree
perfectly with these. Aurivillius sinks the name caffer as a synonym
of mozambica Bertolini, but I am unable to identify the reference given
by Aurivillius in the ‘Rhopalocera A®thiopica,’ though I have made
diligent search.
The expedition brought back five specimens, a male caught at
Medje in August, a pair taken at Niangara in November, and two males
captured at Faradje, one in December 1910, the other in “1911-1912.”
(350) 2. Spindasis crustaria (Holland)
Aphnaus crustaria HOLLAND, 1890, Psyche, V, p. 420; 1893, Ent. News, IV, p. 28, Pl.
1, fig. 10.
Spindasis crustaria Avnivitiius, 1898, Rhop. Aithiop., p, 332.
1920] Holland, Lepidoptera of the Congo 225
There is one somewhat damaged female taken at Medje about the
middle of July.
(351) 3. Spindasis aderna (Ploetz)
Plate XII, Figure 4, ¢7
Zeritis aderna Piarz, 1880, Stett. Ent. Zeit., XLI, p. 203, 7 and @.
Zeritis latifimbriata KE. SuHarpn, 1890, Ann. Mag. Nat. Hist., (6) VI, p. 105; 1890,
Trans. Ent. Soc. London, Pl. xvu, fig. 5, 2.
Spindasis aderna AuRIVILLIUS, 1898, Rhop. Aithiop., p. 332,
One male in not very good condition taken at Niangara about the
middle of November 1910. It agrees on the under side with a female
long in my possession, which was captured by the late Dr. A. C. Good
on the Ogové River thirty years ago. I once took this specimen with
me to London and it was examined by Miss Sharpe, who agreed with me
in my identification of it as her species latifimbriata. Aurivillius sinks
S. (Zeritis) fallax, 3, (Sharpe), as synonym of aderna Ploetz. I am quite
sure he is in error. Z. fallax Sharpe is bright blue on the upper side
of the wings, while S. aderna in the male sex resembles A-xiocerses
perion (harpax), as Ploetz, the author of the species, points out in his
description. The specimen before me accords with what Ploetz says
and might easily be mistaken by a novice for a specimen of A. harpax
upon casual inspection. Ploetz states in his paper that he had male
and female before him. The male of S. aderna on the upper side is
dark red and not blue, and Miss Sharpe’s blue species is distinct,
in spite of the superficial resemblance of the insect on the under side
to that named by Pleetz.
(352) 4. Spindasis chapini, new species
Plate XII, Figure 6, @
&. Frons reddish; a tuft of white hairs at the base of each antenna; antennze
black, short, with a moderately long spindle-shaped club, as in other species of the
genus; eyes dark brown, completely encircled with a ring of white scales; terminal
joint of palpi black, second and third joints heavily clothed with long vermilion
colored scales. Thorax and abdomen black above, deep orange-red inclining to
vermilion below. Legs with the femora and tibize dark brown dorsally, pale red below,
at their point of union with the thorax surrounded by a ring of tuft-like white hairs,
which is again encircled outwardly by a ring of long jet-black hairs. The fore wing
on the upper side is uniformly dark brown or sepia, except for a few dark orange-red
scales just before the hind angle, arranged to form a subtriangular spot, more or less
ill-defined. The hind wings above are of the same ground-color as the fore wings,
but about the middle of the discal area they are darker, being clothed with black
velvety scales. The entire inner margin is laved with deep orange-red, and this
color is extended over the whole posterior extremity of the wing, and along the outer
margin upward as a narrowing line to the extremity of vein 3. The dark brown cilia
226 Bulletin American Museum of Natural History (Vol. XLIII
define this pale area as a fine marginal band. The anal lobe is black at the end; the
two tails, one at the end of vein 1, which is twice as long as the one at the end of vein
2, are very slender, and appear under the microscope to be dark red, heavily dusted
with black scales. On the under side the ground-color of both wings is very deep
orange-red or vermilion. The fore wings on their posterior margin are pale fuscous
from the base, as far upward as the first submedian nervule and as far outward as a
point about two millimeters from the lower angle, where this dark shade vanishes
and is replaced by pale yellow. There is a small sharply defined black spot in the cell
of the fore wing at its base, followed about the middle by two similar spots, one
above the other, and succeeded immediately at the end of the cell by three such spots
fused together to form a moniliform dark bar closing the cell. The hind wings on
this side are devoid of dark markings, except on the inner margin a little above the
anal lobe, where there is an elongated subtriangular spot of small size, which is deep
black, ornamented in the middle by a narrow streak of silvery white scales. The anal
lobe is black ornamented with a few metallic scales. On this, as on the upper side of
the wings, the cilia are dark brown and define the wings outwardly as a narrow
marginal line. Expanse, 28 mm.
There are two males, both captured at Niangara in November
1910. One, the type, is in The American Museum of Natural History;
the other, the paratype, is in the Holland Collection in the Carnegie
Museum. I take pleasure in naming the pp in honor of one of the
leaders of the expedition.
Axiocerses Hiibner
(353) 1. Axiocerses harpax (Fabricius)
Papilio harpax Fasricius, 1775, Syst. Ent., App., p. 829, ?.
Chrysophanus perion Horrrer, 1862, Peters, Reise n. Mossambique, Ins., p. 4038, Pl.
XXVI, figs. 1-3.
Acxiocerses harpax AURIVILLIUS, 1898, Rhop. ASthiop., p. 335.
There are three males taken at Medje which are clearly referable
to this species. One was captured in June, the other two in July.
I confess with Dr. Aurivillius my inability to distinguish clearly
specimens of so-called A. perion (Cramer) which we now and then re-
ceive from correspondents in South and Eastern Africa from A. harpax
(Fabricius). Except that they are darker on the under side, so far as my
specimens show, there is no difference which should cause them to be
regarded as a distinct species. They seem to me at best to be a mere
local race. However, the whole genus is in need of revision, as a number
of species have recently been described.
1920] Holland, Lepidoptera of the Congo 227
CupipestHEs Aurivillius
(354) 1. Cupidesthes thyrsis (Hewitson)
Lycenesthes thyrsis Hewrrson, 1878, Ill. Diurn. Lep., Lycenide, p. 224, Pl. xcrt,
figs. 42-44. AurRIvILiius, 1898, Rhop. A2thiop., p. 349.
Cupidesthes thyrsis BetTHuNE-BaAkER, 1910, Trans. Ent. Soc. London, p. 10, Pl. rv,
fig. 3, genitalia.
- One male of this species, which is very common in Cameroon and
the valley of the Ogové, was captured at Niangara toward the end of
November 1910.
LYCZNESTHES Moore
(355) 1. Lycwnesthes musagetes Holland
Lycenesthes musagetes HOLLAND, 1893, Ent. News, IV, p. 25. AuRIvILLIUs, 1898,
Rhop. Athiop., p. 349. Beraunre-Baxer, 1910, Trans. Ent. Soc. London, p. 21.
Seven males, one taken at Avakubi, October 3, 1909, the rest at
Medje, the dates of capture ranging from April to August.
(356) 2. Lycenesthes ituria Bethune-Baker
Lycenesthes ituria BeTHUNE-BakeErR, 1910, Trans. Ent. Soc. London, p. 22, Pl. 1,
fig. 4; Pl. v, fig. 6.
I refer one badly damaged male taken at Medje, June 26, to this
species, with the figure and description of which it agrees closely, though
somewhat under the size of the specimen figured by the author of the
species.
(357) 3. Lycenesthes lunulata Trimen
Lycenesthes lunulata Trimen, 1894, Proc. Zool. Soc. London, p. 51, Pl. v1, fig. 12. _
(For synonymy consult Bethune-Baker, 1910, Trans. Ent. Soc. London, p. 40.)
One male captured at Faradje in December 1912.
(358) 4. lLycenesthes silvanus (Drury)
Papilio siluanus Drury, 1773, Ill. Exot. Ent., I, p. 5, Pl. m, figs. 2, 3.
(For synonymy consult Bethune-Baker, 1910, Trans. Ent. Soc. London, p. 43.)
The collection contains eight males, one taken at Lubila in Sep-
tember 1909, one at Niangara in November, and the rest at Medje from
May to August 1910.
All of the specimens differ from examples from tropical West Africa
(Sierra Leone, Cameroon, the valley of the Ogové) with which I have
compared them in the fact that the dark markings on the under side
of the wings do not contrast as strongly with the ground-color as in the
specimens from the west coast. It is true that none of the specimens
are in the best condition, and all are somewhat rubbed, but in spite of
228 Bulletin American Museum of Natural History [Vol. XLIII
this it is evident that they all show a tendency on the under side to a loss
of the distinctness of the maculation and a toning down in the direction
of uniformity of surface which causes them to stand in contrast to a
series of specimens from farther west. The markings are identical, in
their form, but the dark bands and spots are only a few shades darker
than the ground-color. The fact is worthy of note.
(359) 5. Lycenesthes larydas (Cramer)
Papilio larydas Cramer, 1780, Pap. Exot., III, p. 160, Pl. cctxxxni, fig. 6.
(For further synonymy consult Bethune-Baker, 1910, Trans. Ent. Soc. London,
p. 44.)
The collection comprises twelve males, one taken at Stanleyville
in September 1909, the rest at Medje from June to September 1910.
(360) 5a. Lycenesthes larydas kersteni Gerstecker
Lyceenesthes larydas var. kersteni GERST®CKER, 1871, Archiv f. Naturg., XX XVII, p.
359; idem, 1873, Von der Decken’s Reise, III, p. 373, Pl. xv, fig. 5.
Three males of this form taken at Faradje in December 1912.
There is a marked difference between the specimens of L. larydas com-
ing from the hot valleys and woodlands of the western part of its range
and those from the eastern parts of the African continent. The latter,
to which Gerstecker gave the name cited above, are always paler blue
on the upper side, and distinctly paler below. The difference is hardly
specific, but is an illustration, one of many, showing that climatic and
other influences are at work in producing variations on the two sides of
the continent.
(361) 6. Lycenesthes lachares Hewitson
Lycenesthes lachares Hewirson, 1878, Ill. Diurn. Lep., Lycenide, p. 225, Pl.
xcl, figs. 33, 34.
(For synonymy consult Bethune-Baker, 1910, Trans. Ent. Soc. London, p. 47.)
One female taken at Medje, July 5, 1910.
(362) 7. Lycenesthes rufomarginata Bethune-Baker
Lycanesthes rufomarginata Betnune-Baker, 1910, Trans. Ent. Soc. London, p. 54,
Pl. 11, fig. 14; PI. 1x, fig. 26.
One male captured at Medje, April 5, 1910.
(363) 8. Lycwnesthes makala Bethune-Baker
Lycanesthes makala Beruune-Baxer, 1910, Trans. Ent, Soc. London, p. 58, Pl. 1,
fig. 16; PI. 1x, figs. 27, 28.
One male taken at Medje about the middle of July.
1920] Holland, Lepidoptera of the Congo 229
(364) 9. Lycenesthes scintillula Holland
Lycenesthes scintillula HoLuAND, 1891, Psyche, VI, p. 50. SmrrH anp Kirsy, 1893,
Rhop. Exot., Afr. Lycenide, p. 98, Pl. xxm, figs. 3, 4. Beraune-Baxker, 1910,
Trans. Ent. Soc. London, p. 60, Pl. 11, fig. 3; Pl. x, figs. 30, 31.
Three males taken at Medje, one in each of the months, May, June,
and July.
(365) 10. Lycenesthes pyroptera Aurivillius
Lycenesthes pyroptera Aurivitiius, 1895, Ent. Nachr., X XI, p. 382; 1899, Rhop.
AMthiop., p. 352, Pl. vi, fig. 3. Beraune-Baxer, 1910, Trans. Ent. Soc. London,
p. 61.
One male taken at Stanleyville, September 5, 1909. The specimen
agrees absolutely with the description and figure given by Aurivillius.
TRICLEMA Karsch
(366) 1. Triclema lutzi, new species
Plate XII, Figure 2, #7
&. Near 7’. rufoplagata Bethune-Baker (cf. Trans. Ent. Soc. London, 1910, p.
72, Pl. 1m, fig. 8) but considerably smaller in size, and differently marked. The thorax
and abdomen are black above, pale chestnut below. The chestnut area of the fore
wings, which in rufoplagata is small and confined to the immediate region about the
origin of veins 2 and 3, is in 7’. lutzi extended so as to cover the greater part of the discal
area, only the base, the costa, and a broad marginal band remaining dark in color.
At the end of the cell, which in its lower half is distally invaded by the chestnut color,
there is a narrow black vertical bar, which merges into the dark costal area. The
secondaries are very dark brown (sepia) almost black, with a fine light line followed
by a very thin black line on the outer border, the fine dark line showing against the
light line within, and the paler cilia. There is an imperfect ocellus at the anal angle
and a more regularly formed ocellus in the space between veins 2 and 3 at their
extremities. These eye-spots are defined inwardly by a few scales slightly lighter
than the rest of the wing. On the under side the ground-color of the wings is mod-
erately dark brownish fuscous, crossed by darker lines and bands, which are defined
more or less clearly on either side by lighter lines. The costa of the primaries at the
base is black for a short distance, there is a light line following, and distally a rather
prominent black triangular spot with its apex pointing toward the costa; there isa .
dark vertical bar near the end of the cell, defined inwardly and outwardly by paler
lines; the median transverse line is curved somewhat irregularly, and widens and
grows blacker as it approaches the inner margin; the postmedian macular band is
widest and darkest between veins 3 and 5; the submarginal band is accentuated by
black spots before the apex in spaces 6 and 7; there is a thin light line just before the
equally thin dark marginal line which latter is clearly defined against the rather light
colored cilia. The hind wings below have several dark basal and subbasal short trans-
verse spots. At the end of the cell is a double bar of dark color defined by paler lines
before and behind; the median band, which runs from the costa to the inner margin
in an irregular curve is accentuated at the middle of the costa by a dark quadrate spot
230 Bulletin American Museum of Natural History (Vol. XLII
and terminates on the inner margin in the usual V-shaped spot; the postmedian
macular band is darkest and broadest opposite the end of the cell, and is edged in-
wardly and outwardly by lighter lines; the submarginal line which is somewhat
irregularly curved is thinner and not as conspicuous as the other lines; it is sueceeded
just before the margin by a series of semilunate darker spots in the interspaces from
the upper angle as far down as vein 3; between veins 2 and 3 there is a conspicuous
ocellus, defined above and on the sides, but not outwardly, by red. The pupilis deep
black irrorated with a few bright metallic blue scales; at the inner angle there is an
imperfectly formed ocellus, red above, pupilled with black, and dusted with a few
bright scales. The marginal line is thin and dark and contrasts against the paler
shade within and the lighter cilia without. Expanse, 16-20 mm.
The collection contains three specimens, all caught at Medje, one in
April, one in June, and one in July. They are males. The type and a
paratype are in The American Museum of Natural History, one para-
type is in the Holland Collection in the Carnegie Museum. I take plea-
sure in naming this species in honor of the amiable curator of the entomo-
logical collections of the American Museum.
Puiyaria Karsch
(367) 1. Phlyaria cyara (Hewitson)
Lycena cyara Hewr1son, 1876, Exot. Butt., V, Lycena, Pl. 1, figs. 9, 10.
Phlyaria cyara Karscu, 1895, Ent. Nachr., XX1I, p. 303.
Cupido phlyaria Aurivituivs, 1898, Rhop. Acthiop., p. 358.
There are two males, one caught at Risimu in September 1909, the
other at Medje in July 1910. The insect is not uncommon in the
interior of Cameroon, but I have never seen a female and this sex seems,
so far as I can now recall, never to have been thus far described.
(368) 2. Phlyaria heritsia (Hewitson)
Lycena heritsia Hewrrson, 1876, Exot. Butt., V, Lycena, Pl. 1, figs. 11, 12.
Hyreus virgo Bot er, 1896, Proc. Zool. Soc. London, p. 121, Pl. v1, fig. 1, 9.
Cupido heritsia Aurivitur0s, 1898, Rhop. Aithiop., p. 359.
Five males, one taken at Bafwasende, September 23, 1909, the rest
at Medje from April to September.
UranotHauma Butler
(369) 1. Uranothauma falkensteini (Dewitz)
Plebeius falkensteini Dewirz, 1879, Nov. Act. Acad. Nat. Cur., XLI, part 2, p. 205,
Pl xxvi, fig. 5.
Cupido falkensteini Avnivizrus, 1898, Rhop. Aethiop., p. 360.
Four males captured at Medje, June, August, and September.
1920] Holland, Lepidoptera of the Congo 231
Castatius Hiibner
(370) 1. Castalius carana (Hewitson)
Lycena carana Hewson, 1876, Exot. Butt., V, Lycena, Pl. 1, fig. 6.
Cupido carana AuRIVILLIUS, 1898, Rhop. Aithiop., p. 364.
This species, which is quite common in Cameroon and the valley of
the Ogové, was taken in some numbers by the expedition at Medje,
where twenty-three specimens were taken, the dates of capture ranging
from July to September. There is also a specimen taken at Bafwabaka,
January 7, and one caught at Gamangui on February 1, 1910. Some of ©
the females are tinged with yellowish.
The species must not be confounded with C. margaritaceus Em.
Sharpe, paratypes of which are in the writer’s collection, and which it
superficially resembles.
(871) 2. Castalius isis (Drury)
Papilio isis Drury, 1773, Ill. Exot. Ent., I, p. 6, Pl. m1, figs. 4, 5.
Cupido isis AURIVILLIUS, 1898, Rhop. Aithiop., p. 365.
The female of this pretty insect seems not to have been described
hitherto. As there is one example of this sex in the collection I may say
that there is no difference in the form of the markings to distinguish it
from the males but that all the bands and spots instead of being brilliant
blue as in the male, are blackish in the female, there being only a faint
trace of the blue color in this sex at the bases of the fore and hind wings.
The collection contains twenty-four males and one female. Most
of them were taken at Medje from April to September, but two or three
were captured at Gamangui in February and June, and several at
Niangara in November 1910.
Tarucus Moore
(372) 1. Tarucus telicanus (Lang)
Papilio telicanus LANG, 1789, Verzeichniss meiner Schmetterlinge, part 2, p. 47.
Hesperia (Rurales) plinius Fasricius, 1793, Ent. Syst., III, part 1, p. 284.
Cupido telicanus var. plinius AURIVILLIUS, 1898, Rhop. Aithiop., p. 364.
There are three males and one female. One of the males was caught
at Niangara, the rest were taken at Medje in July and August.
Azanus Moore
(373) 1. Azanus mirza (Ploetz)
Lycena mirza Piarz, 1880, Stett. Ent. Zeit., XLI, p. 203.
Cupido mirza AuRIVILLIUS, 1898, Rhop. Athiop., p. 366.
232 Bulletin American Museum of Natural History [Vol. XLIII
The species is represented in the collection by thirty-one specimens,
twenty-six taken at Gamangui in February, three at Medje, one in each
of the months, June, July, and September, one at Bafwabaka in Dec-
ember 1909, and one at Faradje, labelled “1911-1912.”
NacapusBa Moore
(374) 1. Nacaduba ethiops (Mabille)
Kharsanda ethiops MaBIL1e, 1877, Bull. Soc. Zool. France, II, p. 219.
Nacaduba stratola HoLLAND, 1891, Psyche, VI, p. 52.
Cupido ethiops AuURIVILLIUS, 1898, Rhop. Aithiop., p. 367.
There are thirteen males, eleven caught at Gamangui in February,
the other two at Medje in July 1910.
Potyommatus Moore
(375) 1. Polyommatus beticus (Linneus)
Papilio beticus Linnxvs, 1767, Syst. Nat., 12th Ed., p. 789.
Cupido beticus AURIVILLIUS, 1898, Rhop. Athiop., p. 367.
Two males, one caught at Medje in August 1910, the other labelled
“Faradje, 1911-1912.”
Cupipo Schrank
(376) 1. Cupido asopus (Hopffer)
Lycena asopus Horrrer, 1855, Monatsber. Akad. Wiss. Berlin, p. 642.
Cupido asopus AuRIvILLIus, 1898, Rhop. A®thiop., p. 373. (As synonym of C. mala-
thana Boisduval.)
Two damaged specimens, a male taken at Bafwabaka, December
1909, and a female caught at Medje, July 1910.
Aurivillius sinks asopus Hopffer as a synonym of C. malathana
Boisduval, but I cannot bring myself to agree with him, in the light of the
material before me, which I believe to be correctly identified. As species
go in this group, there seem to be reasons for regarding the two as
distinct. But I will not discuss the matter except to say that the form
malathana from Madagascar is in my opinion at least a well-marked
insular variety and easily separable from the continental form asopus.
(377) 2. Cupido osiris (Hopffer)
Lycana osiris Horrrer, 1855, Monatsber, Akad. Wiss. Berlin, p. 622.
Cupido osiris Aurtvitires, 1898, Rhop. Aithiop., p. 374.
There are four males, all taken at Faradje, one without date, one
caught in December 1912, and two labelled 1911-1912.” Besides,
there is a dwarfed female captured, according to the label, at Bumba,
July 1909.
1920] Holland, Lepidoptera ‘of the Congo 233
(378) 3. Cupido patricia (Trimen)
Lycena patricia Trimen, 1881, 8. Afr. Butt., II, p. 20.
Cupido patricia AurivILLIUs, 1898, Rhop. Athiop., p. 374.
There are eight specimens, one male, and seven females; one fe-
male taken at Kwamouth, July 1909, the rest at Niangara in November
1910. They have been carefully compared with specimens of C. (Ly-
cena) patricia (Trimen), received from Mr. Trimen, the author of the
species, and appear to agree with them. I may say, in passing, that I
think that the species represents a varietal form of C. parsimon (Fab-
ricius) and it is hard to define the difference between the two forms
without very carefuly scrutiny.
Cupipopsis Karsch
(379) 1. Cupidopsis hippocrates (Fabricius)
_ Hesperia (Rurales) hippocrates Fasricius, 1793, Ent. Syst., III, part 1, p. 288.
Cupido hippocrates AuRIvILLIUS, 1898, Rhop. Aethiop., p. 376.
Cupidopsis hippocrates NBaveE, 1910, Proc. Zool. Soc. London, p. 601.
Two males taken at Medje in August 1910.
I follow Neave in referring this species to the genus Cupidopsis
Karsch, the type of which is C. jobates. It might as well be placed here
as anywhere, unless a new genus be erected for its reception. In its
markings it shows little resemblance to C. jobates, but structurally it is
very close to that species. The female is figured in Mabille’s work on
the Lepidoptera of Madagascar, included in Grandidier’s magnificent
series of volumes on the natural history of that island. It differs from
the male in lacking the white at the apices of the fore wings, and in
having the hind border of the secondaries narrowly margined with
white upon which the marginal ocelli are conspicuous. I have an ex-
tensive series of this species from various parts of Africa, west, east, and
south.
Everes Hiibner
(380) 1. Everes togara (Plotz)
Lycena togara Piarz, 1880, Stett. Ent. Zeit., XLI, p. 202.
Cupido micylus var. togara AuRIVILLIUS, 1898, Rhop. 2thiop., p. 377.
' This species, which is common in Cameroon and the valley of the
Ogové, is represented in the collection by three tattered specimens, a
male taken at Medje in September and two females caught at Medje,
one in April, the other in August.
Aurivillius regards this insect as a varietal form of E. micylus
(Cramer) but Mr. Bethune-Baker informs me that a careful study of
the genitalia made by him reveals that togara (Ploetz) is specifically
distinct from micylus (Cramer).
234 Bulletin American’ Museum of Natural History [Vol. XLIII
ZizERA Moore
(381) 1. Zizera antanossa (Mabille)
Lyceena antanossa MaBIL_e, 1877, Ann. Soc. Ent. France, (3) VII, Bull., p. 72.
Cupido antanossa AURIVILLIUS, 1898, Rhop. Aithiop., p. 378.
Zizera antanossa BUTLER, 1900, Proc. Zool. Soc. London, p. 100, Pl. x1, figs. 14, 15.
Two poorly preserved males taken at Faradje, one, without date,
the other labelled “1911-1912.”
(382) 2. Zizera gaika (Trimen)
Lycena gaika TRIMEN, 1862, Trans. Ent. Soc. London, p. 403.
Cupido gaika Aurtvituius, 1898, Rhop. Aithiop., p. 378.
Zizera gaika ButLER, 1900, Proc. Zool. Soc. London, p. 109.
One male taken at Bumba, July 29, 1909.
(383) 3. Zizera lysimon (Hiibner):
Papilio lysimon Hitpner, 1798-1803, Europzische Schmett., figs. 534, 535.
Cupido lysimon AurIviLutus, 1898, Rhop. Aithiop., p. 379.
Zizera lysimon But_ER, 1900, Proc. Zool. Soc. London, p. 108, Pl. x1, fig. 9.
Two specimens captured at Freetown, Sierra Leone, July 14, 1909.
Osporonia Karsch
(384) 1. Oboronia plurilimbata (Karsch)
Thermoniphas plurilimbata Karscu, 1895, Ent. Nachr., X XI, p. 303.
Cupido plurilimbata Aurtvituius, 1898, Rhop. Aithiop., p. 380.
Three males taken at Medje, one in April, one in August, and one in
September. They agree completely with the description given by Karsch,
and I refer them to his species, of which no figure has been published,
so far as I can recall.
(385) 2. Oboronia punctata (Dewitz)
Plebeius punctatus Dewrtz, 1879, Nov. Act. Acad, Nat. Cur., XLI, part 2, p. 205,
Pl. xxvi, fig. 15.
Cupido punctatus AuRivILLIUs, 1898, Rhop. Aithiop., p. 381.
One male caught at Munie Katoto, September 10, 1909.
This is a very common species in southern Cameroon and the
valley of the Ogové.
(386) 3. Oboronia ornata (Mabille)
Lycana ornata MawiiLe, 1890, Ann, Soc. Ent. France, (6) X, p. 24, Pl. uy, fig. 6.
Cupido ornatus Avrivitiius, 1898, Rhop. Aithiop., p. 381.
Three specimens taken at Medje, two in April. one in June.
1920] Holland, Lepidoptera of the Congo 235
Like the foregoing, this is a very common insect in southern
Cameroon and the valley of the Ogové River. We have in the Museum
at Pittsburgh more than one hundred examples, including the slight
varietal form named vestalis by Aurivillius, in which the hind marginal
band of the secondaries is broader than in the type figured by Mabille,
and several specimens in which the white areas of the wings are broadly
beautiful straw-yellow. To this yellow form, which has not heretofore
been described or figured, I apply the varietal name flava. The type is
in the Holland Collection in the Carnegie Museum and, with ‘several
cotypes, was taken at Benito, Spanish Guinea, by the late Dr. A. C. Good.
In this connection I may state that this insect is the one to which
the Fabrician name elorea has in the past been applied by authors. The
Fabrician description, like many of those given by early authors, is too
brief and concise to enable a student to reach a proper conclusion. The
only figure of elorea is given by Donovan in the ‘Naturalist’s Repository,’
Pl. ti, in reference to which Aurivillius makes the comment that
it may be “ species fictitia.”” 1 confess that among the hundreds of speci-
mens of this genus I have had in my hands I have never found one to
agree with Donovan’s figure, and that I share in the view of Aurivillius
that if it is elorea Fabricius, the species is now probably extinct, if
indeed it ever existed.
Papilionide
Pierinse
Leptrosia Hiibner
(387) © 1. Leptosia medusa (Cramer)
Papilio medusa CRAMER, 1777, Pap. Exot., II, p. 86, Pl. ct, fig. F.
Leptosia medusa AURIVILLIUS, 1898, Rhop. Aithiop., p. 387.
Eight specimens, one taken at Bafwasende in January, one at
Gamangui in February, the rest at Medje in July and August.
(388) la. Leptosia medusa immaculata (Aurivillius)
Nychitona medusa immaculata Aurtviuui0s, 1895, Ent. Tidskr., XVI, p. 257.
Leptosia medusa ab. immaculata Auriviturus, 1910, Seitz, Gross-Schmett., XIII,
p. 31, Pl. xa.
Three examples, one taken at Ngayu in April, one at Medje in
August, and one at Niangara in November.
236 Bulletin American Museum of Natural History [Vol. XLIIT
(389) 2. Leptosia nupta (Butler)
Nychitona nupta BuTLER, 1873, Cist. Ent., I, p. 175.
Leptosia nupta AuRIvILuI0s, 1910, Seitz, Gross-Schmett., XIII, p. 31, Pl. xb.
Twenty-two specimens, of which all were captured at Medje
except two caught at Gamangui in February and one in June. The
specimens from Medje were mostly taken in June, but there are indivi-
duals labelled as captured there in every month from March to August
inclusive.
Mytorturis Hiibner
(390) 1. Mylothris chloris (Fabricius)
Papilio chloris Fasricrus, 1775, Syst. Ent., p. 473.
Mylothris chloris AuRtviL1i10s, 1910, Seitz, Gross-Schmett., XIII, p. 32, Pl. xd.
A male and a female taken at Niangara in November, and another
female caught at Faradje “1911-1912.”
(391) 2. Mylothris sjéstedti Aurivillius
Mylothris sjéstedti AuRtviLui0s, 1895, Ent. Tidskr., XVI, p. 260, Pl. m1, fig. 2; 1910,
Seitz, Gross-Schmett., XIII, p. 33, Pl. xia.
I refer to this species a single male, taken at Medje, August 24,
1910. It differs somewhat from the type, in that the blue-gray color
extends from the base of the fore wing almost entirely over the wing,
and is not restricted to its basal area. In other respects it agrees en-
tirely with the figures and descriptions given by the author of the species.
(392) 3. Mylothris spica (Movschler)
Tachyris spica Maescuuer, 1883, Verh. zool.-bot. Ges. Wien, X XXIII, p. 277, &.
Papilio rhodope Donovan, 1824, Naturalist’s Repository, II, Pl. uxxxvi (non
rhodope Fabricius), 9 .
Mylothris spica Aurtvi1110s, 1910, Seitz, Gross-Schmett., XIII,p. 33, Pl. xd, co”.
There has existed some confusion as to this species. I have in my
possession a long series of males and females collected for me by the late
Dr. A. C. Good in the valley of the Ogové River. A number of pairs
were taken in coitu. This series shows clearly that the insect, which was
in 1883 described by Meeschler under the name spica, is the male of the
butterfly which Donovan in 1824 figured as the female of P. rhodope
Fabricius, which itis not. For the form figured by Donovan I now propose
the name donovani to distinguish it from other female forms of the same
species. Besides the females belonging to this varietal form there are
numerous other females, also taken in coitu with males of M. spica, in
1920] Holland, Lepidoptera of the Congo 237
which the marginal borders more nearly resemble those of M. poppea
(Cramer). In these, however, the fore wings are either white or very
faintly laved with yellow. The females of this species are therefore
polymorphic.
In the Lang-Chapin Collection there is only one female which be-
longs to the form M. donovani Holland. It was taken at Niangara in
November. There are twenty-seven malesof typical spica, one captured
at Kwamouth in July 1909, another at Avakubi in October 1909, a couple
at Gamangui in June, and the rest at Medje in June, July, and August
1910.
(393) 4. Mylothris sulphurea Aurivillius
Mylothris sulphurea Aurivituius, 1895, Ent. Tidskr., XVI, p. 259, Pl. m1, fig. 3;
1910, Seitz, Gross-Schmett., XIII, p. 33, Pl. xf. :
Six males and two females, all taken at Medje, except one male
caught at Munie Katoto in September. The specimens from Medje
were taken from June 20 to August 24, 1910.
(394) 5. Mylothris rubricosta (Mabille)
Pieris rubricosta MaBiLE, 1890, Ann. Soc. Ent. France, (6) X, p. 28.
Mylothris rubricosta AuRIvILLIUs, 1910, Seitz, Gross-Schmett., XIII, p. 35, Pl. xtc.
Six males and two females taken at Medje in June, July, and August.
Thiss pecies is not very far removed from M. bernice (Hewitson), from
which it differs in having the marginal border and spots of both wings
smaller, and the female not suffused so broadly with fuscous upon the
lighter areas of the fore and hind wings. I have a long series of M.
bernice from the Ogové, from which this form differs in the respects
noted, but I am not convinced that they are specifically distinct.
Appias Hiibner
(395) 1. Appias rhodope (Fabricius)
Papilio rhodope Fasricius, 1775, Syst. Ent., p. 473.
Appias rhodope AuRIvVILLIUs, 1910, Seitz, Gross-Schmett., XIII, p. 37, Pl. x1d, 7 ;x1e,9.
Seventeen males and four females. One male was taken at Batama
in September 1909; one female at Risimu in September of the same
year; one female at Niangara in November 1910; and the rest at
Medje from June to August inclusive.
(396) 2, Appias phaola (Doubleday)
Pieris phaola DousuepAy, 1847, Ann. Mag. Nat. Hist., (1) XX, p. 63.
Appias phaola AuRIvILLIvs, 1910, Seitz, Gross-Schmett., XIII, p. 37, Pl. x1c, #7; xd,
Q
A single male taken at Medje, August 20, 1910.
238 Bulletin American Museum of Natural History [Vol. XLIII
(397) 3. Appias sabina (Felder)
Pieris sabina FELDER, 1865, Reise Novara, Lep., III, p. 165.
Appias sabina AURIVILLIUS, 1910, Seitz, Gross-Schmett., XIII, p. 38, Pl. x1f, 3;
xe, 9.
The collection contains twenty-three males and two females. One
male was taken at Kwamouth in July 1909. One female from Lubila,
two males from Risimu, and another from Munie Katoto are, according
to the labels, captures made in September 1909. One male was taken
at Bafwasende in January, and four at Gamangui in February 1910.
All the rest were caught at Medje at dates running through June, July,
and August.
(398) 4. Appias epaphia (Cramer)
Papilio epaphia Cramer, 1779, Pap. Exot., III, p. 26, Pl. ccvn, figs. D, B.
Appias epaphia Aurtvitttus, 1910, Seitz, Gross-Schmett., XIII, p. 38, Pl. x1f, 3.
One male taken at Kwamouth, July 15, 1909.
(399) 5. Appias sp. (?)
There is a single female taken at Gamangui, February 1, 1910,
which agrees with that sex of A. epaphia with the exception that the
base of the primaries and the limbal area of the secondaries lack the dark
markings which are characteristic of this sex in that species. As the
female of A. epaphia is somewhat variable, I hesitate to describe this
even as a new variety, though it differs widely in appearance from any-
thing the figures and descriptions of which are known to me. It evi-
dently is a very near ally of the foregoing species. The apical markings
on both sides of the fore wing are identical with those found in the fe-
males of that species, but the fact that the remaining areas of both the
fore and hind wings are white, except for a little dark squamation near
the base, makes the specimen contrast markedly with the long series of
A. epaphia in my collection, with which I have compared it.
Preris Godart
(400) 1. Pieris creona (Cramer)
Papilio creona Cramer, 1776, Pap, Exot., I, p. 148, Pl. xcv, figs. C, F.
Pieris creona Aunivitii0s, 1910, Seitz, Gross-Schmett., XIII, p. 40, Pl. x1mb.
Two males taken at Faradje in November, one female caught at
Niangara in the same month, and a female captured at Medje, August
24, 1910, constitute the only representatives of this species in the
collection.
1920] Holland, Lepidoptera of the Congo 239
(401) 2. Pieris infida Butler
Pieris infida Butter, 1888, Proc. Zool. Soc. London, p. 77. AurtviLtiius, 1910,
Seitz, Gross-Schmett., XIII, p. 41, Pl. xm.
There are two males and two females of this species in the collec-
tion. The males were taken at Faradje and the females at Niangara.
One male bears the date ‘‘ November 1910"; the rest of the specimens
are simply marked “1911-1912.”
(402) 3. Pieris calypso (Drury)
Papilio calypso Drury, 1773, Ill. Exot. Ent., II, p. 30, Pl. xvm, figs. 3,4, 2.
Belenois (Pieris) welwitschii RoGeNHoFER, 1889, Ann. K. K. Hofmuseums Wien, IV, p.
548, Pl. xxi, fig. 2, 9.
Pieris calypso AURIVILLIUs, 1910, Seitz, Gross-Schmett., XIII, p. 42, Pl. xd.
Of this common and well-known species there are forty-two males,
two females of the typical form with white hind wings, and four females
of the variety named welwitschi by Rogenhofer, in which the hind wings
are orange. The specimens were mostly taken at Medje, a few at
Gamangui, some at Niangara, and others at scattered localities. The
dates of capture are those associated with these localities on all of the
labels.
(403) 4. Pieris theora Doubleday
Pieris theora DouBLEDAY, 1846, Ann. Mag. Nat. Hist., (1) XVII, p. 25. Aurrviiurus,
1910, Seitz, Gross-Schmett., XIII, p. 43, Pl. x1e.
Five males, one taken at Basoko in July 1909, one at Bafwasende
in September of the same year, and three captured at Medje, one in
each of the months, June, July, and September.
(404) 5. Pieris theuszi Dewitz
Pieris theuszi Dewirz, 1889, Ent. Nachr., XV, p. 107, Pl. u, figs. 6-9. AuRIvILLIUs,
1910, Seitz, Gross-Schmett., XIII, p. 44, Pl. xrva.
Nine males and one female. The female, which is in not very good
condition, was taken at Batama in September 1909; one male was
caught at Bafwasende in the same month and year; a couple were taken
at Niangara in November; and the remaining six were captured at
Medje in June, July, and August.
(405) 6. Pieris solilucis Butler
Pieris solilucis BuTLER, 1874, Trans. Ent. Soc. London, p. 433. AurtviLiius, 1910,
Seitz, Gross-Schmett., XIII, p. 44, Pl. xtva.
All of the seventeen specimens are males. One was taken at Medje
in August, the remaining sixteen were caught at Niangara in November.
240 Bulletin American Museum of Natural History [Vol. XLIIT
TERACOLUS Swainson
(406) 1. Teracolus evippe (Linnus)
Papilio evippe Linn=vus, 1764, Mus. Lud. UIr., p. 239.
Teracolus evippe AURIVILLIUS, 1910, Seitz, Gross-Schmett., XIII, p. £7, Pl. xvime.
_ [refer a specimen without abdomen and with badly tattered wings
to this species. It was taken at Matadi, June 20, 1909. It is singular
that this specimen should be the only representative of this great genus,
which has its metropolis in Africa and which should be represented by a
number of species in the eastern parts of the region visited by the ex-
pedition, the land of the ‘White Rhinoceros.” It is absent from the
densely wooded jungles of the great river-valleys, but appears in the
grass-lands, wherever they occur south of the Sahara.
TERIAS Swainson
This genus, owing to the seasonal and loeal variations which occur
in every species, presents many puzzling problems, some of which still
await satisfactory solution. A multitude of so-called species, some of
which are of doubtful validity, have been erected upon slight but ap-
parently more or less constant differences. The test of breeding will
alone serve to decide finally what is the exact relationship of some of the
forms. The writer has had in his hands at various times, and still
possesses, vast numbers of specimens representing the various forms
occurring in Africa, but within certain limits has been hitherto unable
to satisfy himself as to the proper classification of some of the varieties.
The species appear to run into each other, and to be subject to great
variation both in size, coloration, and markings. The following account
of the specimens brought home by the American Museum Congo
Expedition represents the results of very careful study and comparison
with other specimens, many of which had been originally determined
by direct comparison with the types in European collections, but never-
theless it leaves something to be desired, especially in the case of 7.
senegalensis and its varieties, which is extremely variable.
(407) 1. Terias brenda Doubleday and Hewitson
Terias brenda DovusLepay AND Hewrrson, 1847, Gen. Diurn. Lep., I, p. 79, Pl. rx,
fig. 6. Aunivitiies, 1910, Seitz, Gross-Schmett., XIII, p. 64, Pl. xx1te,
Seventy-six males and two females, taken at almost every place
where collections were made. The dates of capture represent every
month in the year.
1920] Holland, Lepidoptera of the Congo 241
(408) la. Terias brenda maculata Aurivillius
Terias brenda var. maculata Aurtviuuius, 1910, Seitz, Gross-Schmett., XIII, p. 64,
Pl. xxue.
Two males taken at Gamangui in February, and an aberrant female
caught at Bafwasende in January. This is the winter form of the species.
(409) 2. Terias senegalensis Boisduval
Terias senegalensis BoispuvAL, 1836, Spec. Gén. Lép., I, p. 672. Aurtvitiius, 1910,
Seitz, Gross-Schmett., XIII, p. 64, Pl. xx1e.
The specimens which I refer to this species fall into two groups, one
in which the marginal band of the prinfaries at the outer angle is broad
and strongly produced on vein 2 as an inwardly projecting tooth, the
other in which it is narrow and only slightly angulated on vein 2. Of
the first group there are in the present collection twenty-two males, the
majority taken at Faradje in November. One is labelled as from Munie
Katoto, one from Kwamouth, one from Gamangui, and one from Basoko.
The four last mentioned were taken in the months of June and July.
Of the second group there are twenty-six males and six females. Of
these, thirteen males and two females were taken at Medje, one male and
one female in January, one male in September, all the rest in June and
July. Two males and a female are labelled as from Gamangui, taken in
February, the remainder at Niangara and Faradje in November, so
far as the labels indicate the date of capture.
(410) 2a. Terias senegalensis bisinuata (Butler)
Terias bisinuata Butrier, 1876, Ann. Mag. Nat. Hist., (4) XVIII, p. 485. Aurivit-
Lius, 1910, Seitz, Gross-Schmett., XIII, p. 64, Pl. xx1id.
This, which is claimed to be the winter form of 7. senegalensis and
is characterized by heavy markings on the under side of the wings, is re-
presented in the collection by fifteen males, twelve taken at Faradje in
November, so far as the labels are dated, though many of the labels
lack any indication of the month. One was captured at Niangara in the
month named, one at Kwamouth on July 15, 1909, and one at Medje
in August. Most of the specimens have the marginal band of the
primaries broad at the outer angle and strongly toothed inwardly, but
there are several in which it is narrow and only slightly angulated on
vein 2, in this respect resembling the specimens in the second group re-
ferred to under the preceding form.
e
242 Bulletin American Museum of Natural History [Vol. XLIII
(411) 3. Terias floricola ceres (Butler)
Terias floricola Botspuvat, 1833, Faune Ent. Madagr., p. 21.
Terias ceres BUTLER, 1875, Ann. Mag. Nat. Hist., (5) XVII, p. 218, Pl. v, fig. 3.
AvRIVILLIUS, 1910, Seitz, Gross-Schmett., XIII, p. 64, Pl. xxud.
There are two specimens, a male caught at Faradje, “1911-1912,”
and a female taken at Niangara in November.
(412) 4, Terias hapale Mabille
Terias hapale MABILLE, 1882, Le Naturaliste, II, p. 99. AUAIVILLIUS, 1910, Seitz,
Gross-Schmett., XIII, p. 65, Pl. xx1d.
Of this species there are four males and one female, all taken at
Faradje. There is no clue to the month, the specimens being simply
labelled ‘‘ 1911-1912.”
(413) 5. Terias desjardinsi Boisduval
Terias desjardinsi BoispUVAL, 1833, Faune Ent. Madagr., p. 22, Pl. 11, fig. 6. AURIVIL-
Lius, 1910, Seitz, Gross-Schmett., XIII, p. 65, Pl. xxtte.
Of this species there are four typical males, all taken at Niangara
in November. There are nineteen other males taken at Niangara and
Faradje, which are mostly smaller in size and with broader marginal
' bands, marking a transition in the direction of the following form, 7’.
regularis, and intermediate between it and typical desjardins?.
(414) 6. Terias regularis Butler
Terias regularis Butuer, 1876, Ann. Mag. Nat. Hist., (4) XVIII, p. 486. AurrviL-
Lius, 1910, Seitz, Gross-Schmett., XIII, p. 65, Pl. xxue.
This is regarded by Professor Aurivillius as an extreme summer
form of the preceding, and is characterized by the great enlargement of
the width of the marginal bands.
There are in the collection thirty-seven males and four females.
Some, including all the females, were taken at Medje in June and July,
some are labelled as taken at Niangara in November, one at Bafwabaka
in January, another at Gamangui in February, and a number at Faradje,
“1911-1912.”
(415) 7. Terias brigitta (Cramer)
Papilio brigitia Cramen, 1780, Pap. Exot., IV, p. 82, Pl. cccr, figs. B, C.
Terias brigitta Aunivitius, 1910, Seitz, Gross-Schmett., XIII, p. 65, Pl. xxuf,
One damaged male taken at Matadi, June 24, 1909.
1920] Holland, Lepidoptera of the Congo 242
(416) 8. Terias zoé Hopffer
Terias zoé Hoprrer, 1855, Monatsber. Akad. Wiss. Berlin, p. 640. AURIVILLIUS,
1910, Seitz, Gross-Schmett., XIII, p. 65, Pl. xxu/f.
One male taken at Niangara in November, and five males and two
females caught at Faradje, 1911-1912.”
Eronia Boisduval -
(417). 1. Eronia pharis (Boisduval)
Pieris pharis Botspuvau, 1836, Spec. Gén. Lép., I, p. 443.
Eronia pharis Auriviuui0s, 1910, Seitz, Gross-Schmett., XIII, p. 62, Pl. xxte.
Twelve males and six females, two of the former taken at Niangara
in November, and one of the latter at Gamangui in June. All the other
specimens were captured at Medje, the dates ranging from early in
June to the end of August.
(418) 2. Eronia thalassina (Boisduval)
Pieris thalassina BoispuvAL, 1836, Spec. Gén. Lép., I, p. 443.
Eronia thalassina AuRIvILLIvs, 1910, Seitz, Gross-Schmett., XIII, p.62, Pls.xve, xx1e.
Twenty-five males, one of which was taken at Bafwasende in Sep-
tember, all the rest at Medje, a few in April, and the others from June to
September.
(419) 3. Eronia argia (Fabricius)
Papilio argia Fasricivus, 1775, Syst. Ent., p. 470.
Eronia argia Aurtvitiius, 1910, Seitz, Gross-Schmett., XIII, p. 62, Pl. x1e, d.
Twenty-two males and one female, all taken at Medje, except one
male caught at Boyulu in September 1909, and another at Niangara
in November 1910. Of the specimens captured at Medje a few are
dated as of April, the remainder as of June to September.
Catopsitia Hiibner
(420) 1. Catopsilia florella (Fabricius)
Papilio florella F asricius, 1775, Syst. Ent., p. 479.
Catopsilia florella AuRIvILLIUS, 1910, Seitz, Gross-Schmett., XIII, p. 63.
Sixty-nine males and six females. Most of the specimens, including
all of the females, were caught at Niangara in November. There are a
few taken at Gamangui in February and at Stanleyville and Medje in
July and August.
244 Bulletin American Museum of Natural History [Vol. XLIII
Papilioninze
Papitio Linnzeus
There are about sixty species and subspecies of this genus which are
recorded from the African continent. Of these, twenty-three are re-
presented in the collection upon which I am reporting.
(421) 1. Papilio antimachus Drury
Papilio antimachus Drury, 1782, Ill. Exot. Ent., III, p.1, Pl.1. Aurtviiiius, 1908,
Seitz, Gross-Schmett., XIII, p. 11, Pl. 1a.
One good and two somewhat damaged males. One is labelled as
from Batama, taken in September 1909, another from Avakubi, May
1910, the third from Bafwasende, September 1910.
(422) 2. Papilio zalmoxis Hewitson
Papilio zalmoxis Hewitson, 1864, Exot. Butt., III, Papilio, Pl. v1, fig. 18. Aurtvit-
Lius, 1908, Seitz, Gross-Schmett., XIII, p. 12, Pl. 1b.
There are five males, one caught at Avakubi, October 4, 1909, a
second taken at Gamangui’in February, two captured at Bafwasende
in September, and the fifth caught at Niangara in November 1910.
(423) 3. Papilio dardanus Brown
Papilio dardanus Brown, 1776, New Ill. Zool., p. 52, Pl. xxm. Avurtvitirus, 1909,
Seitz, Gross-Schmett., XIII, p. 13, Pl. ua, 3.
Papilio hippocoén F anrictvs, 1793, Ent. Syst., III, part 1, p.38, 9. Hewrrson, 1868,
Exot. Butt., IV, Papilio, Pl. xu, fig. 38. Avurtvitiius, 1908, Seitz, Gross-
Schmett., XIII, p. 13, Pl. mb.
Twenty-eight males of this form, most of them taken at Medje, a
few in April and a few in September, but most of them in July and
August; one is labelled “Stanleyville, August 23, 1909,’ another
“Munie Katoto, November 1909,” and a third is marked as caught at
“Niangara, November 1910.’’
One specimen of the female form hippocodn was taken at Niangara,
November 17, 1910.
(424) 4. Papilio cynorta Fabricius
Papilio cynorta Fasricros, 1793, Ent. Syst., III, part 1, p. 37. Westrwoop, 1843,
Arcana Ent., I, p. 15, Pl. x1, figs. 3, 4, o.
Papilio boisduvalianus Westwoon, 1843, Arcana Ent., I, p. 151, Pl. x1, figs. 1, 2.
There are twelve males and two females. One greatly dwarfed
male was caught at Niangara in November; all the rest are from Medje,
two males taken in April, the rest in August and September.
1920 Holland, Lepidoptera of the Congo 245
(425) 5. Papilio zenobia Fabricius
Papilio zenobia F asricivus, 1775, Syst. Ent., p. 503. AurtviLirus, 1908, Seitz, Gross-
Schmett., XIII, p. 16, Pl. mb.
Thirty-one specimens, including a couple of females. One male is
somewhat aberrant, having the pale bands yellowish, not white as is
usual. The specimens were mostly taken at Medje, a few in April,
the rest from July to September inclusive, although several are ticketted
as caught both at Medje and Niangara in November.
A number of the males have a small white spot at the end of the
cell of the fore wing, as is always the case with the females, and there-
fore belong to the so-called “variety,” to which Suffert has given the name
Papilio nobicea (Nomen vix conservandum).
(426) 6. Papilio mechowianus Dewitz
Papilio cypreafila var. mechowiana Drwitz, 1885, Ent. Nachr., XI, p. 305, fig...
AvRIVILLIUs, 1908, Seitz, Gross-Schmett., XIII, p. 16, Pl. ma, 9.
Papilio mechowianus AuRIVILLIUS, 1908, Seitz, Gross-Schmett., XIII, p. 16, Pl. ma.
There are four specimens, one caught at Medje in July, the others
at Niangara in November. They are all more or less defective, rubbed,
and lacking antenne.
(427) 7. Papilio gallienus Distant
Papilio gallienus Distant, 1879, Proc. Zool. Soc. London, p. 649.
There is a small army of specimens which I refer to this species,
although the males have the transverse light bands of the wings on the
upper side narrower than is shown in the figure given by Aurivillius (cf.
Seitz, XIII, Pl. 1). The females agree more nearly with the figure cited,
and indeed this figure may represent the female. The author has un-
fortunately failed to indicate in the text or on the plate what sex his
figure represents.
I count one hundred and twelve specimens of this species, mostly
taken at Medje, or near by, at Gamangui, Bafwasende, and Bafwaboli.
A few were caught in April, but the great majority were captured in
July, August, and September.
(428) 8. Pap'‘lio hesperus Westwood
Papilio hesperus Westwoop, 1843, Arcana Ent., I, p. 189, Pl. xtvut.
There are nine males taken at Medje from July to September.
(429) 9. Papilio nireus Linnzus
Papilio nireus Linnamus, 1758, Syst. Nat., 10th Ed., p. 464. Auriviniius, 1908,
Seitz, Gross-Schmett., XIII, Pl. va, vb.
246 Bulletin American Museum of Natural History (Vol. XLIII
This species is represented by sixty-six males, some of which by the
reduction in width of the blue transverse band on the upper side of the
wings approach in appearance the form lyeus Doubleday, which is
prevalent in southern and eastern Africa.
A couple of the specimens are labelled as taken at Gamangui in
February, a few as caught at Avakubi and Niangara in November, but
by far the greater number were collected at Medje, some in April, the
rest from June to September.
(430) 10. Papilio bromius Doubleday
Papilio bromius DouBLEpay, 1845, Ann. Mag. Nat. Hist., (1) XVI, p. 176. Aurivit-
Lius, 1908, Seitz, Gross-Schmett., XIII, Pl. vb, ve.
There are fifty-three males of this species in the collection. One
was taken at Matadi, the rest at Medje, several in April, the rest in the
mid-summer months.
(431) ; 1l. Papilio demodocus Esper
Papilio demodocus Esper, 1798, Auslindische Schmett., p. 205, Pl. x1, fig. 1.
This is the African race of P. demoleus Linnzus, which latter is
common in the Asiatic tropics and also occurs in Arabia. It principally
differs by being somewhat larger and darker and having the ocellus
at the anal angle of the hind wing less conspicuously marked with red
scales.
There are thirty-one males and six females. Twenty-seven of the
specimens were taken at Niangara in the month of November, five at
Medje in June and July, two at Stanleyville in August, one at Risimu
in September, and one at Ngayu in December.
One of the males taken at Niangara on November 9, 1910, is re-
markable because all the light spots on the upper side of the wings are
brilliant lemon-yellow, and not creamy white, as is usually the case.
One female belongs to the varietal form to which Capronnier gave the
name nubila (ef. Aurivillius, Seitz, loc. cit., Pl. 1va) in which the light
spots on the upper side of the wings are ochreous. Aurivillius suggests
that this so-called subspecies is based upon discolored (“verfirbte’’)
specimens. I find it, however, difficult to agree with him, inasmuch as
I have before me as I write, quite a long series of such specimens, coming
from many different localities from all over the continent, and all in
such state of preservation as to militate against the idea that the color
is other than natural. In every larger series of this species which has
come into my hands I have found a few specimens which have the pale
markings ochreous and, therefore, belong to the form nubila. Though
1920] Holland, Lepidoptera of the Congo 247
merely a color-variety, it occurs constantly, and I should say that it is
no exaggeration to affirm that at least two per cent of the specimens
collected belong to it.
(432) 12. Papilio menestheus lormieri Distant
Papilio menestheus Drury, 1773, Ill. Exot. Ent., II, p. 15, Pl. rx, figs. 1, 2.
Papilio menestheus var. lormieri Distant, 1874, Ent. Mo. Mag., XI, p. 129. AurtvIL-
Livus, 1908, Seitz, Gross-Schmett., XIII, p. 20, Pl. vin1b.
All of the specimens in the collection belong to the varietal form
described by Distant, which differs from typical P. menestheus in having
the transverse band of pale spots on the fore wing nearly straight, and
not curving inwardly near the costa in the direction of the base of the
wing. The typical form occurs in Sierra Leone and the northern parts
of the range of the insect, but the variety lormieri is most prevalent from
southern Cameroon and the valley of the Ogové eastward into the hot
wooded interior of the continent and reaches as far south as Mashona-
land. .
There are seventy-five specimens of this insect in the collection,
including a couple of females. Most of them were taken at Medje, a
few in April, the rest from June to September, but chiefly in the months
of July and August, A small number are labelled as having been taken
at Niangara in November.
(433) 13. Papilio ridleyanus White
Plate VI, Figure 3, typical @
Papilio ridleyanus Wuite, 1843, Ann. Mag. Nat. Hist., (1) XII, p. 262, fig.
AURIVILLIvs, 1908, Seitz, Gross-Schmett., XIII, p. 21, Pl. 1a.
Twenty-three male specimens. Of these nine are labelled as having
been taken at Gamangui in February, six at Medje in June and July,
three at Bafwasende in September, four at Niangara in November, and
one ‘“‘near Barumbu, July 31, 1909.”
(434) 13a. Papilio ridleyanus fumosus, new variety
Plate VI, Figure 4, °
P. ridleyanus is dimorphic. Two males from Medje and two from
Bafwasende belong to a variety in which the red color of both wings dis-
appears and is replaced by dull smoky gray. This is true also of a certain
proportion of the females of the species. I have before me as I write a
number of females taken by Antisdel in the Belgian Congo not far from
Leopoldville, some of which are colored like the typical males and are
248 Bulletin American Museum of Natural History [Vol. XLII
marked with bright red spots, but one of which is precisely like the
smoky-winged males from Medje and Bafwasende. For this form I
propose the subspecific name given above.
Type & from Bafwasende in The American Museum of Natural
History; allotype 2 in Carnegie Museum (Coll. Antisdel). Paratyvpes
in The American Museum of Natural History and Holland Collection in
the Carnegie Museum.
(435) 14. Papilio pylades Fabricius
Papilio pylades Fasricius, 1793, Ent. Syst., IIT, part 1, p. 34. AVELYPLIUB, 1908,
Seitz, Gross-Schmett., XIII, p. 21, Pl. vite.
There are seventy-eight specimens, a'most all of which are males.
They were mostly taken at Niangara in Hovember. Six are labelled as
from Faradje, one of which bears date as of November, and the rest are
without designation of day or month. Only one specimen is recorded as
from Medje, and is dated August 1, 1910.
There is not much to be remarked about the specimens, except that
they run very small in size, which suggests that they represent the wet-
season form.
(436) 15. Papilio angolanus Goeze
Papilio angolanus Gaze, 1779, Ent. Beitr., III, part 1, p. 87. Aurtvriirius, 1908,
Seitz, Gross-Schmett., XIII, p. 21, Pl. vib.
A solitary male, taken at Matadi, June 1909, represents this species.
(437) 16. Papilio tyndareus Fabricius
Papilio tyndareus Fasricius, 1793, Ent. Syst., III, part 1, p.35. Aurtviiiius, 1908,
Seitz, Gross-Schmett., XIII, p. 22, Pl. via.
There are nine males, one of which is discolored. One was taken at
Bafwabaka, January 7; two at Gamangui in February; five at Medje,
two of these in April, the other three one each in June, August, and
September; one was captured at Niangara in November.
(438) 17. Papilio theorini Aurivillius
Papilio theorint Aurnivitiius, 1881, Ent. Tidskr., II, p. 45; 1908, Seitz, Gross-
Schmett., XIII, p. 21, Pl. md.
Represented by four males captured at Medje, two in April, one in
July, and one in August 1910.
(439) 18. Papilio ucalegonides Staudinger
Papilio ucalegonides StauvincER, 1884, Exot. Schmett., I, p. 10, Pl. v (as P. ucalegon).
Papilio ucalegonides Avwivit.ivs, 1908, Seitz, Gross-Schmett., XIII, p. 23.
Two males taken at Medje in September.
1920] Holland, Lepidoptera of the Congo 249
(440) 19. Papilio charcedonius Karsch
Papilio charcedonius Karscu, 1895, Ent. Nachr., X XI, p. 285. Aurrvitirus, 1908,
Seitz, Gross-Schmett., XIII, p. 24, Pl. rxe.
Two males, one caught at Medje in August, the other at Niangara
in November.
(441) 20. Papilio leonidas Fabricius
Papilio leonidas Fasricius, 1793, Ent. Syst., III, part 1, p. 35. Aurrvitirus, 1908,
Seitz, Gross-Schmett., XIII, p. 22, Pl. vid.
Eleven specimens caught at Medje, two in April: the rest from July
to September; nine captured at Niangara in November; one at Risimu,
September 8, 1909; one at Avakubi in November; and two at Faradje
in December.
(442) 21. Papilio antheus Cramer
Papilio antheus Cramer, 1779, Pap. Exot., III, p. 72, Pl. ccxxxtv, figs. B,C. Auri-
VILLIUS, 1908, Seitz, Gross-Schmett., XIII, p. 25, Pl. rxa.
There are thirty-seven specimens, one caught at Medje in April,
two taken at the same place in July, and all the rest captured at Gaman-
gui in February.
(443) 22. Papilio policenes Cramer
Papilio policenes CRAMER, 1775, Pap. Exot., I, p. 61, Pl. xxxvn, figs. A, B. Aurtvit-
Lius, 1908, Seitz, Gross-Schmett., XIII, p. 25.
There are one hundred and six examples of this common species in
the collection. Most of them were taken at Gamangui in February,
but a score or more are labelled as from Medje, a few taken in April,
the rest in the mid-summer months. A few were caught at Niangara
in November, and one is labelled ‘‘ Ngayu, December.’
(444) 23. Papilio illyris Hewitson
Papilio illyris Hewitson, 1873, Ent. Mo. Mag., LX, p. 232. Aurtvitiius, 1908,
Seitz, Gross-Schmett., XTIT, p. 26, Pl. virta.
A single male taken at Medje in the first week of September.
Hesperiidez
In view of the large number of species, nearly three hundred, belong-
ing to the family Hesperiide, which are already known to occur in the
region of the Congo, the collection brought home by the Lang-Chapin
Expedition is somewhat disappointing. Like the collection of the
Lyczenide reported upon on preceding pages, it shows that the native
collectors who were employed to gather insects concentrated their
250 Bulletin American Museum of Natural History [Vol. XLIII
attention upon the larger and showier species in other families. In com-
parison with the really splendid collection of Nymphalide, the Hesperiidze
make a rather poor showing. Nevertheless, some excellent things turn
up, as for instance a specimen of Procampta rara Holland, hitherto
only known by the type, which until now was unique, and two new
species, Abantis rubra and Leptalina niangarensis. The collection con-
tains ninety-four specimens of the Hesperiide, representing thirty-
seven species, belonging to twenty-three genera. Many species are re-
presented by but one specimen.
Hesperiinz
Sarancesa Moore
Subgenus Hypa Mabille
(445) 1. Sarangesa (Hyda) grisea (Hewitson)
Pterygospidea grisea Hewitson, 1878, Ann. Mag. Nat. Hist., (5) I, p. 344.
Hyda micacea MABILLE, 1889, Ann. Soc. Ent. France, (6) IX, Bull., p. elxviii.
MABILLE AND VUILLO1, 1893, Novit. Lep., p. 93, Pl. x1, fig. 3.
Hyda grisea HOLLAND, 1896, Proc. Zool. Soc. London, p. 6. Masriie, 1904, Gen.
Ins., XVII, p. 55.
The species is represented by a single male specimen captured at
Medje, July 19, 1910.
Subgenus Sarge Mabille
(446) 2. Sarangesa (Sape) maculata (Mabille)
Sape maculata Masitxe, 1891, C. R. Soc. Ent. Belgique, p. lxviii. Hottanp, 1896,
Proc. Zool. Soc. London, p. 10. Masriie, 1904, Gen. Ins., XVII, p. 55.
There are two somewhat worn male specimens from Niangara
which agree so closely with a series of finely preserved examples collected
for me at Mombasa by Doherty and which are undoubtedly referable
to S. maculata Mabille, that I do not hesitate to place them here. In
1896 at the time I prepared my ‘Synonymic Catalogue of the Hesperiide
of Africa’ I only knew S. maculata by the descfiption published by
Mabille. The species is well defined and thoroughly valid, as shown by
the material before me.
CeL@norrRuinvus Hiibner
(447) 1. Celenorrhinus galenus (Iabricius)
Hesperia galenus Fannicius, 1793, Ent. Syst., LIT, part 1, p, 350.
Hesperia galena Donovan, 1800, Ins. India, Pl. 1, fig. 3, 9.
Plesione uragalenus STAUDINGER, 1888, Exot, Schmett., part 1, p. 300; part 2, Abbild.,
Pl. c, o’.
1920] Holland, Lepidoptera of the Congo 251
Celenorrhinus galenus HOLLAND, 1896, Proc. Zool. Soc. London, p. 12. MAsBrixe,
1904, Gen. Ins., XVII, p. 51.
The species is represented in the collection by four males and two
females, all of which were taken at Medje, the dates of capture ranging
from April to August 1910.
(448) 2. Celewnorrhinus rutilans (Mabille)
Pardaleodes rutilans MaBILLe, 1877, Bull. Soc. Zool. France, p. 235, 9. MABILLE
AND VUILLOT, 1893, Novit. Lep., p. 96, Pl. x11, fig. 7, o.
Pterygospidea tergemira Hewitson, 1877, Ann. Mag. Nat. Hist., (4) XX, p. 323.
Tagiades wermanni Patz, 1879, Stett. Ent. Zeit., XL, p. 362, 2.
Celenorrhinus rutilans HOLLAND, 1896, Proc. Zool. Soc. London, p. 12. MABILe,
1904, Gen. Ins., XVII, p. 51.
Four males and one female of this species were taken at Medje in
the months of July and August 1910.
(449) 3. Celenorrhinus boadicea (Hewitson)
Pterygospidea boadicea Hewirson, 1877, Ann. Mag. Nat. Hist., (4) XX, p. 323.
Celenorrhinus atratus HOLLAND, (part), 1894, Ent. News, PI. m1, fig. 5.
Celenorrhinus boadicea HOLLAND, 1896,.Proc. Zool. Soc. London, p. 14, Pl. m1, fig. 1.
Masirixe, 1904, Gen. Ins., XVII, p. 51.
One male specimen was caught at Medje in August 1910,
(450) 3. Celenorrhinus chrysoglossus (Mabille)
Ancistrocampta chrysoglossa M aBIL.E, 1891, C. R. Soc. Ent. Belgique, XX XV, p. evii.
Celenorrhinus chrysoglossa HOLLAND, 1896, Proc. Zool. Soc. London, p. 14, Pl. m1,
fig. 5. Mapriie, 1904, Gen. Ins., XVII, p. 51.
_ There is a male in somewhat damaged condition caught at Medje,
April 6, 1910.
TaaciapDEs Hiibner
(451) 1. Tagiades flesus (Fabricius)
Papilio flesus Fasrictus, 1771, Spec. Ins., II, p. 135, No. 621.
Papilio ophion Drury, 1782, Ill. Exot. Ent., III, Pl. xvu, figs. 1, 2.
Nisoniades flesus BUTLER, 1869, Cat. Fabr. Diurn. Lep., p. 286.
Nisoniades ophion TRIMEN, 1866, Rhop. Afr. Austr., IT, p. 313.
Tagiades flesus WATSON, 1893, Proc. Zool. Soc. London, p. 54. HoLianp, 1896, Proc.
Zool. Soc. London, p. 16. Masruye, 1904, Gen. Ins., XVII, p. 53.
There are two males and four females of this well-known and widely
distributed species; the males taken at Niangara in November; two
females at Medje, one in July, the other in August; a third female at
Niangara in November 1910; and a fourth at Stanleyville, in March 1915.
252 Bulletin American Museum of Natural History [Vol. XLIII
Eacris Guenée
(452) 1. Eagris lucetia (Hewitson)
Leucochitonea lucetia Hewitson, 1875, Ill. Exot. Butt., V, July, Leucochitonea, Pl. u,
fig. 21.
Biigrts tsi HOo.uanp, 1896, Proc. Zool. Soc. London, p. 19. Masruue, 1904, Gen.
Ins., XVII, p. 54. ~
There are four males of this well-defined species, all taken at Medje,
one in April, two in June, and one in August 1910. Hewitson originally
described the insect from Angola.
Procampta! Holland
(453) 1. Procampta rara Holland
Procampta rara Houuanp, 1892, Ann. Mag. Nat. Hist., (6) X, p. 298. Watson,
1893, Proc. Zool. Soc. London, p. 59. Hoxuanp, 1894, Ent. News, Pl. 1m, fig. 7;
1896, Proc. Zool. Soc. London, p. 19.
There is a single specimen of this scarce insect, which is labelled as
taken at Medje, August 24, 1910. It has been compared with the type,
and agrees with it in every particular. So far as I know, it is the second
specimen which has thus far turned up. The type came from the valley
of the Ogové River. The butterfly seems to have been overlooked by
collectors, probably because of its small size and obscure coloration.
Caprona Wallengren
(454) 1. Caprona pillaana Wallengren
Caprona pillaana W ALLENGREN, 1857, Kongl. Sv. Vet.-Akad. Handi., N.S., II, No. 4,
p. 51.
Stethotrix heterogyna Maxni.ie, 1889, Ann, Soc. Ent. France, (6) LX, Bull., p. clxxxiv.
Caprona adelica Karscu, 1892, Ent. Nachr., XVIII, p. 242; 1893, Berl. Ent. Zeit.,
XXXVIII, p. 243, Pl. v1, fig. 2.
Caprona pillaana HouLaNnn, 1896, Proc. Zool. Soc. London, p. 19. Mastin, 1904,
Gen. Ins., XVII, p. 76.
There is one ragged male specimen of this species which, according to
the label, was taken at “Faradje 1911-1912.”
'Mabille in the ‘Genera Insectorum,’ Fase. XVII, in the Index gives the genus Procampta Holland
and refers top. 59''; he also cites in the Index the specific name rara “sp. gen. Pree. and
refers top. 131, Neither of these pages contains the slightest allusion either to the genus or the species.
On page 43, in one of the keys to the genera he cites Procampta as belonging to Section 11, Iam curious
to know how the Index of the work was fabricated, so as to cite the genus and species and the es on
which they are recorded, without their being there at all, I may say in passing that while I have the
warmest personal regard for the author of thiswork, which must be constantly referred to, it so abounds
in ty utraphical errors and omissions ae to make its use peepecinaly trying. It is the ‘despair’ of
working hesperidologists. So far as the Hesperiidw of temperate and boreal North America are con~-
cerned, it complete y senores every thing of importance which had been done by students in the United
States and Canada during the quarter of a century which had elapsed prior to its appearance,
1920] Holland, Lepidoptera of the Congo 253
Asantis! Hopffer
(455) 1. Abantis elegantula (Mabille)
Sapea elegantula MasiiiE, 1890, Ann. Soc. Ent. France, (6) X, p.32. MasILie
AND VUILLOT, 1891, Novit. Lep., p. 23, Pl. m1, fig. 6.
This species is represented by a solitary male taken at Medje,
March 8, 1910.
(456) 2. Abantis efulensis Holland
Abantis efulensis HOLLAND, 1896, Proc. Zool. Soc. London, p. 21, Pl. v, fig. 12.
There is a single male specimen from Medje which differs from the
type, with which it has been compared, only in the fact that the outer
border of the hind wings on the upper side is clouded with dark scales to
a greater extent than is the case in the individual which I originally
described and figured. In this respect, however, it does not differ from
other individuals collected since my first description was published.
(547) 3. Abantis rubra, new species
Plate XII, Figure 1, #7
3. Frons and inner margins of palpi reddish orange; outer margins of palpi deep
black with a white spot at the base of each just below the eye. Thorax and patagia
dark brown, the patagia, so far as the specimen reveals, without light markings (the
patagia are in bad case). Abdomen dark brown, with a black median longitudinal
line above, and the edges of the segments on the sides adorned with pale reddish
vertical spots; lower side of abdomen paler brown than above. The primaries on the
upper side are pale semitranslucent fuscous, with all the veins dark brown and clearly
defined against the paler ground-color. There is a small subcircular blackish spot at
the base, defined outwardly by a few orange-red scales. The ground-color of the
secondaries on the upper side is orange-red, the veins are black and clearly defined
against the ground-color. At the end of the cell about the middle of the wing is a
sharply defined velvety black spot, from which through the middle of the cell there
runs to the base a narrow black ray of the same color as the spot at the end of the cell.
The outer margin of the secondaries on the upper side is bordered by fuscous, defined
inwardly by a narrow band of darker brown which widens a little towards the anal
angle. The inner margin is shaded with fuscous. On the under side both wings are
pale reddish inclining to fuscous, with all the markings of the upper side reproduced,
but in paler tints than on the upper side. Expanse, 33 mm.
1Fairmaire in the Annals of the Entomological Society of Belgium, XXXVIII, 1894, p. 395,
announces that he has been informed by his learned colleague, Dr. Berg, that the generic name Abantis
Hopffer should be replaced by the generic name Abantiades Herrich-Scheeffer, and states that he hastens
to make announcement of the fact.. Unfortunately for this supposed correction I discover that the
generic name Abantiades was proposed by Herrich-Scheeffer for one of the large Australian Hepialids,
and he makes Epiolus (Pielus Walker) argenteus Donovan the type of the genus. As the Hepialide
and the Hesperiide are rather far apart in any system of classification, which has thus far been pro-
osed (about as far apart as tigers and squirrels among mammials), the correction provokes a smile.
he name Abantis Hopffer stands in spite of the Choowery of Doctors Berg and Fairmaire. The t
of the genus is A. tettensis Hopffer, as pointed out by Scudder (Proc. Amer. Acad. Arts and Sciences, X,
1875, p. 99).
254 Bulletin American Museum of Natural History [Vol. XLII
The type, which is in The American Museum of Natural History, is
unique. It was taken at Medje, April 6, 1910.
The genus Abantis is thus far known to include eighteen species,
all of which are found in the Aithiopian subregion. .They are the
following:
1. Abantis tettensis Hoprrer, 1855, Monatsber. Akad. Wiss. Berlin, p. 643; 1862,
Peters, Reise n. Mossambique, Ins., p. 415, Pl. xxv1, figs. 16, 17.
Eastern tropical Africa.
2. Abantis paradisea (BUTLER), 1870, Trans. Ent. Soc. London, p. 499; 1874, Lep.
Exot., p. 167, Pl. urx, fig. 8.
Abantis namaquaana (WEsTwoop), 1874, Thes. Ent. Oxon., p. 183, Pl.
xxxIv, fig. 10, 9.
Southern Africa.
3. Abantis zambesiaca (WEstwoop), 1874, Thes. Ent. Oxon., p. 183, Pl. xxxrv, fig. 9.
Abantis zambezina TRIMEN, 1889, S. Afr. Butt., III, p. 344.
Abantis trimeni (Bu1LER), 1895, Proc. Zool. Soc. London, p. 264, Pl. xv,
fig. 5.
Southern tropical Africa.
4. Abantis bismarcki Karscu, 1892, Ent. Nachr., XVIII, p. 228; 1893, Berl. Ent.
Zeit., XX XVIII, p. 242, Pl. v1, fig. t.
Topsland.
5. Abantis bicolor (TRImEN), 1864, Trans. Ent. Soc. London, (3) I, p. 180; 1866,
Rhop. Afr. Austr., II, p. 307, Pl. vy, fig. 1.
South Africa,
6. Abantis venosa TRIMEN, 1889, 8. Afr. Butt., III, p. 339; 1891, Proc. Zool. Soe.
London, p. 105, Pl. rx, fig. 24.
Abantis umvulensis (E. Sharpe), 1890, Ann. Mag. Nat. Hist., (6) VI,
p. 348.
Southern tropical Africa, Transvaal.
7. Abantis elegantula (Mapiuie), 1890, Ann. Soc. Ent. France, (6) X, p. 32.
MABILLE AND VuILLoT, 1891, Novit. Lep., p. 24, Pl. m1, fig. 6.
Sierra Leone, Cameroon, Belgian Congo.
8. Abantis efulensis HOLLAND, 1896, Proc. Zool. Soc. London, p. 21, Pl. v, fig. 12.
Cameroon, Belgian Congo.
9. Abantis plerotica Karscu, 1896, Ent. Nachr., XXII, p. 374.
German East Africa.
10. Abantis leucogaster (MaBiLLE), 1890, Ann. Soc. Ent. France, (6) X, p. 32.
MABILLE AND VurLiorT, 1891, Novit. Lep., p. 22, Pl. mt, fig. 5.
Sierra Leone.
11. Abantis nigeriana Burier, 1901, Ann. Mag. Nat. Hist., (6) VIII, p. 59.
Nigeria.
12. Abantis arctomarginata Laruy, 1901, Trans. Ent. ok. London, p. 34, Pl.
fig. 7.
Zomba, British Central Africa.
13. Abantis ja H. H. Druce, 1909, Proc. Zool. Soc. London, p. 408, Pl. uxvn, fig. z.
Interior of Cameroon,
1920] Holland, Lepidoptera of the Congo 255
14. Abantis lucretia H. H. Deuce, 1909, Proc. Zool. Soc. London, p. 409, Pl. uxvu,
fig. 3.
Interior of Cameroon.
15. Abantis lofu Neave, 1910, Proc. Zool. Soc. London, p. 72, Pl. 1m, figs. 4, 5.
Northern Rhodesia.
16. Abantis leeubu (WALLENGREN), 1857, Kongl. Sv. Vet.-Akad. Hand:., N. 8., I,
No. 4, p.-72. ,
17. Abantis amneris Reset AND RoGEeNnHorer, 1894, in O. Baumann’s ‘Durch
Masailand zur Niiquelle,’ p. 338.
Umbugwe.
18. Abantis rubra HOLLAND, new species.
Belgian Congo.
HesPerra Fabricius
(458) 1. Hesperia dromus (Plcetz)
Pyrgus dromus Puiarz, 1884, Mitth. nat. Ver. Neu-Vorpomm. u. Riig., p. 6.
TrimeEn, 1889, 8. Afr. Butt., III, p. 283.
Hesperia dromus Watson, 1893, Proc. Zool. Soc. London, p. 65. Hoxianp, 1896,
Proc. Zool. Soc. London, p. 23. MasriLue, 1904, Gen. Ins., XVII, p. 81.
Five specimens, three males and two females, were collected. Two
were captured at Medje in the first week of April; one was taken at
Faradje, December 5, 1912; the other two specimens were captured at
Niangara in the second week of November 1910.
(459) 2. Hesperia pletzi Aurivillius
Syrichtus spio PLasrz, 1884, Mitth. nat. Ver. Neu-Vorpomm. u. Riig., p. 21.
Pyrgus spio MaBIL_Z, 1890, Ann. Soe. Ent. France, (6) X, p. 30.
Hesperia pletzi Aurtvitiius, 1891, Ent. Tidskr., p. 227. Karscu, 1893. Berl. Ent.
Zeit., XXVIII, p. 245. Hotuanp, 1896, Proc. Zool. Soc. London, p. 23.
Masixg, 1904, Gen. Ins., XVII, p. 81.
There are two specimens in the collection, one taken at Gamangui,
February 5, the other at Medje, April 6, 1910.
Pamphiline
Actgeros Mabille
(460) 1. Acleros pletzi Mabille
Apaustus leucopygus PLasrz, 1879, Stett. Ent. Zeit., XL, p. 360.
Acleros pletzi MABILLE, 1889, Ann. Soc. Ent. France, (6) IX, Bull., p. elxxiii. Hot-
LAND, 1896, Proc. Zool. Soc. London, p. 29, Pl. 1, fig. 7. Masriue, 1904, Gen.
Ins., XVII, p. 96.
This rather common species is represented in the collection by one
tattered male taken at Medje, June 26, 1910.
256 Bulletin American Museum of Natural History (Vol. XLIII
Osmopes Watson
(461) 1. Osmodes laronia (Hewitson)
Hesperia laronia Hewitson, 1868, Descript. Hesper., p. 35.
Plastingia laronia Patz, 1879, Stett. Ent. Zeit., XL, p. 356; 1884, idem, XLV,
pp. 145-146.
Osmodes laronia Watson, 1893, Proc. Zool. Soc. London, p. 78. HoLianp, 1896,
Proc. Zool. Soc. London, p. 40, Pl. rv, figs. 3, 5. Masinin, 1904, Gen. Ins.,
XVII, p. 102.
Represented by a male and female captured at Medje in August
1910.
Hypotevcis Mabille
(462) 1. Hypoleucis cretacea (Snellen) (?)
Goniloba cretacea SNELLEN, 1872, Tijd. vi Ent., p. 27, Pl. 11, figs. 4-6.
Hesperia gonessa Hewrtson, 1877, Ann. Mag. Nat. Hist., (4) XIX, p. 76. Heron,
1898, Ann. Mag. Nat. Hist., (7) I, p. 256.
Hypoleucis cretacea HOLLAND, 1896, Proc. Zool. Soc. London, p. 47 (where further
synonymy is given).
There is a single somewhat damaged female which I refer with
caution to this species, from typical examples of which it apparently
differs only by a reduction in size of the translucent white spots of the
primaries. The specimen was captured at Medje, July 17, 1910.
Leptatina Mabille
M. P. Mabille, in his Revision of the Family _Hesperiide in the
‘Genera Insectorum,’ published by P. Wytsman, has erected the genus
Leptalina for the reception of Cyclopides unicolor Bremer and Gray,
which is found in China and Japan, and the three African species,
lepeletieri (Latreille), tsita (Trimen), and inornatus (Trimen), which hith-
erto have been commonly regarded as belonging to the genus Cyclopides.
I am quite familiar with the Asiatic species, having personally collected
it in quantity when engaged in research work in Japan in 1887. I am
somewhat dubious as to the propriety of removing it from the genus
Cyclopides. There seems to me to be more propriety in separating the
three African forms from their former association. I propose to add to
the genus Leptalina Mabille another species the description of which
is given herewith.
(463) 1. Leptalina niangarensis, new species
Plate XII, Figure 3,
@. Palpidark fuscous, upper side of thorax and abdomen blackish, lighter on the
under side, inclining to ochraceous, the terminal segments on their posterior edges
marked laterally with pale scales, Both fore and hind wings on the upper side uni-
1920] Holland, Lepidoptera of the Congo 257
formly dark brown, without any traces of translucent spots or other maculation.
Cilia concolorous. On the under side both wings are colored as on the upper side,
but are slightly paler in their ground-color, and near the base are clothed with
ochraceous scales. The hind wings on this side reveal a dark subcircular and some-
what diffuse spot in the cell at its extremity and this is followed externally by a post-
median dark band, which corresponds in its curvature to the outer margin. Expanse,
32 mm.
Two specimens were taken by the Lang-Chapin Expedition at
Niangara in November 1910. One of these, which I designate as the
type, is in The American Museum of Natural History; the other,
which I designate as the paratype, I have the pleasure of retaining for
the Holland Collection in the Carnegie Museum.
I append a brief key for the determination of the four species of the
genus Leptalina now known to occur in Africa.
A. Hind wings marked on the under side by light longitudinal lines.
1. Hind wings marked by two conspicuous silvery white lines, ranning through
the cell from the base to the outer margin, the upper line broader and
SGN CONSTICUGUS. 6. iis de a ees lepeletieri (Latreille).
,2. Hind wings having the nervules, especially in the outer half of the wing
covered with white scales, causing the veins to stand out clearly against
ee carker ground-colof. 4: {06 6s i eee ee ae tsita (Trimen).
B. Hind wings not marked on the under side by light lines, running from the base
toward the outer margin.
1. Fore wings with three minute translucent subapical spots..inornatus (Trimen).
2. Fore wings without translucent spots; hind wings on the under side with an
obscure dark subcircular spot at the end of the cell, followed by a cloudy
dark postmedian transverse band ............. niangarensis Holland.
CuaPpra Moore
(464) 1. Chapra mathias (Fabricius)
Hesperia mathias Fasricius, 1798, Ent. Syst., Suppl., p. 436.
(For synonymy cf. Holland, 1896, Proc. Zool. Soc. London, p. 60.)
One male caught at Boma, June 15, 1915.
ParnakaA Moore
(465) 1. Parnara borbonica (Boisduval)
Hesperia borbonica Botspuvat, 1833, Faune Ent. Madgr., p. 65, Pl. rx, figs. 5, 6.
(For additional synonymy cf. Holland, 1896, Proc. Zool. Soc. London, p. 62.)
There are two specimens of this common insect in the collection,
one taken at Faradje, December 6, 1912, and another at the same place,
labelled ‘ 1911-1912.”
258 Bulletin American Museum of Natural History [Vol. XLIII
(466) 2. Parnara alberti (Holland)
Baoris alberti HOLLAND, 1896, Proc. Zool. Soc. London, p. 67, Pl. m, fig. 21.
Parnara alberti MaBiLx, 1904, Gen. Ins., XVII, p. 136.
Of this species there are three specimens, one taken at Medje, June
27, 1910; one at Matadi, August 24, 1910; and one at Faradje, labelled
“1911-1912.” They all agree with specimens taken on the Ogové
River and in southern Cameroon. .
M. Mabille has transferred this species and several others which
I classified under Baoris Moore to the genus Parnara Moore. The
distinction between the two genera, as is pointed out by Mabille, is
rather fine, based principally upon the length of the antenne and the
sexual brands, which do not always afford good grounds for generic
separation. I follow Mabille provisionally, though feeling that the whole
group needs a more careful revision than it has thus far received and
that a systematic examination of the vast number of species now lumped
under the name Parnara will disclose that the group, as made up, con-
tains a good many incongruous elements.
Sematea' Holland
(467) 1. Semalea pulvina (Ploetz)
Hesperia pulvina Piaerz, 1879, Stett. Ent. Zeit., XL, p. 353.
Cobalus carbo MaBILLE, 1889, Ann. Soc. Ent. France, (6) IX, Bull., p. elxix.
Trichosemeia pulvina Watson, 1893, Proc. Zool. Soc. London, p. 53.
Semalea pulvina HoLuanp, 1896, Proc. Zool. Soc. London, p. 65, fig. and Pl. 1, fig. 14.
MaBixez, 1904, Gen. Ins., XVII, p. 138.
One male specimen caught at Medje, August 24, 1910.
PuaTyLescHes Holland
(468) 1. Platylesches batange Holland
Parnara batanga HOLLAND, 1894, Ent. News, V, p. 92, Pl. 11, fig. 10.
Platylesches batanga Hou.anpd, 1896, Proce. Zool. Soc. London, p. 73. MABrILLE,
1904, Gen. Ins., XVII, p. 138. ,
There is one dwarfed and also somewhat damaged specimen of this
species which I have carefully compared with the type and find to be
identical. It is ticketted as taken at ‘ Basoko, July 3, 1909.”
'The genus Semalea Holland is omitted from the Index in Mabille’s work, loc, cit., but occurs on
p. 138, with notes and comments thereon by M, Mabille.
1920] Holland, Lepidoptera of the Congo 259
PARDALEODES Butler
(469) 1. Pardaleodes edipus (Cramer)
Papilio edipus CraMER, 1782, Pap. Exot., IV, p. 146, Pl. cccixv1, figs. E, F, 3.
Pamphila (?) sator DoUBLEDAY AND swine: 1852, Gen. Diurn. Lep., II, p. 523,
Pl. uxxrx, fig. 4, 9.
(For further synonymy cf. Holland, 1896, Proc. Zool. Soc. London, p. 75.)
There are three males and one female referable to this species. All
of them were taken at Medje in the months of July and August 1910.
(470) 2. Pardaleodes incerta (Snellen)
Pamphila incerta SNELLEN, 1872, Tijd. v. Ent., p. 29, Pl. x, figs. 10-12, #.
Hesperia coanza Piarz, 1883, Stett. Ent. Zeit., XLIV, p. 232, @.
Pardaleodes incerta HOLLAND, 1896, Proc. Zool. Soc. London, p. 75.
Four males; three taken at Medje, two in July, the other in kugabes
the fourth specimen caught at Niangara, November 26, 1910.
(471) 3. Pardaleodes bule Holland
Pardaleodes bule HoLuanp, 1896, Proc. Zool. Soc. London, p. 76, Pl. m1, fig. 21.
Two specimens from Medje, one collected in April, the other in
August 1910.
(472) 4. Pardaleodes fan (Holland)
Osmodes (?) fan Hotuanp, 1894, Ent. News, V, p. 91, Pl. m1, fig. 8.
Pardaleodes fan HoLtuann, 1896, Proc. Zool. Soc. London, p. 79.
Two specimens caught at Medje, August 1910.
CERATRICHIA Butler
(473) 1. Ceratrichia ialemia H. H. Druce
Ceratrichia ialemia H. H. Druce, 1909, Proc. Zool. Soc. London, p. 413, PI. jecainbe
fig. 10.
One male taken at Medje, August 13, 1910, which agrees wibiay
with the description and figure given by Druce. The species is very
near to C. nothus (Fabricius) and is probably only a form of that.
(474) 2. Ceratrichia phocion (Fabricius)
Papilio phocion Fasricius, 1781, Spec. Ins., IT, p. 138.
Ceratrichia phocion ButiER, 1870, Cat. Fabr. Diurn. Lep., p. 274, Pl. mz, fig. 14.
HOo.uanp, 1896, Proc. Zool. Soc. London, p. 79.
Apaustus argyrosticta PLatz, 1879, Stett. Ent. Zeit., XL, p. 358, 9.
Four males and one female of this common insect, all taken at Medje
from July to September 1910. The females are variable in the number
and size of the small light markings on the wings, as are also the males to
a less extent. There are before me as I write-several hundreds of speci-
260 Bulletin American Museum of Natural History [Vol. XLII
mens of the species, representing both sexes, some pairs taken in copula,
and a quite considerable series of females from the Ogové and southern
Cameroon, which conform absolutely to the description of argyrosticta
given by Pletz. I am now convinced that I was in error in 1896 in
regarding C. (Apaustus) argyrosticta as the female of the succeeding
species, C. flava Hewitson = C-charita (Ploetz).
(475) 3. Ceratrichia flava Hewitson
Ceratrichia flava Hewitson, 1878, Ann. Mag. Nat. Hist., (5) I, p. 343. Hoxanp,
1896, Proc. Zool. Soe. London, p. 79, Pl. m1, fig. 14.
Plastingia charita PLetz, 1879, Stett. Ent. Zeit., XL, p. 356.
One male taken at Batama, September 18, 1909.
(476) 4. Ceratrichia wollastoni Heron
Ceratrichia wollastoni Heron, 1909, Trans. Zool. Soc. London, XIX, p. 174, Pl. v,
figs. 13, 14.
There are three specimens referable to this species which were
taken at Medje, two in April, and one in September 1910. It is distin-
guished from C. flava Hewitson by the outline of the inner edge of the
terminal black band of the primaries, which curves regularly from the
inner margin before the lower angle to about the middle of the costa,
whereas in C. flava it is angulated at vein 4, running inwardly for some
distance along that vein towards the cell, which in most specimens it
reaches and the outer extremity of which it covers somewhat diffusely.
The insect was originally described from Ruwenzori; we also have speci-
mens from the interior of Cameroon.
Before dismissing the consideration of the few species of this genus
brought before us by the present collection, it is proper to state that it
seems to the writer that it is much in need of a careful revision, inasmuch
as the material constantly accumulating in his custody seems to indi-
cate that some at least of the so-called species are of questionable
validity.
Acromecis Mabille
Mabille, in the ‘Genera Insectorum,’ separates the species named
neander Plcetz from the genus Andronymus Holland, to which I assigned
it in 1896, and erects the genus Acromecis for its reception.
(477) 1. Acromecis neander (Plcoetz)
Apaustus neander Piarrz, 1884, Stett. Ent. Zeit., XLV, p. 154,
Ancylorypha producta Trimen, 1889, 8. Afr. Butt., IIT, p. 334,
Andronymus neander HOLLAND, 1896, Proc. Zool. Soc, London, p. 82, Pl. u, fig. 23.
Acromecia neander Manitve, 1904, Gen. Ins., XVII, p. 172. «
TE a TT ES ST ITA a
1920] Holland, Lepidoptera of the Congo 261
There are ten specimens, eight males and two females. All were
taken at Medje, except one female which is ticketted “Stanleyville, I,
27, 1915.”’ Six of the specimens taken at Medje were captured in June,
one in May, and one in September.
Czanives Holland
(478) 1. Cenides cenira (Hewitson)
Hesperia cenira Hewrison, 1867, Exot. Butt., 1V, Hesperia, Pl. 11, figs. 15, 16.
Pamphila cenira Kirsy, 1871, Syn. Cat., p. 606.
Hesperia calpis Puarz, 1879, Stett. Ent. Zeit., XL, p. 354; 1882, idem, XLIII, p.
328, 9.
Hidari cenira HOLLAND, 1896, Proc. Zool. Soc. London, p. 82, Pl. u, fig. 3, 2 (calpis
Pleetz).
Cenides cenira (sic) MaBiuie, 1904, Gen. Ins., XVII, p. 182.
One typical male collected at Medje, July 8, 1910.
In my catalogue of the African Hesperiide, 1896, I referred the fore-
going and two other species to the genus Hidariz Distant, with which they
seemed to agree better than with any other genus at that time described.
M. Mabille in his later work relegates them to my genus Cenides, of
which he says they form a section. I cannot take the time at the
moment of writing this to go fully into the matter, which might involve
some detailed investigation of structures under the microscope and
bleaching of wings, but I know that the Hesperia cenira of Hewitson is
certainly very doubtfully congeneric with Hesperia dacela Hewitson,
which is the type of the genus Cenides Holland.
(479) 2. Cenides cylinda (Hewitson)
Hesperia cylinda Hewirson, 1876, Ann. Mag. Nat. Hist., (4) XVIII, p. 449.
Pamphila calpis Karscu (nec Ploetz), 1893, Berl. Ent. Zeit., XX XVIII, p. 252, Pl.
vi, fig. 4 (9 non ).
Cenides cylinda HoLuaNnpd, 1896, Proc. Zool. Soc. London, p. 89, Pl. 1, fig. 12 (9
non 3). MaBILxe, 1904, Gen. Ins., XVII, p. 182.
Two moles, one taken at Leopoldville, July 4, 1909, the other at
Niangara, November 20, 1910.
The figures of this insect given both by Karsch and myself do not
represent the male but the female, which has a postmedian translucent
spot on the hind wing, while the male is destitute of such a spot, as is
shown by numerous specimens which have come into my possession or
custody since 1896.
262 Bulletin American Museum of Natural History (Vol. XLIIT
RHopatocamptTa Wallengren
(480) 1. Rhopalocampta bixe (Linnzeus)
Papilio bixe Linnzxus, 1758, Syst. Nat., 10th Ed., p. 485. Cierckx, 1764, Icones,
Pl. xuu, fig. 4..
Ismene bixe Kirpy, 1871, Syn. Cat., p. 582 (part). :
Rhopalocampta bixe Watson, 1893, Proc. Zool. Soc. London, p. 129. Honianp, 1896,
Proc. Zool. Soc. London, p. 101. Masruue, 1904, Gen. Ins., XVII, p. 88.
One good specimen taken at Medje, July 13, 1910.
(481) 2. Rhopalocampta unicolor (Mabille)
Ismene unicolor MABILLE, 1877, Ann. Soc. Ent. France, (5) VII, Buil., p. xxxix.
Hesperia unicolor TRimEN, 1889, 8. Afr. Butt., IIT, p. 375.
Rhopalocampta unicolor HOLLAND, 1896, Proce. Zool. Soc. London, p. 97. MaBILLE,
1904, Gen. Ins., XVII, p. 88.
Six specimens: two taken at Gamangui in February; three cap-
tured at Medje in July; and one caught at Faradje, ticketted ‘1911~
1912.”
(482) 3. Rhopalocampta forestan (Cramer)
Papilio forestan CRAMER, 1782, Pap. Exot., IV, p. 210, Pl. cccxcr, figs. FZ, F.
Ismene florestan TRIMEN, 1866, (part), Rhop. Afr. Austr., II, p. 318.
Hesperia florestan TRimEN, 1889, 8. Afr. Butt., IIT, p. 368.
Rhopalocampta forestan Watson, 1893, Proc. Zool. Soc. London, p. 129. HoLuanp,
1896, Proc. Zool. Soe. London, p. 98.
Eight specimens are included in the collection; one taken at
Gamangui in February, one at Medje in June, and six at Niangara in
November 1910.
M. Mabille has omitted this species from his list given on page
88 of his work.
HETEROCERA
Amatide Hampson
(Syntomide of authors)
Myopsycxe Hampson
(483) 1. Myopsyche langi, new species
&. Head and thorax black with a faint bluish gloss; antenne black at the base
(tips wanting in the type); frons black; a tuft of orange-red hairs behind each eye;
patagia orange-red with a minute black spot at the base of each; first three segments
of abdomen orange-red, the succeeding segments blue-black, except the last, which is
pale reddish; the ventral surface of the thorax and the first three abdominal segments
are pale fawn-colored, and the legs are of the same color, becoming slightly darker
upon the femora. Fore and hind wings hyaline. The costa of the fore wing is
narrowly blackish, as is also the posterior margin; the border of the external margin
1920] Holland, Lepidoptera of the Congo 263
of the fore wing is broadly black at the apex, but rapidly decreases in width to the
extremity of the second median nervule, and then suddenly widening inwardly and
again diminishing to the inner angle, forms a subtriangular spot on the outer margin
from the second submedian nervule to the angle. This black spot is ornamented by a
minute white or hyaline spot between veins 2 and 3 near their extremities. At the
end of the cell of the fore wing there is a subtriangular black spot. The black outer
margin of the hind wing is relatively wide, and the costal margin is broadly black.
The outer margin has an inwardly projecting tooth-like prolongation on vein 2.
In consequence of the width of the black borders of the hind wing the inner hyaline
area is much reduced, and restricted to the lower half of the cell, a small space beyond
it, and the basal portion of the space between the cell and vein 1. Expanse 2, 27 mm.
This species comes nearest M. ochsenheimeri (Boisduval) but is
totally distinct, differing both in the markings of the
wings and the abdomen. I name it in honor of the
leader of the expedition upon the Lepidoptera col-
lected by which I am reporting. The type, which is
unique, was collected at Medje, August 6, 1910, and is Fig. 2.
in The American Museum of Natural History. Myopsyche langi, 3-}.
\ TRIcHZTA Swinhoe
(484) 1. Tricheta bivittata (Walker)
Syntomis bivittata Watker, 1864, List Lep. Het. B. M., XX XI, p. 66.
Tricheta bivittata Hampson, 1898, Cat. Lep., Phal., I, p. 55.
One specimen taken at Malela, July 8, 1915.
Amata Fabricius
(Syntomis Ochsenheimer and others)
(485) 1. Amata cerbera (Linnzeus)
Sphinx cerbera Linnxvs, 1764, Mus. Lud. Ulr., p. 363.
Syntomis cerbera Hampson, 1898, Cat. Lep., Phal., I, p. 83.
One poor specimen, labelled as taken at Stanleyville, but without date.
(486) 2. Amata marina (Butler)
Syntomis marina ButTLER, 1876, Journ. Linn. Soc. London, Zool., XII, p. 348.
Syntomis ogovensis HOLLAND, 1893, Psyche, VI, p. 374.
One defective specimen caught at Gamangui, June 17, 1910.
Meacanaciia Aurivillius
(487) 1. Meganaclia perpusilla (Walker)
Anace perpusilla WALKER, 1856, List Lep. Het. B. M., VII, p. 1720, #.
Metarctia perpusilla Hampson, 1898, Cat. Lep., Phal., I, p. 144, Pl. v, fig. 19.
Meganaclia carnea Hampson, 1898, Cat. Lep., Phal., I, p. 136, Pl. v1, fig. 27.
Meganaclia perpusilla Hampson, 1914, Cat. Lep., Phal., Suppl., I, p. 61.
One female, taken at Matadi, June 24, 1909.
264 Bulletin American Museum of Natural History (Vol. XLIII
Metarctia Walker
(488) 1. Metarctia invaria (Walker)
Anace invaria WALKER, 1856, List Lep. Het. B..M., VII, p. 1720.
One male, labelled as taken at Stanleyville, but without date.
(489) 2. Metarctia erubescens Walker
Metarctia erubescens WALKER, 1864, List Lep. Het. B. M., XX XI, p. 315.
A pair taken at Medje, July 1910.
(490) 3. Metarctia lutea Holland
Metarctia lutea HoLuANp, 1893, Psyche, VI, p. 396.
A male caught in August and a female captured in June 1910,
both at Medje.
Sir George F. Hampson in his Catalogue treats the last two forms
here listed as being merely varieties of M. invaria (Walker). I find it
difficult to agree with him. There are before me, as I write, some
hundreds of specimens of Walker’s species, representing numerous
localities, and they all agree very closely; there are also some scores of
each of the other two forms from widely separated localities, and these
also agree closely and are easily distinguishable at a glance from typical
M. invaria and from each other. Until the test of breeding shall have
demonstrated the fact that these three forms are derived from one and
the same batch of eggs, it appears to me preferable to distinguish them
as species, instead of “lumping” them under Walker’s name.
(491) 4. Metarctia hematica Holland
Metarctia hematica HOLLAND, 1893, Psyche, VI, p. 396. Hampson, 1898, Cat. Lep.,
Phal., I, p. 147, Pl. v, fig. 26.
One male caught at Gamangui, February 5, 1910.
(492) 5. Metarctia chapini, new species
Plate XIII, Figure 4, 9, type
9. Antenne and eyes black; frons, head, and thorax on the upper side black;
abdomen on the upper side with the first six segments very pale pinkish, margined
terminally with deep black, the remaining segments deep black; thorax and abdomen
on the under side pale fuscous, the banding which is very conspicuous on the upper
side being only faintly indicated on this side in the case of the anterior segments of
the abdomen; legs dorsad pale pinkish, ventrad black. Fore wings on the upper
side sooty black, without any trace of paler markings, on the under side pale fuscous,
slightly darker near the apex, and at the base near the inner margin passing into pale
pinkish. Hind wings on the upper side pale pink, the nervules and outer margin
being slightly clothed with pale fuscous squamation, on the under side dark fuseous,
except along the inner margin, where they are pale pink. Expanse, 45 mm.
1920] Holland, Lepidoptera of the Congo 265
The type, which is unique, was taken at Medje, June 26, 1910, and
isin The American Museum of Natural History. I take pleasure in
naming the species in honor of Mr. J. P. Chapin, one of the leaders
of the expedition.
The species comes near M. rubripuncta Hampson, but may be easily
distinguished from that species by the entire absence of the pale spots
and markings of the upper side of the primaries, the very strongly con-
trasting banding of the upper side of the abdomen, and the pale color
of the hind wings on the upper side.
(493) 6. Metarctia species
There are two specimens representing a very small species of the
genus, which may possibly be new to science, and which were taken at
Faradje, November 22, 1912. The specimens are, however, too badly
rubbed to permit either of identification or description.
(494) 7. Metarctia species
There is a single specimen bearing the label ‘‘Avakubi, Lieut.
Boyton, 1908,” which I find difficulty in referring to any species
hitherto described or figured. It isa male. Inasmuch, however, as the
specimen does not appear to be in prime condition, I refrain from at-
tempting to name or describe it, further than to say that the prevalent
color of the primaries is pale fawn, with obscure lighter discocellular
and postmedian markings on the primaries, and that it is a rather small
species, less than 40 mm. in expanse of wings.
Batacra Walker
(495) 1. Balacra ehrmanni (Holland)
Automolis ehrmanni HOLuanD, 1893, Psyche, VI, p. 535.
One female, labelled as taken at Bafwabaka in January 1910.
(496) 2. Balacra pulchra Aurivillius
Balacra pulchra Aurivituius, 1892, Ent. Tidskr., XIII, p. 200.
Balacra glagoessa HOLLAND, 1893, Psyche, VI, p. 396.
Pseudapiconoma glagoessa Hampson, 1898, Cat. Lep., Phal., I, p. 151, Pl. v1, fig. 14.
One female caught at Medje, July 1910.
Evucuromia Hiibner
(497) 1. Euchromia lethe (Fabricius)
Zygena lethe Fasricius, 1775, Syst. Ent., p. 553.
Euchromia lethe Hampson, 1898, Cat. Lep., Phal., I, p. 296.
266 Bulletin American Museum of Natural History [Vol. XLIII
The collection contains twenty-one specimens, mostly males, taken
_as follows: Kwamouth, 1, July 1909; Stanleyville, 7, August 1909;
Risimu, 1, September 1909; Avakubi, 1, October 1909; Medje, 2,
July—August 1910; Niangara, 5, November 1910; Faradje, 4, Novem-
ber 1911.
(498) . 2. Euchromia guineénsis (Fabricius)
Zygena guineénsis Fasricrus, 1775, Ent. Syst., p. 551.
Sphinx sperchia CRAMER, 1777, Pap. Exot., I, Pl. cxuvt, fig. C.
Euchromia sperchia Hampson, 1898, Cat. Loot Phal., Ip. 296; 1914, idem, Suppl., be
p. 197.
There are three examples captured as follows: Stanleyville, 1909;
Faradje, November 1912; Niangara, March 1913.
Arctiide
Nolinz
Nota Leach
(499) , 1. Nola banane, new species
@. Frons, tegule, and patagia pure snowy white; upper side of thorax and
abdomen a shade darker, very pale gray; lower side of abdomen and thorax much
darker gray; legs concolorous. On the upper side the fore wing is pale gray, growing
slightly darker externally towards the outer margin; a dark gray
spot of raised scales near the end of the cell; beyond it on the
costa a larger dark gray subtriangular spot; a postmedian trans-
verse, slightly curved line, composed of black dots on the
interspaces, runs from the costa to the inner margin, followed by a
submarginal row.of smaller spots. The hind wings on the upper
Fig. 3. side are evenly pale fuscaus. Both wings on the under side are uni-
Nola banane, #. $. formly gray, of adarker shadethan onthe upper side, but both have
their inner margins narrowly lined with white. Expanse, 16 mm.
The type, which is unique, was taken at Banana, June 21, 1909.
Lithosiinz
Cuionazma Herrich-Schieffer
(500) 1. Chionema delicata (Walker)
Bizone delicata WALKER, 1854, List Lep. Het. B. M., II, p. 550.
Chionama delicata Hampson, 1900, Cat. Lep., Phal., III, p. 325, Pl. xxvun, fig. 28.
One female taken at Medje in June.
Asura Walker
(501) 1. Asura atricraspeda Hampson (?)
Asura atricraspeda Hampson, 1914, Cat. Lep., Phal., Suppl., 1, p. 766.
A specimen taken at Medje, July 17, 1910, is referred to this species
with reasonable certainty, but it is too badly rubbed to make the identi-
fication positive,
1920] Holland, Lepidoptera of the Congo 267
Arctiinz
Driacrisia Hiibner
(502) 1. Diacrisia aurantiaca (Holland)
Alpenus aurantiacus HoLLAND, 1893, Psyche, VI, p. 397.
_ Diacrisia aurantiaca Hampson, 1901, Cat. Lep., Phal., III, p. 275, Pl. xxrv, fig. 19.
One male, minus antenne, caught at Medje, July 24, 1910.
(503) 2. Diacrisia maculosa (Stoll)
Bombyx maculosa Stouu, 1781, Pap. Exot., IV, Pl. cccuxx, fig. B.
Four males and three females of this common species, all taken at
Medje in June and July, except a male and a female caught at Faradje
in December.
(504) --3.“